taxonID	type	description	language	source
EA3E7D706C0EE264FE1CB9D5FE6EFBB8.taxon	diagnosis	Diagnosis Adults: Fixed and movable cheliceral digits with about 14 and with four teeth, respectively. Anterior margin of epistome without extensions, convex, with 14 – 16 denticles in the median region. Seta sc much longer than h 1, in turn longer than the subequal h 2 and h 3. Deutosternal denticles in seven rows. Dorsal shield scantly reticulate, with 16 and 15 pairs of setae on podonotal and opisthonotal regions; with two pairs of setae on unsclerotized cuticle along margins of dorsal shield. Most of J, Z and S setae not reaching the insertion of the next seta in the respective series. All dorsal idiosomal setae ridged and, except for J 5, r 5 and R 1 with distinct lateral barbed flat expansions mostly visible in distal half (usually referred to as tricarinate); J 5 with few basal barbs; r 5 and r 1 smooth. Platelets between genital and ventrianal shield absent. With nine pairs of opisthogastric setae, four of which on ventrianal shield; the latter with a pair of small marginal pores about in transverse line with Jv 4. Tibia I with 13 setae. Gnathosoma (Figure 1 A – C). Cheliceral shaft length (from base of first segment to apex of fixed cheliceral digit) 91 (85 – 100) long; dorsal and lateral lyrifissures as well as dorsal seta distinct (the latter stout, slightly paraxial and somewhat appressed to surface of fixed digit, not easily discernable under phase contrast). Fixed digit 31 (30 – 33) long, with aciculate pilus dentilis and about 14 teeth, including offset subapical tooth, in addition to apical tooth; movable cheliceral digit 41 (36 – 47) long, with four teeth in addition to apical tooth. Anterior margin of epistome without extensions, convex and with 14 – 16 denticles in the median region; apex at level slightly anterior to base of palp trochanter. Corniculi tapered, well separated and subparallel to each other. Deutosternum relatively wide 7.5 (7 – 8); deutosternal denticles in seven rows; the first five of uniform lengths, delimited by lateral lines, with two to four denticles each; sixth row with 10 – 18 denticles divided in two separate groups that extend beyond lateral lines; seventh row with four denticles. Palpus 172 (145 – 200) long; numbers of setae from trochanter to tibia: 2, 5, 6, 14; apotele two-tined. Hypostomal and palp-coxal setae aciculate: h 1 31 (27 – 37), h 2 21 (20 – 22), h 3 23 (15 – 30) and sc 35 (30 – 39). Dorsum of idiosoma (Figure 1 D). Holodorsal shield 401 (347 – 460) long and 207 (185 – 245) wide at level slightly posterior to r 3; with distinct reticulation between J 1 and Z 4 and with irregular and scantly reticulation elsewhere. Podonotal region of dorsal shield with 16 pairs of setae (j 1 – j 6, z 2 – z 6, s 3 – s 6, r 3), five pairs of lyrifissures and two pairs of pores. Opisthonotal region of dorsal shield with 15 pairs of setae (J 1 – J 5, Z 1 – Z 5, S 1 – S 5), 11 pairs of lyrifissures and two pairs of pores. Lateral unsclerotized cuticle with two pairs of marginal setae (r 5 and R 1), a pair of lyrifissures and no pores. Lengths of dorsal setae: j 1 28 (25 – 35), j 2 17 (13 – 23), j 3 28 (25 – 33), j 4 21 (19 – 25), j 5 21 (20 – 24), j 6 22 (20 – 24), J 1 21 (20 – 24), J 2 22 (20 – 25), J 3 22 (20 – 25), J 4 22 (20 – 25), J 5 12 (10 – 15), z 2 23 (21 – 26), z 3 23 (21 – 28), z 4 23 (21 – 27), z 5 21 (19 – 24), z 6 21 (19 – 25), Z 1 23 (20 – 27), Z 2 22 (20 – 26), Z 3 28 (25 – 32), Z 4 44 (43 – 45), Z 5 65, s 3 24 (23 – 25), s 4 28 (27 – 30), s 5 25 (23 – 27), s 6 28 (27 - 30), S 1 24 (20 – 29), S 2 26 (23 – 32), S 3 29 (25 – 37), S 