taxonID	type	description	language	source
513E7520D308FFD6FE0D54B9B465FBA3.taxon	type_taxon	Type species Hanleya debilis Gray, 1857 (= Chiton hanleyi Bean in Thorpe, 1844), by monotypy.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D308FFD6FE0D54B9B465FBA3.taxon	description	Stratigraphic range Upper Oligocene – Recent.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D308FFD6FE0D54B9B465FBA3.taxon	distribution	Distribution Atlantic Ocean and adjacent seas, from southern Greenland and the western Barents Sea (74.27 ° N) to the Brazilian coast (25.44 ° S), including the Mediterranean Sea, and the Gulf of Mexico. Bathymetric range extends from the intertidal zone (H. hanleyi) to 1680 m (H. nagelfar).	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D308FFD6FE0D54B9B465FBA3.taxon	diagnosis	Diagnosis The following diagnosis for the genus Hanleya is based on data from Sirenko (1997), Jardim and Simone (2010 b), and new data in this publication.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D308FFD6FE0D54B9B465FBA3.taxon	discussion	Animal small to large. Insertion plate of head valve well developed, a primordium of insertion plates present in intermediate valves in all species. Insertion plates of tail valve well developed in Hanleya harasewychi sp. nov. and H. brachyplax, developed to a variable extent in H. hanleyi, H. nagelfar and H. mediterranea sp. nov., entirely absent in H. tropicalis. Tegmentum with large roundish or oval granules, each granule with one megalaesthete surrounded by 8 – 16 micraesthetes. Girdle covered with spicules or scales and longer, randomly dispersed needles or spines. Heads of major lateral teeth of radula with tridentate cusp, central denticle largest.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D309FFD8FDD55604B369FA09.taxon	description	(Figures 2 – 5)	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D309FFD8FDD55604B369FA09.taxon	materials_examined	Type material Holotype, USNM 93189, BL = 15.0 mm, now disarticulated, consisting of a mount of the shell, perinotum and radula, and 3 paratypes, USNM 323067, 454683, 454684. Type locality 22 mi ESE of Cape Hatteras, North Carolina (35.0800 ° N, 75.0530 ° W) U. S. Fish Commission Steamer Albatross, Sta. 2595, 1885, in 63 fathoms (115.2 m), sand, temp, 75 ° F (holotype). Material examined Holotype, USNM 93189, BL = 15.0 mm, 22 mi ESE of Cape Hatteras, North Carolina (35.0800 ° N, 75.0530 ° W) U. S. Fish Commission Steamer Albatross, Sta. 2595, 1885, in 63 fathoms (115.2 m), sand, temp, 75 ° F; Paratype 1, USNM 323067, BL = 5.0 mm, off Key West, Florida (24.28 ° N, 81.50 ° W), U. S. Fish Commission Steamer Albatross, Sta. 2317, 45 fathoms (81.9 m) on coral, 75 ° F (water column temperature?); Paratype 2, USNM 454683, BL = 3.0 mm, off Fowey Light, Florida (25.35262 ° N, 80.4508 ° W), sponges, Eolis Sta. 188, 48 fathoms (87.4 m), Henderson Coll., 1915; Paratype 3, USNM 454684, BL = 3.0 mm, Off Fowey Light, Florida (25.35262 ° N, 80.4508 ° W), Eolis Sta. 109, 40 fathoms (72.8 m), Henderson Coll., 1914.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D309FFD8FDD55604B369FA09.taxon	distribution	Distribution Off Cape Hatteras, North Carolina to the Florida Keys, in 73 – 115 m.