taxonID	type	description	language	source
1F364B64B67CB915FF38FE12FC22F9AB.taxon	description	(Figs. 2 – 7)	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B67CB915FF38FE12FC22F9AB.taxon	materials_examined	Type locality. Khao Plu cave in Wat [= temple] Tham Khao Plu located in Chumko Subdistrict, Pathiu District, Chumphon Province, southern Thailand (Fig. 1 A, 1 B). The coordinates of the entrance are 10 ° 43 ′ 49.06 ′ N, 99 ° 19 ′ 12.52 ′ E; altitude 27 m a. s. l. The cave is about 50 m long and situated on a small hill, with a single pool filled exclusively with dripping water in the dark zone. The type locality was made by a mixture of stone and cement in order to collect the dripping water for observance. The water temperature during sampling was 24.0 ° C, pH 7.8, and conductivity was 310 μS cm− 1. Material examined. Holotype: an adult female was dissected and mounted on a slide in glycerol and sealed with nail polish, NHMUK 2017.503. Allotype: an adult male was dissected and mounted on a slide in glycerol and sealed with nail polish, NHMUK 2017.504. Paratypes: three undissected females (one female with egg sacs) and males were preserved by 70 % ethanol in a 1.5 mL microtube, NHMUK 2017.505 – 510; three undissected females and males were preserved by 70 % ethanol in a 1.5 mL microtube, NPU 2017.002. Additional materials: four undissected females and males were mounted on SEM Stub (NPU 2017.002 S). All specimens were collected from Khao Plu cave by author on 25 November 2016.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B67CB915FF38FE12FC22F9AB.taxon	diagnosis	Diagnosis. Adult female: anal operculum is ovate, and serrated. P 1 – P 4 show acute projection on distal margins, without inner coxal seta, P 1 has an inner basal spine. P 1 – P 3 have two-segmented Enp and Exp, P 4 has one-segmented Enp and two-segmented Exp. Setal and spine formula are present on swimming legs: Exp- 2 P 1 – P 4 as 5.5.5.4 and 3.3.3.3, respectively; Enp- 2 P 1 – P 3 as 3.3.5 and 1.1.1, respectively. Enp P 4 has four setae and one spine, without spiniform process on the segment. Adult male: P 1, P 2, Exp P 3 - P 4 are the same as in females. Enp- 2 P 3 has three pinnate and one smooth seta. The transformed spine has a well-produced, outstretched part tapering distally and located slightly above the middle of the spine. P 4 Enp is two-segmented, and Enp- 1 has one inner pinnate seta and one spiniform process on the inner distal margin; Enp- 2 has three pinnate setae and one spiniform spine, without a spiniform process.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B67CB915FF38FE12FC22F9AB.taxon	description	Description. Female (Figs. 2, 3), body length: 550 – 625 μm (mean: 610 μm; n = 10), preserved specimens are colourless. Naupliar eye not discernible. Cephalosome wider than rest of body, with transverse scar indicating the fusion region of cephalosome and pediger 1 (Figs. 2 A, 3 A, 3 B). Cephalosome, pedigers 2 – 5, genital double-somite, urosomites and anal somite (incl. anal operculum) covered with fine refractile points. Posterior dorsal margins of cephalosome and pedigers 2 – 4 with smooth hyaline membrane, and with a serrated transverse row of dorsal hyaline frills above its posterior margin; pediger 5 with small serrated hyaline frill. Genital double-somite and urosomite 2 – 3 with irregularly serrate hyaline membrane on entire posterior margins; with a transverse row of small grooves on dorso-lateral surface (Figs. 2 A, 3 C, 3 D). Genital double-somite symmetrical, about 1.5 times wide as long; with a pair of dorsal sclerotised; single copulatory pore situated ventrally near middle of segment. Anal somite (Figs. 2 A – C, 3 C) with strong spinules along entire posterior margin. Anal operculum (Figs. 2 C, 3 C) ovate, coarsely serrated along its free margin, reaching beyond the middle of caudal ramus length, with a pair of small sensillum above the base. Caudal rami (Figs. 2 A, 3 C) symmetrical