taxonID	type	description	language	source
062F616DB930FF9727EEBA00FBA45898.taxon	description	(Figures 1 – 186, Tables 1 – 4)	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB930FF9727EEBA00FBA45898.taxon	materials_examined	TYPE SPECIES. Scorpio carpathicus Linnaeus, 1767	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB930FF9727EEBA00FBA45898.taxon	diagnosis	DIAGNOSIS. Total length 20 – 50 mm. Movable fingers of pedipalp chela with MD aligned in a single straight row in entire length of chelal fingers; ID, IAD (5 in number) and OD present. OD are displaced to outer aspect of fingers. Pedipalp chela flat in appearance. Trichobothrial pattern type C. Two subdistal denticles present on cheliceral movable finger dorsal edge. Ventral edge of cheliceral movable finger smooth; serrula absent. Two pedal spurs present on legs. Tarsal spurs on legs absent. Sternum pentagonal in shape. Hemispermatophore lamelliform in shape with broad trunks. Telson without subaculear tubercle. Metasoma I – IV with single median carina usually absent. Taxonomic characters. Characters important for distinguishing species in the genus Euscorpius are: (1) trichobothriotaxy of the pedipalp patella, i. e., number of trichobothria in the external series eb, eb a, esb, em, est, and et, in combination with the number of ventral trichobothria; (2) shape and dimensions of the carapace, pedipalp and metasomal segments; (3) granulation and carination of metasomal segments; (4) coloration; and (5) pectinal tooth count. Two currently valid subgenera can be separated by the number of ventral trichobothria on the pedipalp chela manus, i. e., 4 in Euscorpius sensu stricto, and more than 6 in Polytrichobothrius Birula, 1917. These characters alone are not always sufficient for distinguishing a total of ca. 60 species, most of which were described or revalidated in the last two decades. Taxonomy of this genus remains complex and challenging. Recently, DNA analysis was suggested to be the only solution to this problem (see e. g. Parmakelis et al., 2013; Fet et al., 2018). Based on our studies of this genus and the application of diagnostic characters to the related genus Megacormus Karsch, 1881 (Euscorpiidae) and to the related family Scorpiopidae Kraepelin, 1905, we here propose application of two additional characters to elucidate taxonomy of Euscorpius.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB936FF9127DFBFA3FB0659DF.taxon	description	(Figures 10 – 45, 184, Table 1) http: // zoobank. org / urn: lsid: zoobank. org: act: 24837678 - 7078 - 4297 - B 4 D 9 - 4026 A 0 C 535 EC	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB936FF9127DFBFA3FB0659DF.taxon	materials_examined	TYPE LOCALITY AND TYPE REPOSITORY. Albania, Vlorë County, Mali i Cikës, Maja e Gurrës, 2010 m a. s. l.; HNHM. TYPE MATERIAL. Albania, Vlorë County, Mali i Cikës, Maja e Gurrës, 2010 m a. s. l., 12. VIII. 2004, HNHM No. 2004 / 101, 1 ♂ (holotype), leg. Z. Fehér; Vlorë County, Cikë Mts, pine forest N of the Llogara Pass, 40 ° 12 ' 43.0 " N 19 ° 34 ' 45.0 " E, 860 m a. s. l., 11. V. 2006, 1 ♀ (paratype), leg. Dányi, Kontschán and Murányi, HNHM.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB936FF9127DFBFA3FB0659DF.taxon	etymology	ETYMOLOGY. The species epithet is a patronym honoring our colleague Alberto Bonacina, the author of important early studies on the genus Alpiscorpius (Euscorpiidae).	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB936FF9127DFBFA3FB0659DF.taxon	diagnosis	DIAGNOSIS (♂ ♀). Total length 35 mm. Color reddish black, pedipalp reddish brown, legs and telson yellow. Pedipalp patellar external trichobothria numbers: 5 eb, 4 eb a, 2 esb, 4 em, 4 est, 6 et, ventral aspect of patella with 8 – 9 trichobothria. Pectinal teeth number 9 in male, 7 in female. Chelicerae yellow, very slightly reticulated. Male with pedipalp fingers type C, female with fingers undulate. Dorsal metasomal carinae on III – IV segments irregularly granulated. Dorsolateral metasomal carinae on II – IV segments absent. Ventrolateral metasomal carinae on II – IV segments present and smooth. Metasoma V ventrally with median carina developed. Chela length / width ratio, 2.5 in male and 2.75 in female. Metasoma IV length / width ratio, 1.99 in male and 1.89 in female. Telson length / depth ratio, 2.50 in male and 3.13 in female.