taxonID	type	description	language	source
03C687E26F17D371FE67F9BED42FF9BB.taxon	type_taxon	Type species: Clubiona pallidula (Clerck, 1757).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F17D375FE48F9FED6A4FAA0.taxon	diagnosis	Diagnosis Four characters stand out for all members of the apiculata - group: (1) the widely separated PME located at least twice their diameter apart (Figure 2 G); (2) the tibiae and metatarsi of the anterior legs carrying pairs of elongated and slender spines (Figure 2 H – I); (3) the presence of the tubercle at dorsal cymbium (Figure 1 G); and (4) the well-developed retrolateral tibial apophysis (Figure 1 A – D). There are three pairs of spines on the ventral surface of tibia I and II (Figure 2 H – I, TS), whereas metatarsus has only one pair, but these spines are much longer, with their base originating on proximal part and their apices almost reaching distal margin of metatarsus (Figure 2 H – I, MS). Pairs of spines can also be found on the femora (Figure 2 H – I, FS). They are curved and elongate, situated on the dorsal side of all femora, whereas those on other leg segments are relatively shorter. Males of the apiculata - group, the hystrix - group and the pahilistapyasea - group possess a simple, U-shaped sperm duct without additional loop. However, the conformation of the palp is uniform in males of the apiculata - group. They can be differentiated from males of all other species-groups by the embolus emerging from a distinct conical base (compare Figure 1 A – D with 1 E), which occupies most of the apico-prolateral extent of the palpal tegulum – embolus with broad base, gradually emerges from the tegulum, and tapering towards apex in hystrix - group (Figure 1 E) and pahilistapyasea - group (see also Barrion and Litsinger 1995: 145, fig. 79 d). The apex of the embolus points apically instead of moving clockwise along the margin of the tegulum base (compare Figure 1 A – D with 1 E). The conductor represented by a transparent lamina, and is located next to the embolic base on apico-retrolateral portion of the tegulum (Figure 3 E) (conductor absent in hystrix - group as per Deeleman-Reinhold 2001). The retrolateral apophysis on the palpal tibia is distinctly enlarged and can vary in shape from broad, heavily sclerotized ridge (Figure 1 A – C) to long, thinnish and point (Figure 1 D). The ventral tibial apophysis is well developed; it is variable in shape from being an elevated ridge (Figure 1 D) to a massive column (Figure 1 A, F). The legs are not lengthened and devoid of a large spine (legs distinctly elongate and provided with a very large apical spine on tibia II in the males of pahilistapyasea - group). The conformation of the female genitalia is uniform and closely resembles that of the hystrix - group in many aspects. Females of both species groups possess sclerotized spermathecae situated anteriorly to hyaline bursae (Figure 2 A, D); their epigynal region is marked with posteriorly situated copulatory orifices. However, the concomitantly copulatory ducts are elongate; these ducts are simple and lack convolution (Figure 2 A, D) in the apiculata - group (insemination ducts usually with convolution in hystrix - group). All known females also possess an additional, glandular head on each spermatheca (Figure 2 C, E). Included species Clubiona apiculata sp. nov., Clubiona cylindriformis sp. nov., Clubiona cultrata sp. nov. and Clubiona conica sp. nov.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F17D375FE48F9FED6A4FAA0.taxon	distribution	Distribution Known only from Borneo.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F17D375FE48F9FED6A4FAA0.taxon	discussion	Remarks No type specimens of Clubiona hystrix Berland, 1938 are known to exist. The first author could not locate them in the collections of RMHN. The original description and illustrations of this species correspond well with those provided by Deeleman-Reinhold (2001: 104, fig. 14 – 16, male from Bali) and the specimen examined here, which was collected near the type locality in New Guinea (MHNG, Irian Jaya, Manokwari Province, Manokwari, Gunnung Meja, 200 m, 30 December 2000 – 1 January 2001, A. Riedel leg., Figure 1 E). The hystrix - group is one of several species-groups in the spider genus Clubiona – for a full listing of species and synonymies see Deeleman-Reinhold (2001: 92). The males of this group have the following characteristics: (1) the palpal tegulum is marked with U-shaped sperm duct without forming additional loop; (2) the conductor is absent (but clearly present in a specimen treated here: Figure 1 E); (3) the embolus is situated on the distal margin of tegulum, with its apex pointed retrolaterally. The females can be recognized by: (1) the copulatory orifices located on posterior half of the epigynal region, hidden by fold or hood; (2) the spermathecae and hyaline bursae are distinct.