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D309FFD8FDD55604B369FA09.taxon	diagnosis	Diagnosis Animal small (to 15.0 mm), elevated (dorsal elevation of valve V 0.36), back rounded. Head valves, lateral areas of intermediate valves, postmucronal areas sculptured with roundish granules arranged without distinct order, pleural areas of intermediate valves sculptured with long granules, ratio of granule length to granule width in pleural areas, 2.5 – 3.0; interspaces between rows of granules on pleural areas twice as large as width of granules; width of jugal sinus equal to width of apophyses in intermediate valves; insertion plate of tail valve well developed and pectinate; each granule contains a single megalaesthete and 12 – 13 micraesthetes. Valves and girdle buff coloured, spotted with brown and white. Dorsal spicules of girdle flat with one central longitudinal rib distally terminated with sharp beak. Radula with 15 transverse rows of mature teeth; exterior denticle of major lateral teeth of radula half the size of interior one, width of central denticle is equal to its length.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D309FFD8FDD55604B369FA09.taxon	description	Description Holotype dry, curled, about 15.0 mm long, 6.0 mm wide when straightened (Figure 2 F and H). Valves rounded (Figure 2 E). Head valve semicircular (Figure 2 A) posterior margin widely V-shaped, tegmentum sculptured with roundish granules arranged without regular pattern. Intermediate valves more or less rectangular (Figure 2 B), side margins evenly rounded, posterior margin nearly straight except for a small projecting apex, lateral areas not elevated, with sculpture as in head valve (Figure 2 G), pleural areas sculptured with elongated oval granules (about 180 × 60 μm) arranged in longitudinal rows, interspaces nearly twice as broad as width of granules, tegmentum of jugal area sculptured with small, elongated granules arranged without pattern (Figure 3 B). Each granule contains a single megalaesthete and 12 – 13 micraesthetes (Figure 4 E). Tail valve as wide as head valve, more or less oval; posterior margin well rounded, anterior margin straight to concave (Figure 2 B); mucro central, highly elevated, prominent; length of antemucronal area equal to that of postmucronal area (Figure 2 D), the latter deeply concave, sculptured like the head valve; antemucronal area sculptured like central area of intermediate valves (Figure 3 D). Tegmentum and girdle buff coloured with white and brown blotches. Articulamentum white, solid, blotched with rose in front of the central portion. Apophyses wide, long, with primordium of insertion plates extending nearly to end of lateral areas (Figures 3 E and 5 C), jugal sinus as wide as apophyses. Insertion plates well developed both in head and tail valves (Figure 3 A and C) (length of insertion plates 0.4 – 0.5 mm in head valve, 0.2 mm in tail valve); insertion plates obsoletely striate in head valve, pectinated in tail valve. Girdle narrow in present condition, densely covered with small, slender, slightly curved, pointed scales (Figures 3 E and 4 B) each with a single, central, longitudinal rib distally terminated by a sharp beak (≤ 60 – 70 × 14 – 18 μm), as well as randomly dispersed straight to slightly curved needles (Figure 4 A) varying in size from 230 × 20 μm to 470 × 30 μm. Marginal needles like dorsal needles (Figure 4 C). Ventral scales narrow, sharply pointed, reaching 80 × 17 μm in middle part of girdle, 120 × 20 μm near its margin (Figure 4 C and D). Radula 4.5 mm long, with 15 transverse rows of mineralised mature teeth (total about 20 rows). Central tooth short, almost rectangular, with narrow blade (Figure 4 F). Major lateral tooth with tridentate cusp (Figure 4 G and H), central denticle largest, as wide as long, exterior denticle smallest, half size of interior denticle. Gills merobranchial, adanal, without interspace, with 14 ctenidia per side, gills extending from the end