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB936FF9127DFBFA3FB0659DF.taxon	description	DESCRIPTION (♀ ♂). Total length about 35 mm in both sexes. The habitus is shown in Figs. 10 – 13. For position and distribution of trichobothria of pedipalps, see Figs. 38 – 44. Sexual dimorphism, see below in the description. Coloration (Figs. 1 – 2). Base color uniformly reddish black, sternites lighter, yellowish brown, legs and telson yellow, pedipalps reddish brown. Chelicerae yellow and very slightly reticulate. Carapace and mesosoma (Figs. 10 – 13, 14 – 17). Carapace finely granulated with several smooth areas; carinae are absent. The anterior margin of the carapace is straight. Carapace with two lateral eyes. Tergites finely granulated, without developed carina. Tergite VII lacking lateral and median carinal pairs. Sternites III – VII smooth and lustrous; VII lacking lateral and median carinae. Stigmata small, narrow ellipical. Pectinal teeth number 9 in male and 7 in female, fulcra are present. Pectines have three marginal lamellae and five middle lamellae. Metasoma and telson (Figs. 22 – 29). Metasoma very sparsely hirsute and finely granulated. Dorsal metasomal carinae on II – V segments distinctively irregularly granulated; dorsolateral carinae absent; ventrolateral carinae present, smooth on segments II – IV and granulated on segment V. Metasoma V ventrally granulated with median carina developed, metasoma I – IV with ventral median carina indicated to absent. Anal arch with small pigmented granules. Telson finely granulated, elongate in female and swollen in male, with annular ring absent in female and developed in male. Aculeus short, more curved in male. Pedipalps (Figs. 30 – 45). Pedipalps very sparsely hirsute. Patella with 25 (5 eb, 4 eb a, 2 esb, 4 em, 4 est, 6 et) external and 8 – 9 (2 x 8, 2 x 9) ventral trichobothria. Chela with 4 ventral trichobothria, of which V 4 is located on external surface near Eb 1. Femur and patella granulated, femur roughly and patella with fine, reticulated granules. Femur has granulated developed carinae; ventroexternal carina is incomplete. Patella with 5 complete carinae including irregular wide externomedian. DPS well developed. Manus dorsally with fine, rounded granules, which do not form median carina; only five chelal carinae are developed, of which dorsoexternal is almost absent in female. Male with pedipalp fingers type C, female with fingers undulate. Legs (Figs. 18 – 21). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules. Measurements. See Table 1. AFFINITIES. E. bonacinai sp. n. is morphologically most similar to E. solegladi Fet et al., 2014 from Bulgaria and Greece. These two species are differentiated according to trichobothrial pattern: E. bonacinai sp. n. has pedipalp patella with 4 eb a and 8 – 9 ventral trichobothria and E. solegladi has 5 – 6 eb a and 9 – 10 ventral trichobothria. DISTRIBUTION. Albania (Fig. 184).	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB939FF852529B87CFDDC5AE9.taxon	description	(Figures 46 – 79, 184 – 185, Tables 2, 4) http: // zoobank. org / urn: lsid: zoobank. org: act: B 71 FEF 8 F- 77 A 6 - 4 CFF-B 1 FE- 1 CCDB 1 F 3503 D	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB939FF852529B87CFDDC5AE9.taxon	materials_examined	TYPE LOCALITY AND TYPE REPOSITORY. North Macedonia, Mavrovo, 41.65 ° N 20.72 ° E; FKCP. TYPE MATERIAL. North Macedonia, Mavrovo, 41.65 ° N 20.72 ° E, VI. 2014, 3 ♂ 1 ♀ (holotype and paratypes, 540, 541, 542), leg. M. MinařÍk, FKCP; Tetovo Province, Shar Planina, road 1209 (Tetovo to Popova Shapka) between Popova Shapka and junction to Shipkovica, 42.020617 ºN 20.903512 ºE, 8. VI. 2014, 1 ♀ (paratype), leg. Petr Janšta et al., FKCP.