sub-rectangular, about 1.3 times long as wide. Dorsal keel long, orientated nearly the entire caudal ramus length. Seta I absent, seta II-V pinnate. Lateral seta (II) shorter than the caudal ramus, inserted at about mid-length of caudal ramus. Outermost terminal seta (III) longer than dorsal seta, about 1.5 times as long as caudal ramus with strong spinules near its insertion on lateral and ventral views. Seta IV and V with fracture plane: outer terminal seta (IV) and inner terminal seta (V) about 4.0 and 6.0 times as long as caudal ramus, respectively. Innermost terminal seta (VI) thin and bare, shorter than 1 / 2 of caudal ramus. Dorsal seta (VII) articulated and bare, about 1.2 times as long as caudal ramus, inserted on distal end of keel with tiny spinules at insertion. Rostrum (Fig. 4 A) fused to cephalosome, free margin round, unornamented. Antennule (Figs. 3 A, 4 B) short, reaching about 1 / 2 of cephalosome, 11 - segmented. Setal formula on each segment as follows (A = aesthetasc): 6.2.5.2.0.2.3.1 + A. 2.2.7 + A. All setae smooth. Antenna (Fig. 4 C) four-segmented, comprising coxobasis and three-segmented Enp. Coxobasis with bare seta on distal inner corner. Enp- 1 with smooth seta on distal inner margin plus row of spinules on outer margin. Enp- 2 with five smooth setae: three lateral and two apical setae. Enp- 3 with seven smooth setae on distal end: one inner, four apical and two outer setae. Mandible (Fig. 4 D) short and robust, with five strongly chitinised teeth on gnathobase plus a smooth seta on innermost margin. Mandibular palp reduced, with one bare seta. Maxillule (Fig. 4 E) with praecoxa, coxobasis and onesegmented Enp. Praecoxal arthrite with four strong apical claws fused to arthrite base, with one strong spine, four short and smooth setae and one spinulate seta on frontal side. Coxobasis with one spiniform and two smooth setae. Exp reduced into one smooth seta. Enp with three smooth setae. Maxilla (Fig. 4 F) with praecoxa, coxa, basis, and twosegmented Enp. Praecoxal endite with two apical setae. Coxa with one seta at near middle of inner margin and two setae on distal endite. Basis with two claw-like expansions plus one seta at insertion. Enp- 1 with one seta and Enp- 2 with three thin setae and one robust seta. Maxilliped (Fig. 4 G) syncoxa and basis with two and one spiniform setae, respectively, each with a row of strong spinules on inner margin. Two-segmented Enp, Enp- 1 with spiniform seta. Enp- 2 reduced, with two smooth setae. Armament of legs 1 – 4 as follows (legend: outer-inner seta / spine; outer-apical-inner; Arabic numerals indicate setae; Roman numerals indicate spines): P 1 (Fig. 5 A) intercoxal plate with acute projections on distal margin. Coxa without seta on inner corner. Basis with bare, slender outer seta and robust, stout inner spine with a cluster of spinules at spine insertion. Two-segmented Exp and Enp. Exp- 1 smaller than Exp- 2, with outer spiniform spine. Exp- 2 twice as long as wide with three outer spiniform spines, two apical pinnate setae and three inner pinnate setae. Enp- 1 with one inner pinnate seta with spiniform process at distal outer corner. Enp- 2 with three subequal pinnate setae and one spiniform spine; one outer seta located between two spiniform processes, one apical spine and one apical seta, and one inner seta. P 2 (Fig. 5 B) intercoxal plate, coxa, basis, and Exp similar to P 1, but basis without spine on inner margin. Two-segmented Enp, and Enp- 1 slightly smaller than Enp- 2. Enp- 1 with one inner pinnate seta and spiniform process at distal outer corner. Enp- 2 with four subequal elements: one apical spiniform spine and three pinnate setae (outer seta located between two spiniform processes, apical seta and inner seta). P 3 (Fig. 5 C) intercoxal plate, coxa, basis, Exp, and Enp- 1 similar to P 2. Enp- 2 with one apical spine and five pinnate setae: shortest outer seta located between two spiniform processes, three inner setae about as