of valve VI to anus. It is not possible to find a nephropore or gonopore in these dry specimens. Age variability With increasing body size, the width / length ratio of intermediate valves measured using valve V decreases from 2.7 (BL = 5.0 mm) to 2.3 (BL = 15.0 mm), as does the ratio of head valve length / head valve insertion plate length, from 6.0 (BL = 5.0 mm) to 4.6 (BL = 15.0 mm) (Figures 2 A, B and 5 A, B, D). The smaller paratype (BL = 5.0 mm) has similar dorsal spicules and needles that are slightly shorter and half as wide as those of the holotype (Figure 4 I and J). Granules on jugal area in paratype and holotype are similar but on the pleural area of paratype a little bit longer than that of holotype (Figures 2 G, 3 B, D, 5 E, F). Comparative remarks (Table 1) Hanleya harasewychi sp. nov. is readily distinguished from its congeners by its buff colour with white and brown blotches, rounded back, longer granules and larger interspaces on pleural areas of valves, fewer rows of mature teeth in the radula, and a shorter central denticle on the major lateral tooth of the radula. The colour of the tegmentum remains visible after preservation and is a reliable taxonomic character. Hanleya mediterranea sp. nov. has similar dorsal girdle scales, but differs from H. harasewychi in having larger, joined granules on the tegmentum (elongate in H. harasewychi), as well as similar-sized exterior and interior denticles on the major lateral tooth (very small exterior denticle in H. harasewychi). Hanleya nagelfar, which also has wide apophyses, differs from H. harasewychi in having dorsal girdle spicules that are ribbed in young specimens, but nearly smooth and less developed in mature adults, and in lacking a pectinated insertion plate on the tail valve. Hanleya tropicalis, which also has a small exterior denticle on the major lateral tooth of the radula (Figure 8 K), can be distinguished from the new species by its more elevated shell and absence of an insertion plate in the tail valve. Hanleya brachyplax also has a well-developed insertion plate on the tail valve, but can be separated from H. harasewychi by its white colour, shorter granules and fewer interspaces on pleural areas of valves, longer central denticle on the major lateral tooth of the radula, as well as by having longer and narrower apophyses, and a less dorsally elevated shell.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D309FFD8FDD55604B369FA09.taxon	etymology	Etymology This new species honours Dr. M. G. Harasewych, Curator of Marine Molluscs, Department of Invertebrate Zoology, National Museum of National History, Smithsonian Institution for his contributions to malacology and for his help and support during the preparation of this publication.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D307FFC0FDD057D9B106FE5A.taxon	description	(Figures 6 – 8 A – J)	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D307FFC0FDD057D9B106FE5A.taxon	description	length	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D307FFC0FDD057D9B106FE5A.taxon	materials_examined	Type material Holotype, ZISP 2201, BL = 4.7 mm, now disarticulated, consisting of mounts of shell, perinotum and radula, and three paratypes, ZISP 2202. Type locality Mediterranean Sea, off Begur, Girona, Spain, in 200 – 300 m. Material examined Holotype and three paratypes, ZISP 2202, BL = 4.0 – 4.7 mm, Mediterranean Sea, off Prinkipo, Turkey, 40.5139 ° N, 29.0724 ° E, in 50 m.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D307FFC0FDD057D9B106FE5A.taxon	distribution	Distribution Mediterranean Sea off Spain and Turkey, in 50 – 200 m.