4 35 (29 – 44), S 5 37 (31 – 50), r 3 37 (30 – 47), r 5 19 (16 – 25), R 1 17 (15 – 20). Most of J, Z and S setae not reaching the insertion of the next seta in the respective series. All dorsal idiosomal setae ridged and, except for J 5, r 5 and R 1 with distinct lateral barbed flat expansions mostly visible in distal half (usually referred to as tricarinate); J 5 with few basal barbs; r 5 and r 1 smooth. Venter of idiosoma (Figure 1 E). Setae aciculate and smooth, except Jv 5 ridged, stout and barbed, and post-anal seta, stout and barbed. Base of tritosternum 13 (12 – 15) long and 12 (10 – 13) wide at its median length; laciniae fused for about 60 % of their total length [74 (87 – 58)]. Sternal shield smooth, lineate along lateral margins, bearing setae st 1 – st 3, and two pairs of lyrifissures. Metasternal plate rounded, bearing st 4 and a pair of lyrifissures. Genital shield 113 (112 – 115) long, including hyaline anterior flap, lateral margins moderately concave such that narrowest width between coxae IV [50 (45 – 55)] about 0.7 its posterior width [72 (70 – 75)]; truncate posteriorly, smooth, bearing genital seta (st 5); paragenital lyrifissure on unsclerotized cuticle posterolaterad of genital seta. Platelets between genital and ventrianal shields absent. Two pairs of metapodal plates, the anterior elongate and smaller than the posterior, which is oval to subtriangular. Ventrianal shield subtriangular, transversely reticulate with few diagonal connections between striae laterally, about as long [134 (122 – 152)] as wide [137 (125 – 155)], with four pairs of opisthogastric setae (Jv 1 – Jv 3, Zv 2) in addition to circumanal setae and with a pair of minute pores positioned on the margin of the shield, about in transverse line with insertion of Jv 4; postanal seta about 1.5 times as long as para-anal seta, anal opening not enlarged, its length (24 – 30 including frame) about 0.2 the length of ventrianal shield. Unsclerotized cuticle around ventrianal shield with five pairs of opisthogastric setae (Jv 4, Jv 5, Zv 1, Zv 3 and Zv 5) and three pairs of lyrifissures. Lengths of ventral setae: st 1 34 (30 – 42), st 2 29 (28 – 32), st 3 27 (26 – 28), st 4 25 (20 – 30), st 5 25 (20 – 32), Jv 1 15 (13 – 18), Jv 2 16 (14 – 28), Jv 3 16 (15 – 17), Jv 4 16 (14 – 18), Jv 5 35 (30 – 40), Zv 116 (13 – 19), Zv 2 15 (13 – 18), Zv 3 12 (10 – 16), Zv 5 19 (18 – 20), para-anal 20, post-anal 30. Endopodal plate distinct between coxae III and IV. Exopodal plate distinct, running from posterior margin of coxa IV to region between coxae I and II. Peritreme and peritrematic shield. Peritreme extending anteriorly to region between j 1; peritrematic shield fused to dorsal shield at level between s 3 and s 4 and to exopodal shield at level of coxa IV. Spermatheca (Figure 1 F). Major duct inflated in a pouch immediately after solenostome, follows as a narrow duct that flares towards a pronounced curve and then as a complex section that apparently corresponds to the vesicle, which is viewed as a variously shaped structure according to position in mounted specimens, seemingly with side projections and with a needle-like blind-ended structure. Legs (Figure 1 G – I). Pretarsi of legs I – IV consisting of elongate stalk, a pair of claws and pulvillus with three rounded lobes. Leg I shorter than, and leg IV about as long as, dorsal shield; lengths of legs: I, 356 (332 – 400); II, 313 (275 – 370); III, 280 (245 – 350); IV, 385 (320 – 475). All setae aciculate. Chaetotaxy of legs I – IV: coxa 2, 2, 2, 1; trochanter 6 (1 - 0 / 1, 1 / 2 - 1), 5 (1 - 0 / 1, 0 / 2 - 1), 5 (1 - 1 / 1, 0 / 2 - 0), 5 (1 - 1 / 