long as spine and longest apical seta. P 4 (Fig. 5 D) intercoxal plate with acute projections on distal margin. Coxa without inner seta. Basis with slender outer seta. Two-segmented Exp, Exp- 1 with outer spiniform spine and Exp- 2 with three outer spiniform spines, two apical pinnate setae and two inner pinnate setae. One-segmented Enp, with one spine and four pinnate setae; outer seta about as long as two inner setae, apical one longest, about 3.0 times as long as spine. P 5 (Figs. 2 C, 2 D, 3 E) completely fused to somite, with three pinnate setae: outermost seta longest, inserted on small prominence; middle and innermost seta directly inserted on somite, former seta shorter than latter one. P 6 (Figs. 2 C, 3 D) reduced to simple plate on genital doublesomite laterodorsally, with two minute spinules and short pinnate seta. Adult females each have a pair of egg sacs, each with three large eggs. Male (Figs. 3 F – I, 6) smaller than female. Body length, excluding caudal setae, 460 – 550 μm (mean: 530 μm; n = 10). Body shape similar to female, except in genital segment and urosomal segmentation (i. e. the males have one more urosomite). Antenna and mouthparts (not figured), caudal rami (Fig. 6), P 1 (Fig. 7 A), P 2 (Fig. 7 B), P 3 – P 4 exopods (Figs. 7 C, 7 D) and P 5 (Fig. 6 D) similar to those in female. Anal operculum (Figs. 3 G, 6 B, 6 D) slightly shorter than in female, about mid-length of caudal ramus. Antennule (Fig. 4 H) 15 - segmented, geniculate. Setal formula as follows (A = aesthetasc, S = spine): 8 + 2 A. 3.2.2 + A. 1.2.2.1 + A. 2.2. S. 1 + A. S. 1.8 + A. All setae smooth. P 3 (Fig. 7 C) two-segmented Enp, proximal segment smaller than distal one. Enp- 1 with inner pinnate seta and spiniform process on distal outer margin. Enp- 2 with small outer pinnate seta between two spiniform processes, apical transformed spine and apical bare seta and two inner pinnate setae. Transformed spine with outstretched part tapering distally and well-produced, located slightly above the middle of spine; with several strong spinules on its outstretched part. P 4 (Fig. 7 D) two-segmented Enp. Enp- 1 with inner pinnate seta and spiniform process on inner distal margin. Enp- 2 with one outer pinnate seta, one apical spine and pinnate seta, and one inner pinnate seta. P 6 (Figs. 3 I, 6 C) reduced to simple plate, with two equal pinnate setae and one longer pinnate seta. Variability. Not observed.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B67CB915FF38FE12FC22F9AB.taxon	etymology	Etymology. The specific name muscicoloides, formed with the Latin suffix “ - oides ” for resembling, refers to the similarity of the new species with B. muscicola. The species epitheton is a masculine singular adjective.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B67CB915FF38FE12FC22F9AB.taxon	discussion	Remarks. Bryocyclops muscicoloides, new species, fits to the generic characters as the P 5 is completely fused to the fifth thoracic somite, with three setae, P 2 – 4 have no inner coxal seta, the intercoxal plate of P 4 has acute protuberances on the distal margin, and the male P 3 Enp- 2 has a transformed spine. The new species is similar to the members of Group II sensu Lindberg (1954) (except that P 1 coxa has no inner seta in B. muscicoloides, new species) in the following characters: (1) sexual dimorphism occurs on P 3 Enp- 2 and P 4 Enp- 2, (2) P 1 basis has an inner spine, (3) intercoxal plates of P 1 – P 4 have acute protuberances on the distal margin, (4) the setal and spine formulae of P 1 – P 4 Exp- 2 are 5.5.5.4. and 3.3.3.3, respectively and (5) the P 4 of female has two-segmented Exp and one-segmented Enp, and there are two-segmented Exp and Enp in the male (Lindberg, 1954; Watiroyram et al., 2015 a). Currently, Group II consists of five species: B. bogoriensis (Menzel, 1926), B. muscicola (Menzel, 1926), B. fidjiensis Lindberg, 1953, B. caroli Bjornberg, 1985, and B. campaneri Rocha & Bjornberg, 1987 (Reid, 1999). The new species is most similar to B. muscicola from Indonesia (Menzel, 1926; Bjornberg, 1985; Rocha & Bjornberg, 1987; Watiroyram et al., 2015 a). Apart from the group characteristics as previously mentioned, they share some affinities in the posterior margin of the urosome and anal operculum serrated, the female Enp P 4 has four setae and one spine; the setal formulae on Enp- 2 P 1 and P 3 in female have four and six elements, respectively, and Enp- 2 P 1 of males has four elements. Bryocyclops muscicoloides, new species, differs markedly from B. muscicola in the following characteristics: in both sexes, P 1 has no inner coxal seta (present in B. muscicola) and Exp- 2 P 2 – P 3 have normal setae (each with two blunt setae in B. muscicola); in males, the transformed spine on Enp- 2 P 3 shows the outstretched part gradually tapering to the tip, which is located above the middle of its length (semi-circular and located at about halfway along the length in B. muscicola); in female, Enp P 4 has no spiniform process on the outer margin (present in B. muscicola). Regarding species descriptions by Reid (1999), the other differences between the species are (a) the prosome and urosome have refractile points including anal operculum (only prosome in female and included urosomite 2 in male with respective characters in B. muscicola), (b) caudal ramus has spinules at the insertion of dorsal seta, and (c) caudal ramus of male has dorsal keel (both absent in B. muscicola). Bryocyclops muscicoloides, new species, differs from B. bogoriensis in that the female of the new species has three setae on P 2 Enp- 2 but there are five setae in B. bogoriensis; P 4 Enp has no spiniform process on the outer margin of B. muscicoloides, new species, whereas it is present in B. bogoriensis; and apical seta are longest in the new species but are almost as long as the distal inner seta in B. bogoriensis. The transformed spine on the male P 3 Enp- 2 has an outstretched part gradually tapering to the distal end, but it is rounded in B. bogoriensis. The new species has a serrated and rounded anal operculum, but this is triangular and smooth in B. fidjiensis. Bryocyclops muscicoloides, new species’ P 4 Exp and Enp are two- and one-segmented in female and two-segmented in male, but are one- and two-segmented in both sexes of B. fidjiensis. There are three spines on the distal exopod of P 4 in the new species but four spines on the same segment in B. fidjiensis. Bryocyclops muscicoloides, new species, can be distinguished from B. caroli by P 1 of the new species: it has no coxal inner seta, which is present in the later species. The anal operculum is regularly serrated in the new species compared to those in B. caroli. The shape of the female P 4 Enp diferentiates the new species from B. caroli by lack of a spiniform process on the outer margin and the transformed spine of male P 3 Enp- 2 has well-developed outstretched part and acute tip in the new species, while those spine in B. caroli has lessdeveloped outstretched part and blunt tip. Bryocyclops muscicoloides, new species, is different from B. campaneri in that the new species has no inner coxal seta on P 1, which is present in B. campaneri; the P 1 Exp- 2 of the new species has five setae compared to four for B. campaneri; the female P 4 Enp of the new species has no any spiniform process and the apical seta is typically longer than the apical spine while those segment in B. campaneri has outer spiniform process and apical seta is slightly longer than the spine; and the transformed spine on the male P 3 Enp- 2 of the new species has an outstretched part tapering to its tip compared to a semi-circular shape for B. campaneri. The new species is easily distinguished from Group VII, proposed by a single species (B. maholarnensis Watiroyram, Brancelj & Sanoamuang, 2015) from Thailand, in that the distal Enp of P 4 in B. muscicoloides, new species, has four setae and one