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F13D37BFE0DFA1AD522FC6C.taxon	description	(Figures 1 A, F, G, 2 D – I, 3 A – G)	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F13D37BFE0DFA1AD522FC6C.taxon	diagnosis	Diagnosis Males of C. apiculata sp. nov. can be recognized by the presence of a basal tubercle on the dorso-retrolateral side of the cymbium (Figures 1 G, 3 C, D), in combination with a colossal, rectangular, sclerotized VTA (Figures 1 F, Figure 3 B). Males are distinguished from other members of the species-group by the short triangular embolus, and by the heavily sclerotized RTA (compare Figure 1 A with Figure 1 B – D). Females can be recognized by the oblong spermathecae and the distinctly larger spherical bursae connected to thick-walled insemination ducts (Figure 2 D, 3 G).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F13D37BFE0DFA1AD522FC6C.taxon	materials_examined	Type material Holotype. ♂, Malaysia, Borneo, Sabah State, Kinabalu National Park, 2590 m, humid ravine below Layang Layang, cloud forest, 1 May 1987, D. Burckhardt & I. Löbl leg. (MHNG, 10 a). Paratype. One ♂, 2 ♀, same data as holotype (MHNG, 10 a).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F13D37BFE0DFA1AD522FC6C.taxon	etymology	Etymology The specific epithet is derived from the Latin adjective apiculatus, referring to the triangular embolus and its conical base, which are situated near the apex of the tegulum.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F13D37BFE0DFA1AD522FC6C.taxon	description	Description Male (holotype). Total length 6.9; prosoma 3.3 long, 2.0 wide; opisthosoma 3.6 long, 1.8 wide. Prosoma ovoid, in profile distinctly higher just in front longitudinal fovea; integument smooth; carapace uniform yellow; chelicerae yellowish brown, with three promarginal and three retromarginal teeth; labium and gnathocoxae pale brown; sternum yellow. Eye sizes and interdistances: AME 0.08, ALE 0.12, PME 0.18, PLE 0.16, AME – AME 0.08, AME – ALE 0.10, PME – PME 0.32, PME – PLE 0.18; MOQ: 0.42 long, 0.24 anterior width, 0.64 posterior width. Legs pale yellow, anterior metatarsi and tarsi brown, distinctly darker than other segments; scopulae and tufts indistinct. Leg formula 4213. Leg measurements: leg I 6.9 (1.9, 1.0, 1.8, 1.6, 0.6); leg II 7.6 (2.4, 1.0, 2.2, 1.4, 0.6); leg III 6.6 (1.6, 1.4, 1.4, 1.4, 0.6); leg IV 10.0 (2.8, 1.0, 2.4, 2.8, 1.0). Spines on anterior legs elongate and slender; anterior tibiae armed with three pairs of elongated ventral spines; anterior metatarsi armed with a pair of very long spines, originating proximally, apices almost reaching tarsus. Opisthosoma ovoid, pale, without colour marking; lightly sclerotized dorsal scutum extending half length of opisthosoma. Palp (Figures 1 A, F, G, 3 A – E). Palpal tibia with distinctly elevated VTA in ventral (Figure 3 A) and retrolateral views (Figure 3 C), broad and massive rectangular in prolateral view (Figures 1 F, 3 B); RTA petal-shaped, heavily sclerotized, reaching almost half length of cymbium; cymbium apically with shallow dorsal excavation, covered with thick patch of hairs; digitiform tubercle originating on baso-retrolateral side of cymbium, directed horizontally, pointing ectad; tegulum oblong; conductor digitiform, translucent, originating on apico-retrolateral area of tegulum; sperm duct U-shaped; embolic base conical, situated apico-prolaterally; embolus triangular, minute, apex sharply pointed, directed anteriad, emerging subterminally on embolic base. Female (paratype). Total length 5.6; prosoma 3.0 long, 2.6 wide; opisthosoma 2.6 long, 1.1 wide. General appearance as in male, including spination on anterior tibiae and metatarsi. AME 0.08, ALE 0.10, PME 0.18, PLE 0.14, AME – AME 0.08, AME – ALE 0.10, PME – PME 