1, 0 / 2 - 0); femur 12 (2 - 2 / 1, 2 / 3 - 2), 11 (2 - 3 / 2, 2 / 1 - 1), 6 (1 - 2 / 1, 1 / 0 - 1), 6 (1 - 2 / 1, 2 / 0 - 0); genu 12 (2 - 3 / 1, 3 / 1 - 2), 10 (2 - 3 / 1, 2 / 0 - 2), 7 (2 - 2 / 0, 2 / 0 - 1), 9 (2 - 2 / 1, 3 / 0 - 1); tibia 13 (2 - 3 / 2, 3 / 1 - 2), 10 (2 - 1 / 1, 3 / 1 - 2), 8 (1 - 2 / 1, 2 / 1 - 1), 10 (2 - 1 / 1, 3 / 1 - 2); tarsus II – IV 18, 18, 18. Adult male (one specimen measured) (Figure 2) Gnathosoma (Figure 2 A, B). Lyrifissures, dorsal seta and pilus dentilis of cheliceral shaft as in adult female; fixed cheliceral digit 19 long, and seven or eight teeth, including offset subapical tooth, in addition to apical tooth; movable cheliceral digit 23 long, with apparently one tooth in addition to apical tooth. Spermatodactyl 32 long, sinuate, tapering distally, with a small subapical heel. Anterior margin of epistome without extensions, denticulate as in adult female. Corniculi tapered, well separated and subparallel to each other. Deutosternum relatively wide, 4 (4 – 5); deutosternal denticles in seven rows; number of denticles in the first five rows not possible to determine; sixth row with 8 – 12 denticles; seventh row with four denticles. Hypostomal and palp-coxal setae aciculate: h 1 23, h 2 16, h 3 18 and sc 25. Dorsum of idiosoma (Figure 2 C). Holodorsal shield 290 long and 172 wide at level of r 3; with distinct reticulation on opisthonotal region of the shield and with irregular and scant reticulation elsewhere. Podonotal region of dorsal shield with 17 pairs of setae (j 1 – j 6, z 2 – z 6, s 3 – s 6, r 3, r 5), four pairs of lyrifissures and two pairs of gland pores. Opisthonotal region with 16 pairs of setae (J 1 – J 5, Z 1 – Z 5, S 1 – S 5, R 1), eight pairs of lyrifissures and one pair of pores. Form and relative lengths of dorsal shield setae as in adult female. Unsclerotized cuticle laterad of dorsal shield without setae but with a pair of lyrifissures. Lengths of dorsal setae: j 1 20, j 2 13, j 3 22, j 4 16, j 5 16, j 6 17, J 1 16, J 2 17, J 3 17, J 4 15, J 5 9, z 2 16, z 316, z 417, z 5 15, z 6 16, Z 1 17, Z 2 18, Z 3 23, Z 4 38, Z 5 45, s 3 16, s 4 19, s 5 20, s 6 20, S 1 18, S 2 20, S 3 20, S 4 20, S 5 21, r 3 29, r 5 14, R 1 10. Shape and relative length of dorsal idiosomal setae as in adult female. Venter of idiosoma (Figure 2 D). Base of tritosternum 10 long and 10 wide at its median length; laciniae fused for about 50 % of their total length (45). Sternogenital shield 127 long and 63 wide at level of st 2, smooth, lineate along lateral margins, bearing st 1 – st 5 and three pairs of lyrifissures. Ventrianal shield subtriangular, transversely striate with few diagonal connections between striae laterally, 130 long, 162 wide at anterolateral corners, covering most of opisthogaster, not fused with peritremal and sternogenital shields, bearing (Jv 1 – Jv 3, Zv 1 – Zv 3) in addition to circumanal setae, five pairs of lyrifissures and a pair of pores next to the margin of the shield, about in transverse line with Zv 5. Postanal seta about 1.2 times as long as para-anal seta. Unsclerotized cuticle around ventrianal shield with two pairs of opisthogastric setae (Jv 5, Zv 5) and a pair of lyrifissures. Lengths of ventral setae: st 1 25, st 2 22, st 3 20, st 4 15, st 5 16, Jv 1 15, Jv 2 12, Jv 3 13, Jv 5 16, Zv 1 15, Zv 2 12, Zv 3 10, Zv 5 10, para-anal 17, post-anal 20. Setae aciculate and smooth, except post-anal seta, barbed. Exopodal plates as in adult female. Peritreme and peritrematic shield. Peritremal shield fused to dorsal shield at level of r 3 and to exopodal shield at level of caxa IV; peritreme as in adult female. Legs. Pretarsi of legs I – IV, shape of setae and chaetotaxy as in adult female. Lengths of legs: I, 282; II, 237; III, 225; IV, 312. Setae aciculate. Leg chaetotaxy as in female.	