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D307FFC0FDD057D9B106FE5A.taxon	diagnosis	Diagnosis Animal very small, to 4.7 mm long, elevated (dorsal elevation 0.37) subcarinate, valves beaked; anterior margin of intermediate valves splayed laterally, with large inward curve in jugal area. Several granules of tegmentum in pleural areas joined, each granule with single megalaesthete and 9 – 16 micraesthetes; girdle covered dorsally with long, pointed scales that are smooth or with one central rib, as well as with numerous long needles. Insertion plates well developed both in head and tail valves.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D307FFC0FDD057D9B106FE5A.taxon	description	Description Holotype small, BL = 4.7 mm. Valves subcarinated, beaked (Figure 6 D), shell and girdle white. Head valve semicircular, posterior margin wide, V-shaped, notched in the middle, tegmentum sculptured with roundish granules (diameter 70 – 80 µm), arranged without pattern (Figure 6 A). Intermediate valves trapeziform, anterior margin greatly splayed laterally, with large inward curve in jugal area, posterior margin with small beak, lateral areas sculptured like head valve with rare round granules, jugal area sculptured with oval granules (about 80 × 50 µm) arranged without interspaces or pattern (Figure 6 B). Pleural areas sculptured with oval granules (120 × 70 µm) arranged without pattern, several granules join to form larger granules (up to 220 × 160 µm) (Figures 6 E and 8 I, J). Intermediate valves VI and VII have indistinct, short, longitudinal rows of granules in pleural areas near their posterior margin that are interrupted by large, joined granules in other part of pleural area. Each granule contains one megalaesthete surrounded by 9 – 16 micraesthetes (Figure 8 I). Tail valve narrower than head valve, more or less oval, mucro submedian, clearly indicated, postmucronal slope decidedly concave directly behind mucro, straightening near the posterior margin, antemucronal area sculptured like central area of intermediate valves, postmucronal area like head valve (Figure 6 C and F). Articulamentum strongly developed, insertion plates only on head valve 1 / 5 the length of the valve, apophyses wide, oriented laterally, connected to primordium of insertion plates in intermediate valves; insertion plates obsoletely striate in head and tail valves (Figure 6 G). Girdle narrow, 1 / 4 width of intermediate valve, covered dorsally with long pointed scales that may be smooth or with one central rib (50 – 86 × 23 µm) and numerous long smooth needles (330 – 550 × 24 µm). Marginal and dorsal needles similar. Ventral scales (50 – 55 × 18 µm) smooth typically, shorter than dorsal scales (Figures 7 A, B, C, and 8 C – E). Radula 2.2 mm long, with 28 transverse rows of mature teeth (total about 34 transverse rows). Central tooth short, nearly rectangular, with narrow blade; major lateral tooth with tridentate cusp, central denticle largest, exterior and interior cusps approximately equal (Figures 7 D and 8 F – H). Gills merobranchial, adanal, without interspaces, five ctenidia on each side, gills extending from valve VII to anus.