0.30, PME – PLE 0.16; MOQ: 0.38 long, 0.24 anterior width, 0.66 posterior width. Leg measurements: leg I 5.9 (1.7, 0.8, 1.6, 1.2, 0.6); leg II 7.0 (2.2, 0.8, 2.2, 1.2, 0.6); leg III 6.3 (1.6, 1.4, 1.2, 1.5, 0.6); leg IV 9.0 (2.6, 0.8, 2.2, 2.6, 0.8). Genitalia (Figures 2 D – F, 3 F, G). Epigynal region circular, yellow, weekly sclerotized; copulatory orifices situated underneath heavily sclerotized, V-shaped ridge protruding over epigastric furrow; insemination ducts thick-walled, twisted proximally to form semicircular loop, with funnel-shaped atrium internally, ducts ascending anteriorly and opening into base of spermathecae; spermathecae oblong, basolaterally with glandular head (Figure 2 E); acicular fertilization ducts membranous, located on dorsal side near base of spermathecae (Figure 2 F); bursae spherical, hyaline, approximately twice diameter of spermathecae, connected to insemination ducts at mid length between copulatory orifices and spermathecae. Natural history Clubiona apiculata sp. nov. is considered a high-altitude species. The type specimens were collected in a cloud forest (2590 m) by sifting decomposing plant and organic material in a humid ravine.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F13D37BFE0DFA1AD522FC6C.taxon	distribution	Distribution Known only from the type locality (Kinabalu National Park, Borneo; separated from C. conica sp. nov. by altitudinal gradient).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1DD378FE3DFC59D4AAFE41.taxon	description	(Figures 1 D, 2 A – C, 4 A – E)	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1DD378FE3DFC59D4AAFE41.taxon	diagnosis	Diagnosis The male of C. conica sp. nov. can be recognized by the long, sickle-shaped retrolateral apophysis on the palpal tibia, and by the very short, spiniform embolus (Figures 1 D, 4 A – C). The female is recognized by the thin-walled insemination ducts connected to pyriform spermathecae, and by the membranous bursae with distinct, elongated stalk (Figures 2 A, 4 D, E).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1DD378FE3DFC59D4AAFE41.taxon	materials_examined	Type material Holotype. ♂, Malaysia, Borneo, Sabah State, Kinabalu National Park, Poring Hot Spring, 500 m, 8 May 1987, D. Burckhardt & I. Löbl leg. (MHNG, 16 H). Paratype. One ♀, same data as for holotype (MHNG, 16 H).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1DD378FE3DFC59D4AAFE41.taxon	etymology	Etymology The specific epithet is derived from the Greek (κωΝΟϛ = conus = cone-shaped), alluding to a conical base of embolus.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1DD378FE3DFC59D4AAFE41.taxon	description	Description Male (holotype). Total length 5.8; prosoma 3.1 long, 2.8 wide; opisthosoma 2.7 long, 1.3 wide. Prosoma ovoid, pars cephalica broad, almost as wide as carapace width, anterior margin straight, in profile flat; integument smooth; fovea shallow, longitudinal; carapace orange brown, with distinctly brown anterior margin; chelicerae dark brown, with three promarginal and four retromarginal teeth; labium and gnathocoxae brown; sternum pale yellow, bordered with brown margin. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.22, PLE 0.20, AME – AME 0.10, AME – ALE 0.10, PME – PME 0.62, PME – PLE 0.30; MOQ: 0.40 long, 0.30 anterior width, 0.80 posterior width. Legs uniform pale yellow; scopulae on tarsi and metatarsi I – II distinct, in III – IV sparse. Leg formula 4213. Leg measurements: leg I 5.3 (1.6, 0.8, 1.4, 1.1, 0.4); leg II 6.6 (2.0, 0.8, 2.1, 1.2, 0.5); leg III 6.0 (1.5, 1.4, 1.2, 1.4, 0.5); leg IV 7.9 (2.2, 0.7, 2.0, 2.2, 0.8). Opisthosoma ovoid, pale, without colour marking; dorsal scutum lightly sclerotized, located on cardiac region, extending half length of opisthosoma, followed by pair of muscle apodemes. Palp (Figures 1 D, 4 A – C). Palpal tibia short, approximately half length of cymbium; dorsal surface with blunt tubercle; ventro-prolateral surface with broad, triangular, sclerotized VTA; RTA sickle-shaped, greatly elongate, extending anteriorly then curving backwards, tapering towards sharply pointed apex; cymbium with apicodorsal excavation, covered with thick patch of hairs; small, blunt tubercle located on meso-retrolateral side of cymbium; tegulum oblong, apical region weakly sclerotized; sperm duct U-shaped; embolic base dark, conical, situated medially on apical surface of tegulum; embolus spiniform, minute, apex sharply pointed, directed retrolaterad, emerging on apex of embolic base. Female (paratype). Total length 6.9; prosoma 3.5 long, 3.0 wide; opisthosoma 3.4 long, 1.8 wide. General appearance as in male, but entire body pale yellow, dorsal scutum on opisthosoma absent. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.20, PLE 0.20, AME – AME 0.10, AME – ALE 0.10, PME – PME 0.64, PME – PLE 0.32; MOQ: 0.46 long, 0.30 anterior width, 0.86 posterior width. Legs uniform yellow; scopulae on tarsi and metatarsi I – II distinct, in III – IV sparse. Leg formula 4213. Leg measurements: leg I 5.9 (1.8, 0.8, 1.6, 1.2, 0.5); leg II 7.3 (2.2, 0.8, 2.2, 1.4, 0.7); leg III 6.4 (1.6, 1.6, 1.2, 1.5, 0.5); leg IV 8.4 (2.4, 0.9, 2.0, 2.3, 0.8). Genitalia (Figures 2 A – C, 4 D, E). Epigynal region weakly sclerotized, posteriorly with heavily sclerotized, oval depression situated near epigastric furrow; copulatory orifices circular, separated from one another by their diameter apart, located at centre of epigynal depression; insemination ducts thin-walled, ascending anteriorly, turning outwards and connected to dorsal surface of pyriform spermathecae; spermathecae ventrally with obliquely located falcate projection (Figures 2 B, 4 D), apically with digitiform glandular head (Figures 2 C, 4 D, E); fertilization ducts ribbon-shaped, distinctly elongate (Figure 2 B, C); bursae membranous, with tubular stalk originating on dorsal surface of spermathecae. Natural history Clubiona conica sp. nov. is a lowland species. Type specimens were collected by sifting leaf and organic litter under dipterocarp trees.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1DD378FE3DFC59D4AAFE41.taxon	distribution	Distribution Known only from the type locality (Kinabalu National Park, Borneo).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1ED37EFDD8FE45D416FDA4.taxon	description	(Figures 1 B, 5 A – C)	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1ED37EFDD8FE45D416FDA4.taxon	diagnosis	Diagnosis Males of C. cylindriformis sp. nov. closely resemble those of C. cultrata sp. nov. in having the petal-shaped RTA; the cylindrical base of embolus freely emerging from the tegulum (cone-shaped in C. cultrata sp. nov.); the embolus being shorter and more slender (embolus blade-shaped and more elongate in C. cultrata sp. nov.). Males of both species possess a conspicuous, conical tubercle situated on the dorsal side of cymbium (Figure 5 C, E, F).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1ED37EFDD8FE45D416FDA4.taxon	materials_examined	Type material Holotype. ♂, Malaysia, Borneo, Sarawak State, Serian District, Penrissen Road 12 miles (19 km) of Kuching, Sernonkok Wildlife Rehabilitation Centre: Nursery Centre of the Forest Department, 20 – 50 m, 8 December 1987, C. Lionhard leg. (MHNG, Sar- 87 / 62).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1ED37EFDD8FE45D416FDA4.taxon	etymology	Etymology The specific epithet is a derived combination of Greek and Latin words (Gk. κυΛΙΝδΡΟϛ = cylindrus = cylindrical), referring to the shape of the embolic base.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1ED37EFDD8FE45D416FDA4.taxon	description	Description Male (holotype). Total length 5.8; prosoma 3.4 long, 3.0 wide; opisthosoma 3.2 long, 1.4 wide. Prosoma ovoid, convex, in profile highest mid-distance between PME and longitudinal fovea; integument smooth, devoid of hairs; carapace yellowish brown; chelicerae brown, with three promarginal and three smaller retromarginal teeth; labium and gnathocoxae pale brown; sternum pale yellow. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.20, PLE 0.18, AME – AME 0.10, AME – ALE 0.10, PME – PME 0.40, PME – PLE 0.32; MOQ: 0.30 long, 0.32 anterior width, 0.66 posterior width. Legs uniform pale yellow; scopulae on tarsi and metatarsi I – II distinct, in III – IV sparse. Leg formula 4213. Leg measurements: leg I 5.2 (1.6, 0.8, 1.2, 1.0, 0.6); leg II 6.8 (2.2, 0.8, 2.0, 1.2, 0.6); leg III 6.4 (1.6, 1.4, 1.4, 1.4, 0.6); leg IV 8.4 (2.4, 0.8, 2.2, 2.0, 1.0). Opisthosoma cylindrical, pale, without colour marking; dorsal scutum absent; one pair of muscle apodemes located mid-dorsally. Palp (Figures 1 B, 5 A – C). Palpal tibia compressed on both sides, with heavily