en	da Silva, Renan V., Narita, João Paulo Z., Vichitbandha, Patchanee, Chandrapatya, Angsumarn, Konvipasruang, Ploychompoo, Kongchuensin, Manita, de Moraes, Gilberto J. (2013): Prospection for predatory mites to control coconut pest mites in Thailand, with taxonomic descriptions of collected Mesostigmata (Acari). Journal of Natural History (J. Nat. Hist.) 48 (11 - 12): 699-719, DOI: 10.1080/00222933.2013.839842, URL: http://dx.doi.org/10.1080/00222933.2013.839842
EA3E7D706C0EE264FE1CB9D5FE6EFBB8.taxon	materials_examined	Material examined Female holotype, two female and one male paratypes, on coconut palm leaves, Cocos nucifera L. (Arecaceae), Don Sak, Thailand. All collected in March 2011, by Gilberto J. de Moraes. Holotype female, paratype male and paratype female deposited in the Insect Museum, Department of Entomology, Kasetsart University, Bangkok, Thailand. Paratype female deposited in the Acarology Collection of Departamento de Entomologia e Acarologia, Escola Superior de Agricultura “ Luiz de Queiroz ”, Universidade de São Paulo, Piracicaba, São Paulo, Brazil.	en	da Silva, Renan V., Narita, João Paulo Z., Vichitbandha, Patchanee, Chandrapatya, Angsumarn, Konvipasruang, Ploychompoo, Kongchuensin, Manita, de Moraes, Gilberto J. (2013): Prospection for predatory mites to control coconut pest mites in Thailand, with taxonomic descriptions of collected Mesostigmata (Acari). Journal of Natural History (J. Nat. Hist.) 48 (11 - 12): 699-719, DOI: 10.1080/00222933.2013.839842, URL: http://dx.doi.org/10.1080/00222933.2013.839842
EA3E7D706C0EE264FE1CB9D5FE6EFBB8.taxon	etymology	Etymology The name of the species was given after the former name (Siam) of the country where the types were collected.	en	da Silva, Renan V., Narita, João Paulo Z., Vichitbandha, Patchanee, Chandrapatya, Angsumarn, Konvipasruang, Ploychompoo, Kongchuensin, Manita, de Moraes, Gilberto J. (2013): Prospection for predatory mites to control coconut pest mites in Thailand, with taxonomic descriptions of collected Mesostigmata (Acari). Journal of Natural History (J. Nat. Hist.) 48 (11 - 12): 699-719, DOI: 10.1080/00222933.2013.839842, URL: http://dx.doi.org/10.1080/00222933.2013.839842
EA3E7D706C0EE264FE1CB9D5FE6EFBB8.taxon	discussion	Remarks This new species is closely related to Lasioseius cuppa Walter and Lindquist, 1997. Female of L. siamensis sp. nov. differs from L. cuppa by having: 13 setae on tibia I (instead of 12; with two instead of three ventral setae); no platelets between genital and ventrianal shield (instead of with fragmented strip); pre-anal pores on the margin of the ventrianal shield, in transverse line with insertion of Jv 4 (instead of anterolaterad of and slightly anterior to insertion of Jv 4). Male of L. siamensis differs from L. cuppa by having pre-anal pores next to the margin of the shield, well posterior and laterad of insertion of Jv 3 (instead of away from the margin, slightly posterior and mesad of insertion of Jv 3).	en	da Silva, Renan V., Narita, João Paulo Z., Vichitbandha, Patchanee, Chandrapatya, Angsumarn, Konvipasruang, Ploychompoo, Kongchuensin, Manita, de Moraes, Gilberto J. (2013): Prospection for predatory mites to control coconut pest mites in Thailand, with taxonomic descriptions of collected Mesostigmata (Acari). Journal of Natural History (J. Nat. Hist.) 48 (11 - 12): 699-719, DOI: 10.1080/00222933.2013.839842, URL: http://dx.doi.org/10.1080/00222933.2013.839842