spine, as in other members of Group II; however, only two setae are present in B. maholarnensis and P 3 Enp- 2 of male in B. muscicoloides, new species, have remarkable transformed spine which is normally found in Group II and other Bryocyclops species but is absent in B. maholarnensis. The new species also differs from B. maewaensis Watiroyram, Brancelj & Sanoamuang, 2012, found in Thailand, by the new species lacking inner coxal seta on P 1, P 2 Enp- 2 of the new species having three setae (compared to four setae in B. maewaensis), the female P 4 Enp being one-segmented and having four setae and one spine in the new species (compared to B. maewaensis being two-segmented and having three setae and one spine on the distal Enp), the male P 3 Enp- 2 of the new species having a longer distance between its tip and the outstretched part in the new species being smaller and differently shaped to B. maewaensis.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B674B90CFC69F9F3FD51F92B.taxon	description	(Figs. 8 – 13)	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B674B90CFC69F9F3FD51F92B.taxon	materials_examined	Type locality. Khao Pina cave in Wat [= temple] Tham Khao Pina located in Nawong Subdistrict, Huai Yot District, Trang Province, southern Thailand (Figs. 1 A, 1 C). The coordinates of the entrance are 07 ° 44 ′ 55.30 ′ N, 99 ° 31 ′ 36.81 ′ E; altitude 34 m a. s. l. The cave is about 150 m long, with several pools filled exclusively with water dripping from stalactites. A rimstone pool in the dark zone, with about 10 – 20 L was selected as the location type. The water temperature during sampling was 22.0 ° C, pH 8.0, and conductivity was 350 μS cm− 1. Material examined. Holotype: an adult female was dissected and mounted on a slide in glycerol and sealed with nail polish, NHMUK 2017.511. Allotype: an adult male was dissected and mounted on a slide in glycerol and sealed with nail polish, NHMUK 2017.512. Paratypes: three undissected females and males were preserved by 70 % ethanol in a 1.5 mL microtube, NHMUK 2017.513 – 518; three undissected females with egg sacs and males were preserved by 70 % ethanol in a 1.5 mL microtube, NPU 2017.003. Additional materials: four undissected females and males were mounted on SEM Stub (NPU 2017.003 S). All specimens were collected from Khao Pina cave by the author on 27 November 2016.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B674B90CFC69F9F3FD51F92B.taxon	diagnosis	Diagnosis. Adult female: anal operculum is ovate and serrated. P 1 – P 4 have acute projections on the distal margins, P 1 has inner coxal seta, which are lacking on P 2 – P 4. P 1 has an inner basal spine. P 1 – P 3 have two-segmented Enp and Exp; P 4 has one-segmented Enp and two-segmented Exp. Setal and spine formulae on (1) Exp- 2 P 1 – P 4 as 5.5.5.4 and 3.3.3.3, respectively; (2) Enp- 2 P 1 – P 3 as 3.4.5 and 1.1.1, respectively; (3) Enp P 4 has four setae and one spine, with an outer spiniform process. Adult male: P 1, P 2, and Exp P 3 - P 4 are as in female. Enp- 2 P 3 has four pinnate setae and one strong transformed seta and spine. The transformed spine has a less-produced outstretched part, located near the tip of the spine. P 4 Enp is two-segmented; the Enp- 1 has one inner pinnate seta and one spiniform process on the outer distal corner plus two strong, short, spine-like processes along the inner margin; Enp- 2 has three pinnate setae and one spiniform spine, with a spiniform process on the outer distal corner.