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB939FF852529B87CFDDC5AE9.taxon	etymology	ETYMOLOGY. The species epithet is a patronym honoring Petr Janšta, the collector of paratype of the new species.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB939FF852529B87CFDDC5AE9.taxon	diagnosis	DIAGNOSIS (♂ ♀). Total length 33 – 40 mm. Color reddish black, legs and telson yellow. Pedipalp patellar external trichobothria numbers: 4 eb, 4 eb a, 2 esb, 4 em, 4 est, 6 et, ventral aspect of patella with 7 – 8 trichobothria. Pectinal teeth number 8 – 9 in males, 7 in females. Chelicerae yellow, slightly reticulated. Male with pedipalp fingers type A, female with fingers undulate. Dorsal metasomal carinae on III – IV segments granulated. Dorsolateral metasomal carinae on II – IV segments absent. Ventrolateral metasomal carinae on II – IV segments present and smooth. Metasoma V ventrally with median carina developed. Chela length / width ratio, 2.3 in males and 2.6 in females. Metasoma IV length / width ratio, 1.95 in males and 2 in females. Telson length / depth ratio, 2.3 in males and 3 in females.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB939FF852529B87CFDDC5AE9.taxon	description	DESCRIPTION (♀ ♂). Total length about 33 – 40 mm in both sexes. The habitus is shown in Figs. 46 – 49. For position and distribution of trichobothria of pedipalps, see Figs. 72 – 78. Sexual dimorphism, see below in the description. Coloration (Figs. 46 – 49). Base color uniformly reddish black, sternites lighter, brown, legs and telson yellow. Chelicerae yellow, slightly reticulate. Carapace and mesosoma (Figs. 46 – 53). Carapace finely granulated with several large granules in anterior part and several smooth areas; carinae are absent. The anterior margin of the carapace is almost straight. Carapace with two lateral eyes. Tergites finely granulated, without developed carina. Tergite VII lacking lateral and median carinal pairs. Sternites III – VII finely granulated, lacking lateral and median carinae. Stigmata small, narrow ellipical. Pectinal teeth number 8 – 9 (2 x 8, 4 x 9) in males and 7 in females, fulcra are present. Pectines have three marginal lamellae and 4 – 6 middle lamellae. Metasoma and telson (Figs. 56 – 63). Metasoma very sparsely hirsute. Metasoma, mainly in dorsal aspect and whole metasoma V finely granulated more in males. Dorsal metasomal carinae on II – V segments distinctively irregularly granulated; dorsolateral carinae absent; ventrolateral carinae present, smooth on segments II – IV and granulated on segment V. Metasoma V ventrally granulated with median carina developed, on metasoma I – IV ventral median carina is absent. Anal arch with small pigmented granules. Telson finely granulated, elongate in females and swollen in males, with annular ring indicated in females and developed in male. Aculeus short, more curved in male. Pedipalps (Figs. 64 – 79). Pedipalps very sparsely hirsute. Patella with 24 (4 eb, 4 eb a, 2 esb, 4 em, 4 est, 6 et) external and 7 – 8 (3 x 7, 7 x 8) ventral trichobothria. Chela with 4 ventral trichobothria, of which V 4 is located on external surface near Eb 1. Femur and patella are granulated, femur roughly and patella with fine, reticulated granules. Femur has granulated developed carinae; ventroexternal carina incomplete. Patella with 5 complete carinae including irregular wide externomedian. DPS moderately developed. Manus dorsally with fine, rounded granules, which do not form median carina; only five chelal carinae are developed. Male with pedipalp fingers type A, female with fingers undulate. Legs (Figs. 54 – 55). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules. Measurements. See Table 2. AFFINITIES. E. janstai sp. n. is morphologically most similar to E. concinnus (C. L. Koch, 1837) from Italy, which differs by darker colored telson and legs, and E. deltshevi Fet et al., 2014 from Bulgaria and eastern Serbia, which differs by having usually a higher number of pedipalp patellar trichobothria in et series. Both of these species differ in having pedipalp fingers type C in males while E. janstai sp. n. has type A (see Figs. 3 – 9 versus Fig. 65). DISTRIBUTION. North Macedonia (Fig. 184).	