EA3E7D706C05E278FE16BC7EFE7EFB98.taxon	description	Adult female (five specimens measured) (Figure 3) Gnathosoma (Figure 3 A – C). Fixed cheliceral digit 30 (27 – 31) long, with eight teeth; movable cheliceral digit 26 (23 – 29) long, with two teeth near the apex, in addition to apical tooth. Anterior margin of epistome denticulate, with two aciculate projections. Palp-apotele three-tined. Deutosternum narrow, with seven transverse rows of denticles, anterior six rows with two to eight denticles, and the posterior row with 17 denticles. Corniculi two-tined, sub-parallel to each other. Lengths of hypostomal setae: h 1 19 (18 – 20), h 2 22 (20 – 23), h 3 24 (22 – 25), sc 19. Dorsum of idiosoma (Figure 3 D – F). Holodorsal shield 485 (470 – 512) long and 280 (270 – 290) wide at level of s 6, reticulate; reticula formed by simple lines. Podonotal region of dorsal shield with 17 pairs of setae (j 1 – j 6, z 2 – z 5, s 1, s 3, s 4, s 6, r 3 – r 5), three pairs of lyrifissures and three pairs of pores. Opisthonotal region of dorsal shield with 12 pairs of setae (J 2, J 4, Z 1 – Z 5, S 1 – S 5), five pairs of lyrifissures and one pair of pores. Lengths of dorsal setae: j 1 9, j 2 16 (15 – 17), j 3 18 (17 – 19), j 4 19 (16 – 22), j 5 18 (17 – 19), j 6 19, J 2 18 (17 – 19), J 4 22 (20 – 23), z 2 19 (17 – 20), z 3 18 (16 – 23), z 4 21 (19 – 22), z 5 17 (15 – 18), Z 1 20, Z 2 19 (17 – 20), Z 3 21 (20 – 22), Z 4 23 (20 – 25), Z 5 48 (45 – 50), s 1 18 (16 – 20), s 3 20, s 4 21 (20 – 22), s 6 22 (20 – 25), S 1 20 (18 – 22), S 2 20, S 3 21, S 4 21 (20 – 22), S 5 23 (20 – 25), r 3 18 (17 – 20), r 4 20 (18 – 23), r 5 18 (16 – 20). Setae J 4 and Z 4 stout and fringed distally; Z 5 stout and barbed distally; other dorsal shield setae setiform and smooth. Venter of idiosoma (Figure 3 G). Base of tritosternum 16 (15 – 18) long and 10 wide; laciniae fused along about 50 % of their total length (45 long). Sternal shield 127 long at midline and 129 (125 – 130) wide at widest level, with scanty lateral reticulation, centrally smooth, with a triangular posterior projection, bearing st 1 – st 3 and two pairs of lyrifissures, seta st 4 and iv 3 inserted on an elongate and relatively wide plate that might represent the fused metasternal and endopodal plates. Genital shield 75 wide at widest level, reticulate, truncate posteriorly, bearing genital seta (st 5); paragenital lyrifissure on unsclerotized cuticle, posterolaterad of st 5. With a pair of centrally punctuate, rounded metapodal plates. Ventrianal shield subrectangular, with transverse striae connected by few diagonal lines, 156 (140 – 160) long at midline and 171 (160 – 175) wide at widest level, with three pairs of opisthogastric setae (Jv 1, Zv 1 and Zv 2) in addition to circumanal setae. Unsclerotized cuticle of opisthogaster with one pair of setae (Jv 5) and two pairs of lyrifissures. Lengths of ventral setae: st 1 15 (13 – 17), st 2 16 (14 – 17), st 3 17 (15 – 19), st 4 13 (10 – 15), st 5 15 (12 – 18), Jv 1 11, Jv 5 14 (12 – 15), Zv 1 11 (10 – 12), Zv 2 10. All ventral setae setiform and smooth. Peritreme and peritrematic shield (Figure 3 D – G). Peritrematic shield fused to dorsal shield at an undetermined level, and to exopodal shield immediately behind stigma; peritreme reaching level of j 1. Spermatheca (Figure 3 H). Induction pore apparently at postero-proximal region of coxa III; tubulus (70 long) apparently leading to the sacculus. Legs. Pretarsi of legs I – IV consisting of elongate stalk, a pair of claws and pulvillus with three rounded lobes. Lengths of legs: I, 320, II, 285 (280 – 290), III, 278 (275 – 280), IV, 335 (325 – 345). Chaetotaxy of legs I – IV: coxa 2, 2, 2, 1; trochanter 6 (1 - 0 / 1, 1 / 2 - 1), 5 (1 - 0 / 1, 0 / 2 - 1), 5 (1 - 1 / 1, 0 / 2 - 0), 5 (1 - 1 / 1, 0 / 2 - 0); femur 10 (2 - 2 / 2, 1 / 2 - 1), 9 (1 - 2 / 1, 2 / 2 - 1), 6 (1 - 2 / 1, 1 / 0 - 1), 6 (1 - 2 / 1, 1 / 0 - 1); genu 11 (2 - 3 / 1, 2 / 1 - 2), 9 (2 - 3 / 0, 2 / 0 - 2), 7 (2 - 2 / 0, 2 / 0 - 1), 9 (2 - 2 / 1, 3 / 0 - 1); tibia 11 (2 - 3 / 1, 2 / 1 - 2), 10 (2 - 2 / 1, 2 / 1 - 2), 8 (1 - 2 / 1, 2 / 1 - 1), 10 (2 - 1 / 1, 3 / 1 - 2); tarsi not counted, 18, 18, 18. Dorsal setae of femora barbed; posterolaterad seta and one of the posterodorsal seta of genua lightly barbed; other leg setae aciculate and smooth. Adult male (three specimens measured) (Figure 4) Gnathosoma (Figure 4 A). Fixed cheliceral digit 30 long, with eight teeth in addition to apical tooth; movable cheliceral digit 23 long, with a tooth near the apex, in addition spermatodactyl 35 long, slightly and homogeneously curved along its extension. Epistome, apotele, deutosternum and corniculi similar to adult female. Lengths of hypostomal and palp-coxal setae: h 1 19, h 2 21, h 3 20 (19 – 21), sc 18 (17 – 20). Dorsum of idiosoma. Holodorsal shield 470 (460 – 480) long and 288 (283 – 290) wide at level of s 6, ornamented as in adult female. Podonotal region of dorsal shield with 17 pairs of setae (j 1 – j 6, z 2 – z 5, s 1, s 3, s 4, s 6, r 3 – r 5), three pairs of lyrifissures and three pairs of pores. Opisthonotal region of dorsal shield with 12 pairs of setae (J 2, J 4, Z 1 – Z 5, S 1 – S 5), five pairs of lyrifissures and a pair of pores. Lengths of dorsal setae: j 1 9, j 2 17 (16 – 18), j 3 19 (18 – 20), j 4 18, j 5 19 (18 – 20), j 6 20, J 2 20, J 4 21, z 2 17 (16 – 18), z 3 16, z 4 21, z 5 16 (15 – 17), Z 1 21, Z 2 21, Z 3 21, Z 4 19 (18 – 20), Z 5 46 (45 – 47), s 1 17 (16 – 18), s 3 22, s 4 22, s 6 21, S 1 20, S 2 21, S 3 21, S 4 21, S 5 23, r 3 19 (17 – 20), r 4 20, r 5 16. Seta Z 5 stout and fringed distally; other dorsal shield setae setiform and smooth. Venter of idiosoma (Figure 4 B). Base of tritosternum 14 long and 10 wide; laciniae fused along about 50 % of their total length (45). Sternogenital shield 242 (235 – 250) long and 118 (110 – 120) wide at level between st 2 and st 3, reticulate anteriorly and scantly reticulate posteriorly, smooth centrally, bearing st 1 – st 5 and three pairs of lyrifissures; seta st 4 inserted on a lateral strip along coxa III, connected anteriorly to the main body of the sternogenital shield. Paragenital lyrifissure on unsclerotized cuticle posterolaterad of st 5. With a pair of round, smooth metapodal plates. Ventrianal shield subcircular, with transverse striae connected by few diagonal lines, 137 (135 – 143) long at midline and 185 (180 – 187) wide at widest level, bearing Jv 1, Zv 1 and Zv 2. Unsclerotized cuticle around ventrianal shield with a pair of setae (Jv 5). Lengths of ventral setae: st 1 11 (10 – 13), st 2 16 (15 – 17), st 3 16, st 4 14 (13 – 15), st 5 15 (14 – 16), Jv 1 10, Jv 5 12, Zv 1 11, Zv 2 11. Shape of ventral setae similar to female. Peritreme and peritrematic shield. Similar to adult female. Legs. Pretarsi, chaetotaxy and type of setae as in adult female.	en	da Silva, Renan V., Narita, João Paulo Z., Vichitbandha, Patchanee, Chandrapatya, Angsumarn, Konvipasruang, Ploychompoo, Kongchuensin, Manita, de Moraes, Gilberto J. (2013): Prospection for predatory mites to control coconut pest mites in Thailand, with taxonomic descriptions of collected Mesostigmata (Acari). Journal of Natural History (J. Nat. Hist.) 48 (11 - 12): 699-719, DOI: 10.1080/00222933.2013.839842, URL: http://dx.doi.org/10.1080/00222933.2013.839842