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B674B90CFC69F9F3FD51F92B.taxon	description	Description. Female (Figs. 8, 9), body length: 490 – 570 μm (mean: 550 μm; n = 10). Naupliar eye not discernible. Cephalothorax expanded laterally at cephalosome, with transverse scar on posterior end of cephalosome dorsally (Fig. 8 A). Cephalosome, pedigers 2 – 5, and genital double-somite covered with fine refractile points. Posterior dorsal margins of cephalosome and pedigers 2 – 4 with smooth hyaline membrane; pediger 5 with irregularly serrated hyaline membrane (Figs. 8 A, 8 B). Genital double-somite and two succeeding urosomites with irregularly serrated hyaline membrane on the entire posterior margin (Figs. 8 B – D, 9 A). Genital double-somite symmetrical, anterior part expanded laterally, about 1.4 times wide as long, with a pair of dorsal sclerotised; single copulatory pore situated mid-ventrally. Anal somite (Figs. 8 B – D, 9 A, 9 B) with a row of strong spinules on the entire posterior margin except nearly insertion point of anal operculum. Anal operculum (Figs. 8 B, 8 D, 9 B) ovate, small and irregularly serrated along its distal margin, not reaching the middle of caudal ramus, with a pair of dorsal sensilla on its base. Caudal rami (Figs. 8 B – D, 9 A, 9 B) symmetrical rectangular, about 2.3 times long as wide, with well-developed longitudinal dorsal keel, extended nearly the entire caudal ramus length. All seta pinnate, except seta VI; seta VII articulated at its base. Anterolateral accessory seta (I) absent. Anterolateral seta (II) shorter than caudal ramus, inserted near the middle of ramus dorsally. Posterolateral seta (III) about 1.5 times as long as caudal ramus, with strong spinules at its insertion laterally and ventrally. Outer terminal seta (IV) and inner terminal seta (V) with fracture plane, about 5.0 and 9.0 times as long as caudal ramus, respectively. Terminal accessory seta (VI) thin, shorter than 1 / 2 of caudal ramus. Dorsal seta (VII) about 2.0 times as long as caudal ramus, inserted on distal end of keel. Rostrum (Fig. 10 A) fused to cephalosome, free margin round and unornamented. Antennule (Figs. 10 B) short, not reaching posterior margin of cephalosome, 11 - segmented. Setal formula as follows (A = aesthetasc, S = spine): 7.2.5.2.0.2.3.1 + A. 2.2.7 + A. Antenna (Fig. 10 C) four-segmented, comprising coxobasis and three-segmented Enp. Coxobasis with smooth seta on inner distal corner. Enp- 1 with smooth seta on inner medial margin. Enp- 2 with five smooth setae along inner distal margin (one proximal and four distal setae) and row of tiny spinules along outer margin. Enp- 3 with seven smooth setae on distal end: one inner, four apical and two outer. Mandible (Fig. 10 D) short and robust, with four strongly chitinised teeth on gnathobase plus a smooth seta on innermost margin. Mandibular palp reduced, with one smooth seta. Maxillule (Fig. 10 E) with three-segmented. Praecoxal arthrite with two strong apical claws, one spine on distal margin and one spine, five setae on inner margin. Maxillular palp twosegmented, with four and three smooth setae on coxobasis and Enp, respectively. Maxilla (Fig. 10 F) with five-segmented. Praecoxal endite with two apical pinnate setae. Coxa with smooth seta on proximal endite and two smooth setae on distal endite. Basis with one apical claw-like expansion, one strong spine and one seta inserted at base of claw. Enp- 1 with one smooth inner seta. Enp- 2 with one strong and two thin apical setae. Maxilliped (Fig. 10 G) with four-segmented. Syncoxa with one spiniform and one smooth seta on distal inner margin accompanied by row of strong spinules along middle outer surface. Basis with one spiniform seta on middle inner margin and row of strong spinules on distal outer margin. Enp with two-segmented, with one spiniform seta and two smooth setae on proximal and distal segments, respectively. Armament of legs 1 – 4 as follows (legend: outer-inner seta / spine; outer-apical-inner; Arabic numerals indicate setae; Roman numerals indicate spines): P 1 (Fig. 11 A) intercoxal plate with acute projections on distal margin. Coxa with inner seta. Basis with slender seta on outer margin and stout spiniform spine on inner margin. Two-segmented Exp and Enp. Exp- 1 much smaller than Exp- 2, with one outer spine. Exp- 2 twice as long as wide, with three inner pinnate setae, two apical pinnate setae and three outer spines. Enp- 