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D307FFC0FDD057D9B106FE5A.taxon	etymology	Etymology From the Mediterranean Sea, where this species was found. Comparative remarks (Table 1) Hanleya mediterranea is distinguished from other species in the genus, including H. hanleyi, by its lack of longitudinal rows of granules across the entire pleural area, and by the presence of large granules comprising two or more small granules in pleural areas. Small specimens of Hanleya nagelfar, which have a similar radula, differ from H. mediterranea by having dorsal spicules that are oval in transverse section, with six perceptible ribs. In H. mediterranea (BL = 4.7 mm) the head valve has more strongly developed insertion plates that are 16 – 20 % the length of the valve, compared with H. nagelfar, (BL = 5 – 7 mm), in which the insertion plates are 3 – 9 % the length of the head valve. Hanleya tropicalis differs from H. mediterranea in having carinated valves, in lacking an insertion plate in the tail valve, and by having a smaller exterior denticle on the major lateral teeth of the radula (Figure 8 H and K). Hanleya harasewychi has similar dorsal scales on the girdle, but differs from H. mediterranea by having rounded valves, in the colour of the shell and girdle, by having longer granules on pleural areas of intermediate valves, as well by their arrangement, by a wider jugal sinus, a well-developed and pectinate insertion plate in the tail valve, and by a lower number of mineralised transverse rows of radular teeth as well as in the shape of the head of the major lateral teeth. Hanleya brachyplax has a similar radula, but differs from H. mediterranea in being less elevated, having a well-developed insertion plate in its tail valve, and by its larger size. Hanleya multigranosa (Reuss, 1860) from Middle Miocene deposits of Bohemia differs from the new species in having a shorter insertion plate on the head valve, a differently shaped tail valve, and smaller and more numerous granules on the tail valve. A single Pleistocene intermediate valve from Parma, Italy, which was identified as H. multigranosa by Sabelli and Taviani (1979: Table 1, Figure 4) most likely belongs to H. mediterranea. The records of fossil Hanleya from the Miocene of Poland, reported as H. multigranosa by Sulc (1934) and from the Upper Miocene of Italy, reported as H. hanleyi by Dell’ Angelo et al. (1999) should be confirmed by comparison of these fossil specimens with the new species described herein. Hanleya glimmerodensis Janssen, 1978 from the Upper Oligocene of western Germany differs from H. mediterranea in having well-marked longitudinal rows of granules on pleural areas of intermediate valves. Several publications report Hanleya hanleyi as occurring in the Mediterranean Sea (Monterosato 1879; Malatesta 1962; Sabelli 1972, 1974; Dell’ Angelo et al. 2004 and references in Dell’ Angelo and Smriglio 2001). Among these studies, Sabelli (1972) reported this species to reach a body length of 9 mm, and occur at depths ranging from 35 – 40 m to 140 m. Sabelli (1974) reported a body length of about 10 mm, while Dell’ Angelo and Smriglio (2001) reported a maximum body length of about 13 mm. Dell’ Angelo (personal communication) informed me that he has several specimens of Hanleya hanleyi collected in the Mediterranean Sea (BL <7 mm), from depths of 40 – 120 m and single valves dredged at depths up to 400 m in his collection. Dell’ Angelo and Smriglio (2001: pl. 25 C, Figure 36) included several photographs of chitons that have longitudinal rows of granules on central areas of intermediate valves. Although more than one species of Hanleya may inhabit the Mediterranean Sea, I provisionally attribute the above-mentioned records to H. mediterranea. Although the validity of Hanleya tropicalis and H. brachyplax is not in doubt, the relationship between H. hanleyi and H. nagelfar has been questioned. It is very difficult to distinguish small specimens of H. nagelfar from H. hanleyi because they both have a very similar sculpture of the tegmentum (Waren and Klitgaard 1991). As noted in the introduction, the discussion about differences between H. nagelfar and H. hanleyi has continued since the description of H. nagelfar. The relationship between H. hanleyi and H. nagelfar is further explored by a study of age variability in H. nagelfar.