sclerotized, triangular VTA; RTA heavily sclerotized, petal-shaped in ventral view, rectangular in retrolateral view; cymbium with apicodorsal depression, basoretrolaterally with triangular tubercle, apex blunt; tegulum oblong, apical portion weakly sclerotized; conductor broad lamina, beak-shaped in ventral view, originating apico-retrolaterally; sperm duct U-shaped, elongate; embolic base cylindrical, situated apico-prolaterally, gradually tapering towards apex; embolus spiniform, apex sharply point, directed anteriad. Female. Unknown. Natural history Type specimen of C. cylindriformis sp. nov. was collected by beating shrubs and lower vegetation in lowland forest.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F1ED37EFDD8FE45D416FDA4.taxon	distribution	Distribution Known only from the type locality (Kuching, Borneo).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F18D37DFDF9FD26D478FC6D.taxon	description	(Figures 1 C, 5 D – F)	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F18D37DFDF9FD26D478FC6D.taxon	diagnosis	Diagnosis The male of C. cultrata sp. nov. is almost identical to that of C. cylindriformis sp. nov. but can be distinguished by the conical embolic base (cylindrical base in C. cylindriformis sp. nov.); the blade-shaped embolus is longer, and the conductor is situated on a more prominent elevation (broad lamina in C. cylindriformis sp. nov.). Both species have a convex carapace and their PME are widely separated.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F18D37DFDF9FD26D478FC6D.taxon	materials_examined	Type material Holotype. ♂, Indonesia, Borneo, East Kalimantan Province, Berua District, near Kampung Suaran, c. 40 km south of Tanjungredeb (1 ° 59 ′ 42 ″ S, 117 ° 36 ′ 03 ″ E), 50 m, primary forest on limestone, 1 October 2008, P. J. Schwendinger leg. (MHNG, IND- 08 / 11).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F18D37DFDF9FD26D478FC6D.taxon	etymology	Etymology The specific epithet is derived from Latin (cultratus = blade-like), referring to the shape of the embolus.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F18D37DFDF9FD26D478FC6D.taxon	description	Description Male (holotype). Total length 6.0; prosoma 3.0 long, 2.8 wide; opisthosoma 3.0 long, 2.6 wide. Prosoma ovoid, strongly convex, in profile highest just in front longitudinal fovea; integument smooth; carapace orange uniform yellow; chelicerae brown, with three promarginal and three retromarginal teeth; labium and gnathocoxae pale brown; sternum yellow. Eye sizes and interdistances: AME 0.10, ALE 0.12, PME 0.20, PLE 0.18, AME – AME 0.10, AME – ALE 0.10, PME – PME 0.42, PME – PLE 0.30; MOQ: 0.30 long, 0.30 anterior width, 0.82 posterior width. Legs uniform pale yellow; scopulae on tarsi and metatarsi I – II distinct, in III – IV sparse. Leg formula 4213. Leg measurements: leg I 5.4 (1.4, 0.8, 1.6, 1.0, 0.6); leg II 6.8 (2.0, 0.8, 2.2, 1.2, 0.6); leg III 5.6 (1.2, 1.4, 1.2, 1.2, 0.6); leg IV 8.4 (2.4, 0.8, 2.0, 2.4, 0.8). Opisthosoma cylindrical, pale, without colour marking; dorsal scutum absent. Palp (Figures 1 C, 5 D – F). Palpal tibia laterally compressed, with sclerotized, triangular VTA on ventro-prolateral surface; RTA heavily sclerotized, petal-shaped in ventral view, more or less rectangular with rounded apical apex in retrolateral view; cymbium with triangular tubercle situated basally, apex blunt; tegulum oblong, apical region weakly sclerotized; conductor represented by digitiform lamina, originating apico-retrolaterally on elevated mound; sperm duct U-shaped; embolic base conical, situated apico-prolaterally, clearly elevated from tegulum; embolus blade-like, dorsoventrally flattened, apex bluntly pointed, obliquely directed ectad. Female. Unknown. Natural history Type specimen of C. cultrata sp. nov. was collected by sifting decomposing leaf and organic litter in lowland primary forest at the foot of limestone hill.	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
03C687E26F18D37DFDF9FD26D478FC6D.taxon	distribution	Distribution Known only from the type locality (Kalimantan, Borneo).	en	Dankittipakul, Pakawin, Singtripop, Tippawan (2014): A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History (J. Nat. Hist.) 48 (31 - 32): 1923-1936, DOI: 10.1080/00222933.2014.902140, URL: http://dx.doi.org/10.1080/00222933.2014.902140