1 with one inner pinnate seta. Enp- 2 with one apical spiniform spine and three pinnate setae (one inner, one apical and one outer). P 2 (Fig. 11 B) intercoxal plate, and Exp similar to P 1. Coxa without inner seta. Basis with one slender outer seta. Twosegmented Enp, Enp- 1 with inner seta, Enp- 2 with one apical spine and four pinnate setae (two inner, one apical and one outer). P 3 (Fig. 11 C) intercoxal plate, coxa, basis and Exp similar to P 2. Two-segmented Enp, Enp- 1 with inner seta, Enp- 2 with one apical spine and five setae (three inner, one apical and one outer). All setae pinnate. P 4 (Fig. 11 D) intercoxal plate with acute projections on distal margin. Coxa without inner seta. Basis with slender seta on outer margin. Exp with two-segmented, Exp- 1 with outer spine. Exp- 2 with two inner pinnate setae, two apical pinnate setae and three outer spines. Enp with one-segmented, with spiniform process on outer margin, ornamented with apical spine and four pinnate setae (two inner, one apical and one outer). P 5 (Fig. 10 H) fused to fifth thoracic somite, with three pinnate setae: outermost seta longest, inserted on small prominence; middle and innermost setae directly inserted to somite, former seta shorter than latter. P 6 (Figs. 8 B, 8 D, 9 C) reduced to simple plate on genital double-somite laterodorsally, with lateral pinnate seta, middle minute spinule and inner naked seta. Adult females each have a pair of egg sacs, each with three eggs. Male (Figs. 12 – 13) slightly smaller than female. Body length, excluding caudal setae, 500 – 560 μm (mean = 530 μm; n = 5). Body shape similar to female, except in genital segment and urosomal segmentation (Fig. 12 A). Antenna, mouthparts, anal operculum, caudal rami (Figs. 9 E, 10 C – G), P 1 – P 2, P 5 (Figs. 12 C, 13 A, 13 B), P 3 – P 4 exopods (Figs. 13 C – D), similar to those in females. Antennule (Fig. 12 D) 15 - segmented. Setal formula as follows (A = aesthetasc, S = spine): 8 + A. 5.2.2 + A. 0.4.1.0.2.3. S. 1 + A. S. 1.8 + A. All setae smooth. P 3 (Fig. 13 C) Enp two-segmented, Enp- 1 smaller than Enp- 2. Enp- 1 with inner pinnate seta. Enp- 2 with one transformed spine apically, four pinnate setae (two inner, one apical, one outer) and one strong transformed seta on distal inner margin; transformed spine with outstretched part less-produced, located nearly its tip. P 4 (Fig. 13 D) Enp two-segmented. Enp- 1 bigger than Enp- 2, with inner pinnate seta and two short, strong spines on inner surface. Enp- 2 smaller, with inner seta, apical seta and spine and outer seta; all setae pinnate. P 6 (Figs. 9 F, 12 B, 12 C) reduced to simple plate, represented by three pinnate setae. Variability. P 1 Exp- 2 of male with six setae on one ramus (only one individual of ten examined males).	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B674B90CFC69F9F3FD51F92B.taxon	etymology	Etymology. The specific name trangensis refers to the place “ Trang ” where the new species was found, the Latin suffix “ - ensis ” is an adjective for the place.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
1F364B64B674B90CFC69F9F3FD51F92B.taxon	discussion	Remarks. Bryocyclops trangensis, new species, also fits with the generic and Group II characteristics as described for B. muscicoloides, new species. Bryocyclops trangensis, new species, is most similar to B. muscicola by the posterior margin of urosome and anal operculum serrated, the present of inner coxal seta on P 1, Enp- 2 P 1 having four elements, the female Enp- 2 P 3 having six elements, the shape and setation of the female Enp P 4 (five elements), and the male Enp- 2 P 4 of males having four elements. Bryocyclops trangensis, new species, can be easily distinguished from B. muscicola in the morphological differences in both sexes: Enp- 2 P 2 has five elements in B. trangensis, new species, but four elements in B. muscicola, and Exp- 2 P 2 – P 3 of B. trangensis, new species, have normal setae but those in