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D31FFFCAFDDB500CB19DFA6B.taxon	description	(Figures 9 – 19)	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D31FFFCAFDDB500CB19DFA6B.taxon	materials_examined	Type material Lectotype, NHRM, type collection 1329 (designated by Kaas and Van Belle (1985 )). Type locality Finnmark, Norway. Material examined Barents and Norwegian Seas: R / V Andrey Pervozvaniy, 1899, stn 4, 70.0230 ° N, 31.49 ° E, 144 – 178 m, 2 spms; stn 38, 69.58 ° N, 32.40 ° E, 230 m, 1 spm; 1900, stn 102, 74.27 ° N, 22.04 ° E, 180 – 211 m, 1 spm (Table 2, No. 8); stn 462, 71.20 ° N, 33.30 ° E, 304 – 280 m, 1 spm; stn 464, 71.21 ° N, 3159 ° E, 268 m, 1 spm. R / V Persey, 1924, stn 228, 70.3030 ° N, 31.13 ° E, 175 m, 1 spm (Table 2, No. 10); 1927, stn 677, 69.31 ° N, 34.39 ° E, 173 m, 1 spm; 1935, stn 3337, 71.35 ° N, 31.464 ° E, 313 m, 1 spm (Table 2, No. 11). R / V SRT 4225, cruise 1, 1955, stn 38 / 104, 61.308 ° N, 04.166 ° E, 300 m, 1 spm. R / V Sevastopol. RT- 97, cruise 8, 1958, Lofoten Ids., 68.08 ° N, 13.50 ° E, 1 spm (Table 2, No. 7); cruise 9, 1958, stn 1453, 62.002 ° N, 6.146 ° W, 112 m, 1 spm; stn 1488, 72.291 ° N, 23.072 ° E, 335 m, 2 spms. R / V Maslov, cruise 5, 1969, stn 180 / 201, 70.40 ° N, 17.275 ° E, 510 m, 1 spm; stn 182 / 203, 71.30 ° N, 16.46 ° E, 300 m, 1 spm (Table 2, No. 13); stn 183 / 204, 72.02 ° N; 17.02 ° E, 360 m, 1 spm (Table 2 No. 12); stn 188 / 209, 73.20 ° N, 15.25 ° E, 500 m, 1 spm. R / V Tunets, cruise 105, 1978, stn 4, 70.06 ° N, 32.325 ° E, 230 m, 1 spm; stn 11, 70.110 ° N, 17.350 ° E, 225 m, 2 spms; stn 12, 70.330 ° N, 16.540 ° E, 1100 m, 5 spms. R / V Dalnie Zelentsi, cruise, 16.1982, stn 64, 67.559 ° N, 10.080 ° E, 280 m, 3 spms. F. D. Behrens, 1948, Lofoten, 2 spm (ZIM). F. D. Saturn, 1958, 64.44 ° N, 35.06 ° W, 500 m, 1 spm. (ZIM); 1923, Folden, 50 – 100 m, 1 spm, (TM 13350); Skattora, Tromso, 54 m, 2 spms, (Table 2, No. 5), (TM 12923); 1951, Senja, Steinavar, 360 – 365 m, 1 spm, (TM 13335); 1954, Morsdalsfjord, 50 – 110 m, 1 spm, (TM 13196); 1954, Balsnes, 28 m, 1 spm, (TM 13258); 1954, Brokskar, Rystr, 50 – 120 m, 1 spm (Table 2, No. 3), (TM 12962); Beiarn, 40 – 90 m, 1 spm, (TM 13237); 1955, Tjelsund, 60 – 110 m, 1 spm, (TM 12955); 1955, Holandsfjord, 45 – 770 m, 1 spm, (TM 13241); 1955, Eiet, 52 – 58 m, 6 spms, (TM 13001); Masoy, 125 – 90 m, 2 spms, (TM 13299); 1955, Bjarandfjord, 76 – 92 m 1 spm, (TM 12997); 1955, Rana, 35 – 42 m, 1 spm, (TM 13304). Eastern Greenland Sea: R / V Atlantida, cruise 10, 1973, stn 18 / 1668, 74.50 ° N, 16.00 ° E, 360 m, 1 spm. Southeastern Greenland: R / V Sevastopol, RT- 97, cruise 5, 1957, stn 1168, 66.162 ° N, 31.356 ° W, 345 m, 2 spms; stn 1609, 65.526 ° N, 29.586 ° W, 425 m, 1 spm. Central and North Atlantic Ocean: R / V Akademik Keldysh, cruise 4, 1982 stn 513, 34.261 ° N, 30.030 ° W, 690 m, 1 spm; stn 444, 58.208 ° N, 31.363 ° W, 1635 – 1535 m, 1 spm; stn 364, Reykjanes Ridge, 1330 m, 1 spm; st. 385, Reykjanes Ridge, 1680 – 1800 m, 1 intermediate valve; R / V Ichtiandr cruise 8, 1982, stn 39, 09.038 ° N, 21.101 ° W, 610 – 700 m, 1 spm; stn 50, 33.141 ° N, 29.425 ° W, 700 – 740 m, 1 spm. Newfoundland Bank: R / V Sevastopol, RT- 97, cruise 16, 1960, stn 2805, 49.477 ° N, 50.185 ° W, 318 – 324 m, 1 