EA3E7D706C05E278FE16BC7EFE7EFB98.taxon	discussion	In the original description, Womersley (1956) reported all four pairs of opisthogastric setae of this species to be on the ventrianal shield. Similarly to what was found in this study, Halliday (1997) observed that the female holotype of B. leveri has four pairs of opisthogastric setae, three pairs of which on the ventrianal shield. Halliday (1997) indicated that this species can be distinguished from B. marianneae by having the sternal shield with a triangular posterior process (slightly convex in B. marianneae), dorsal shield much less ornate and dorsal shield setae short, except for Z 5. Other characteristics that can be used to separate these species are the denticulate anterior margin of the epistome, which also has two aciculate projections in B. leveri (smooth and with a single and acute anterior projection in B. marianneae) and ventrianal shield with transverse striae connected by few diagonal lines in B. leveri (distinctly reticulate in B. marianneae).	en	da Silva, Renan V., Narita, João Paulo Z., Vichitbandha, Patchanee, Chandrapatya, Angsumarn, Konvipasruang, Ploychompoo, Kongchuensin, Manita, de Moraes, Gilberto J. (2013): Prospection for predatory mites to control coconut pest mites in Thailand, with taxonomic descriptions of collected Mesostigmata (Acari). Journal of Natural History (J. Nat. Hist.) 48 (11 - 12): 699-719, DOI: 10.1080/00222933.2013.839842, URL: http://dx.doi.org/10.1080/00222933.2013.839842
EA3E7D706C19E27AFEC9BC5FFCFEF91E.taxon	description	Adult male (three specimens measured) (Figure 5) Gnathosoma (Figure 5 A). Fixed cheliceral digit 25 long, with two large basal teeth and a small subapical tooth, and with an inflated structure tapered distally; movable cheliceral digit 23 long, edentate. Spermatodactyl 50 long, slightly curved, of about uniform diameter and with a sharp distal process. Lengths of hypostomal and palp – coxal setae: h 1 23, h 2 20, h 3 25, sc 20. Dorsum of idiosoma (Figure 5 B). Holodorsal shield 395 long and 240 wide at level of s 6, reticulate laterally and smooth centrally, between j 3 and J 4. Podonotal region of dorsal shield with 19 pairs of setae (j 1 – j 6, z 2, z 4 – z 6, s 1, s 2, s 4 – s 6, r 2 – r 5), with four pairs of lyrifissures and four pairs of pores. Opisthonotal region of dorsal shield with 10 pairs of setae (J 2, J 4, Z 1, Z 2, Z 4, Z 5, S 2 – S 5), with nine pairs of lyrifissures and three pairs of pores. Lengths of dorsal setae: j 1 32 (30 – 35), j 2 18 (15 – 25), j 3 17 (15 – 20), j 4 15, j 5 13, j 6 15, J 2 49 (40 – 63), J 4 13 (10 – 15), z 2 17 (15 – 20), z 4 17 (15 – 20), z 5 17 (15 – 18), z 6 17 (15 – 18), Z 1 18 (15 – 20), Z 2 37 (30 – 48), Z 4 34 (25 – 45), Z 5 35 (30 – 40), s 1 23 (20 – 30), s 2 23 (20 – 30), s 4 37 (30 – 50), s 5 37 (30 – 50), s 6 37 (30 – 50), S 2 40 (30 – 50), S 3 44 (35 – 58), S 4 45 (35 – 60), S 5 47 (40 – 60), r 2 42 (35 – 55), r 3 38 (30 – 50), r 4 40 (30 – 55), r 5 43 (30 – 60). Setae j 1 leaf-shaped and barbed; setae J 2 stout and barbed; setae Z 2, Z 4, Z 5, s 4 – s 6, S 2 - S 5, r 2 – r 5 stout, barbed and broader distally; other dorsal setae setiform and smooth. In five of nine males of N. ceylonicus examined in this study seta J 4 was inserted posterolaterad of seta J 2, as in the illustration; in the remaining, J 4 was inserted anterior of J 2. Stridulating organ on unsclerotized cuticle at level between s 6 and S 3, with a large anterior lobe followed by a series of small lobes. Venter of idiosoma (Figure 5 C). Base of tritosternum 7 long and 14 wide; laciniae fused along about a third of their total length (60 long). Sternogenital shield 193 (185 – 200) long and 87 (85 – 90) wide at level of st 3, bearing five pairs of setae and three pairs of lyrifissures, with two pairs of rounded metapodal plates. Anal shield oval, smooth, 105 (100 – 110) long at midline and 125 (120 – 140) wide at widest level, unsclerotized cuticle around anal shield with six pairs of setae (Jv 1 - Jv 5, Zv 2). Lengths of ventral setae: st 1 22 (20 – 25), st 2 22 (20 – 25), st 3 21 (20 – 22), st 4 19 (17 – 20), st 5 22 (18 – 27), Jv 1 18 (15 – 20), Jv 2 17 (15 – 18), Jv 3 15, Jv 4 40 (35 – 50), Jv 5 52 (45 – 65), Zv 2 17 (15 – 20). Setae Jv 4 and Jv 5 stout and barbed; other ventral setae setiform and smooth. Endopodal plate represented by an arched platelet at level between coxae II and III, and an elongate platelet at level between coxae III and IV. Peritreme and peritrematic shield (Figure 5 C, D). Peritrematic shield fused to dorsal shield at level between j 2 and j 3, and to exopodal shield immediately behind stigma; with a large pore at level between coxa III and IV (gd 3 of Athias-Henriot 1975; gp 2 of Lindquist and Moraza 2009); peritreme reaching the level of j 1. Legs (Figure 5 D). Pretarsi of legs I – IV consisting of elongate stalk, a pair of claws and pulvillus with three rounded lobes. Lengths of the legs: I 350, II 300, III 305 (300 – 310), IV 390 (370 – 410). Chaetotaxy of legs I – IV: coxa 2, 2, 2, 1; trochanter 6 (1 - 0 / 2, 1 / 1 - 1), 5 (1 - 1 / 2, 0 / 1 - 1), 5 (1 - 1 / 2, 0 / 1 - 0), 5 (1 - 0 / 2, 1 / 1 - 0); femur 12 (2 - 3 / 1 - 2 / 2 - 2), 10 (2 - 2 / 1,2 / 2 - 1), 6 (1 - 2 / 1, 1 / 0 - 1), 6 (1 - 2 / 1, 1 / 0 - 1); genua 12 (2 - 3 / 2, 2 / 1 - 2), 11 (2 - 3 / 1, 2 / 1 - 2), 9 (2 - 2 / 1, 2 / 1 - 1), 9 (2 - 2 / 1, 3 / 0 - 1); tibiae 12 (2 - 3 / 2, 2 / 1 - 2), 10 (2 - 2 / 1, 2 / 1 - 2), 8 (2 - 1 / 1, 2 / 1 - 1), 9 (2 - 2 / 1, 2 / 1 - 1); tarsi not counted, 17, 17, 17. With one to four lobe-like structures on the dorsal surface of femur, genu and tibia IV. Each of femur and genu IV with a posterodistal spur probably used to rub stridulating organ.	en	da Silva, Renan V., Narita, João Paulo Z., Vichitbandha, Patchanee, Chandrapatya, Angsumarn, Konvipasruang, Ploychompoo, Kongchuensin, Manita, de Moraes, Gilberto J. (2013): Prospection for predatory mites to control coconut pest mites in Thailand, with taxonomic descriptions of collected Mesostigmata (Acari). Journal of Natural History (J. Nat. Hist.) 48 (11 - 12): 699-719, DOI: 10.1080/00222933.2013.839842, URL: http://dx.doi.org/10.1080/00222933.2013.839842
EA3E7D706C19E27AFEC9BC5FFCFEF91E.taxon	discussion	A detailed account of the morphology of the female was presented in the original description of this species; hence, this sex is not re-described in this paper. Narita et al. (2011) distinguished N. ceylonicus from N. stridulans (Evans 1955) based on the sternal shield, wider than long in N. ceylonicus. Other characteristics that separate the females of these species are the presence of a tooth near the apex of the fixed cheliceral digit of N. ceylonicus (absent in N. stridulans) and tritosternal base wider than long in N. ceylonicus (longer than wide in N. stridulans).	en	da Silva, Renan V., Narita, João Paulo Z., Vichitbandha, Patchanee, Chandrapatya, Angsumarn, Konvipasruang, Ploychompoo, Kongchuensin, Manita, de Moraes, Gilberto J. (2013): Prospection for predatory mites to control coconut pest mites in Thailand, with taxonomic descriptions of collected Mesostigmata (Acari). Journal of Natural History (J. Nat. Hist.) 48 (11 - 12): 699-719, DOI: 10.1080/00222933.2013.839842, URL: http://dx.doi.org/10.1080/00222933.2013.839842