B. muscicola have two blunt setae. The female P 4 Enp of B. trangensis, new species, has subequal three setae but B. muscicola has apical seta remarkably longer than the others. Bryocyclops trangensis, new species, has certain typical characteristics in the male P 3 – P 4: Enp- 1 P 4 has two strong, spine-like outgrowths on its inner surface, which are absent in B. muscicola; Enp- 2 P 3 of B. trangensis, new species, has four pinnate setae and one transformed seta while B. muscicola has one smooth and three pinnate setae; the outstretched part of the transformed spine on Enp- 2 P 3 is not well-produced in B. trangensis, new species, but is well-produced in B. muscicola. Bryocyclops trangensis, new species, differs from B. bogoriensis by a P 1 – P 4 intercoxal plate with an acute projections on distal margin that is more rounded in B. bogoriensis and the P 1 Enp having inner coxal seta only in the new species. The male P 3 Enp- 2 of the new species has larger setae than those in B. bogoriensis; there are one transformed and four normal setae in the new species but only four normal setae in B. bogoriensis. The outstretched portion of the transformed species on the male P 3 Enp- 2 is less produced compared to those in B. bogoriensis, which are semi-circular. The male P 4 Enp- 1 of the new species has two spiniform outgrowths on the inner margin but is smooth in B. bogoriensis. Bryocyclops trangensis, new species, is different from B. fidjiensis by its anal operculum ovate, which is serrated but triangular and smooth in the later species, and P 4 ’ s twosegmented Exp and one-segmented Enp in the female in the new species but one-segmented Exp and two-segmented in both sexes in B. fidjiensis. The female P 4 Enp of the new species has one spine and four setae but one spine and three setae in B. fidjiensis; and the distal exopod of P 4 has three spines in the new species but four spines in B. fidjiensis. The male P 4 Enp- 1 of the new species has two outgrowths on the inner margin that are absent in B. fidjiensis. Bryocyclops trangensis, new species, differs from B. caroli in both sexes by the following: anal operculum is finely serrated in the new species but coarsely serrated in B. caroli; Enp- 2 P 2 of the new species has four setae but three in B. caroli; and Exp- 2 P 2 – P 3 of the new species have no blunt setae compared to two in B. caroli. The female P 4 Enp of the new species has four subequal setae but one typically long seta and three equal setae in B. caroli. The new species has two spiniform outgrowths on the inner margin in the male P 4 Enp- 1, but this is absent in B. caroli. The male P 3 Enp- 2 of B. trangensis, new species, has four setae, one transformed spine and seta compared to four setae and one transformed spine in B. caroli. The transformed spine of the new species has an acute tip and a short distance between the outstretched part and its tip, but there is a blunt tip and a long distance in B. caroli. Bryocyclops trangensis, new species, and B. campaneri are different from each other as follows: the posterior margins of pediger 2 – 4 dorsally are smooth in B. trangensis, new species, but B. campaneri has transverse crenulated frills; P 2 Enp- 2 has four setae in the new species but three setae in B. campaneri; and P 2 – 3 Exp- 2 has no blunt setae compared to two in B. campaneri. The female P 4 Enp of the new species has no outgrowth near the innermost seta but there is a large rounded outgrowth in B. campaneri. The transformed spine on the male P 3 Enp- 2 of the new species has an outstretched part that is less developed and located close to its tip, but it is well-produced into a semi-circular shaped and located near the mid-length of the spine for B. campaneri.	en	Watiroyram, Santi (2018): Two new species of the genus Bryocyclops Kiefer, 1927 (Copepoda: Cyclopoida: Cyclopidae) from southern Thailand. Raffles Bulletin of Zoology 66: 149-169, DOI: http://doi.org/10.5281/zenodo.4502937