spm; R / V Nereida, 2009, cruise 0509, stn 6, cod 5, 48.2915 ° N, 44.0609 ° W, 1348 m, 1 spm; stn 10, cod 54, 48.0005 ° N, 43.7607 ° W, 1554 m, 1 spm; stn 12, cod 16, 47.7772 ° N, 43.5748 ° W, 1462 m, 1 spm; stn 19, cod 21, 47.1643 ° N, 43.5338 ° W, 1358 m, 2 spms (Table 2, No. 4), stn 22, cod 12, 46.8400 ° N, 43.6404 ° W, 956 m, 1 spm (Table 2, No. 6); stn 23, cod 22, 46.7749 ° N, 43.8651 ° W, 1127 m, 3 spms; stn 24, cod 27, 46.6942 ° N, 43.9686 ° W, 1104 m, 1 spm; 2010, cruise 0610, stn 63, cod 8, 46.3697 ° N, 44.9944 ° W, 1406 m, 2 spms; stn 67, cod 7, 46.2677 ° N, 46.5429 ° W, 613 m, 1 spm; stn 69, cod 10 a, 46.1543 ° N, 46.4503 ° W, 502 m, 1 spm; stn 69, cod 10 c, 46.1543 ° N, 46.4503 ° W, 502 m, 2 spms; stn 71, cod 10, 46.0800 ° N, 46.3421 ° W, 888 m, 1 spm; cruise 0710, stn 75, cod. 24, 45.8692 ° N, 46.8210 ° W, 1227 m, 1 spm (Table 2, No. 1); stn 96, cod. 30, 43.9894 ° N, 48.9512 ° W, 863 m, 1 spm; cruise 0810, stn 99, cod 11, 43.6147 ° N, 49.0781 ° W, 548 m, 1 spm; stn 100, cod 12, 43.4844 ° N, 49.1855 ° W, 694 m, 2 spm; stn 103, cod 20, 429433 ° N, 49.5621 ° W, 1154 m, 1 spm. FN 3 L, 2006, stn 2, 47.9415 ° N, 47.3725 ° W, 411 m, 1 spm; FN 3 L, 2006, stn 9, 47.9370 ° N, 47.1883 ° W, 460 m, 1 spm; FN 3 L, 2008, stn 37, 47.6480 ° N, 47.1340 ° W, 503 m, 2 spms; FN 3 L, 2008, stn 84, 46.4488 ° N, 47.2603 ° W, 396 m, 1 spm; FN 3 L, 2009, stn 89, 461100 ° N, 47.3333 ° W, 1005 m, 2 spms; PLA, 2008, stn 98, 43.6298 ° N, 48.9907 ° W, 1436 m, 4 spms (Table 2, No. 9). North America: Off Eastport, Maine (Jeffreys coll.), 1 spm, (USNM, 177511); Off Martha’ s Vineyard, Massachussetts, 1 spm, (USNM); Georges Bank, 270 m, 1 spm (USNM); Nova Scotia, 72 m, 1 spm (USNM).	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D31FFFCAFDDB500CB19DFA6B.taxon	distribution	Distribution North and Central Atlantic Ocean, the Barents, Norwegian and Greenland seas, near southern Greenland, North America, Europe and northern Africa, on Mid-Atlantic Ridge from 74.27 ° N to 09.038 ° N at depths from 28 to 1680 m, mostly deeper than 200 m.	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
513E7520D31FFFCAFDDB500CB19DFA6B.taxon	discussion	Large specimens of Hanleya nagelfar are easily distinguished from H. hanleyi (see Kaas and Van Belle, 1985), but small chitons (<10 mm) are difficult to identify. In order to study age variability in H. nagelfar, seventeen specimens of different sizes (BL = 5.0 – 73.0 mm), from different regions of the Atlantic Ocean were measured (Table 2, Figures 8 – 18). These measurements revealed that important changes occur in the shape of the valves, the thickness of the girdle, and the shape, size and ribbing of the dorsal spicules of the girdle. With increasing body length, the ratio of head valve width / tail valve width decreased from 1.27 (BL = 5.0 mm) to 0.84 (BL = 70.0 mm); the ratio of valve V width / valve V length decreased from 2.56 (BL = 5.0 mm) to 1.36 (BL = 70.0 mm); and the ratio of valve V width / width of adjacent girdle also decreased from 5.9 (BL = 5.0 mm) to 1.36 (BL = 70.0 mm) (Table 2). Small and medium size specimens (BL = 5.0 – 29.0 mm) have flattened, short and pointed spicules with 6 – 7 ribs arranged around the spicule (Figures 10 A, C, 12 A, B, 14 A, C, D, E, 16 A – C, 17 D, 18 A, C). With increasing age (specimen size) the spicules become longer and the ribs become less prominent or obsolete (Figures 18 B, D, 19 E).	en	Sirenko, Boris (2014): Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History 48 (45 - 48): 2913-2945, DOI: 10.1080/00222933.2014.963722, URL: http://dx.doi.org/10.1080/00222933.2014.963722