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB923FF832515B8E3FDE0582A.taxon	description	(Figures 80 – 114, 184, Table 2) http: // zoobank. org / urn: lsid: zoobank. org: act: A 813698 C- 63 F 9 - 40 B 2 - B 2 EA- 45 DE 501 FF 7 D 4 Euscorpius sicanus (in part): Parmakelis et al. 2013: fig. 1 (Locality 60), fig. 2 – 3 (FESP 5). Euscorpius sicanus complex (in part): Fet et al., 2018: 129, fig. 3.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB923FF832515B8E3FDE0582A.taxon	materials_examined	TYPE LOCALITY AND TYPE REPOSITORY. Greece, Mt. Parnassos, Ski Center Parnassos, 38 ° 32.4 ‘ N 22 ° 34.8 ‘ E, 1800 m a. s. l.; FKCP. TYPE MATERIAL. Greece, Mt. Parnassós, Ski Center Parnassou, 38 ° 32.4 ' N 22 ° 34.8 ' E, 1800 m a. s. l., 2 ♂ 4 ♀ 1 ♀ juv. (holotype and paratypes), 5 – 6. VI. 2007, leg. P. Kabátek, 6. VI. 2007, 2 ♀ 1 ♀ juv. (paratypes), leg. V. Ryjáček, FKCP.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB923FF832515B8E3FDE0582A.taxon	etymology	ETYMOLOGY. The species epithet is a patronym honoring Petr Kabátek, the collector of holotype of the new species.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB923FF832515B8E3FDE0582A.taxon	diagnosis	DIAGNOSIS (♂ ♀). Total length 28 – 37 mm. Color reddish yellow. Pedipalp patellar external trichobothria numbers: 5 – 6 eb, 4 eb a, 2 esb, 4 em, 4 est, 7 et, ventral aspect of patella with 9 – 11 trichobothria. Pectinal teeth number 9 – 10 in males, 7 – 9 in females. Chelicerae yellow, without reticulation. Males with pedipalp fingers type C, females with fingers undulate. Dorsal metasomal carinae on III – IV segments granulated. Dorsolateral metasomal carinae on II – IV segments absent. Ventrolateral metasomal carinae on II – IV segments present and smooth. Metasoma V ventrally with median carina indicated. Chela length / width ratio, 2.3 in males and 2.4 – 2.5 in females. Metasoma IV length / width ratio, 2.1 – 2.2 in males and 1.7 – 1.8 in females. Telson length / depth ratio, 2.2 – 2.3 in males and 3.2 – 3.3 in females.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB923FF832515B8E3FDE0582A.taxon	description	DESCRIPTION (♀ ♂). Total length about 28 – 37 mm in both sexes. The habitus is shown in Figs. 80 – 83. For position and distribution of trichobothria of pedipalps, see Figs. 107 – 113. Sexual dimorphism, see below in the description. Coloration (Figs. 80 – 83). Base color uniformly reddish yellow to orange. Chelicerae yellow, without reticulation. Carapace and mesosoma (Figs. 80 – 88). Carapace finely granulated with several smooth areas; carinae are absent. The anterior margin of the carapace is almost straight. Carapace with two lateral eyes. Tergites finely granulated, without developed carina. Tergite VII lacking lateral and median carinal pairs. Sternites III – VII smooth and lustrous, lacking lateral and median carinae. Stigmata small, narrow ellipical. Pectinal teeth number 9 – 10 (1 x 9, 3 x 10) in males and 7 – 9 (2 x 7, 3 x 8, 7 x 9) in females, fulcra are present. Pectines have three marginal lamellae and 4 – 5 middle lamellae. Metasoma and telson (Figs. 91 – 98). Metasoma very sparsely hirsute. Metasoma, mainly in dorsal aspect and whole metasoma V very finely granulated. Dorsal metasomal carinae on II – V segments distinctively irregularly granulated; dorsolateral carinae absent; ventrolateral carinae present, smooth on segments II – IV and granulated on segment V. Metasoma V ventrally granulated with median carina developed, on metasoma I – IV ventral median carina is absent. Anal arch with small pigmented granules. Telson smooth, elongate in females and swollen in males, with annular ring indicated in females and developed in male. Aculeus short more curved in male. Pedipalps (Figs. 99 – 114). Pedipalps very sparsely hirsute. Patella with 26 – 27 (5 – 6 eb [8 x 5, 12 x 6], 4 eb a, 2 esb, 4 em, 4 est, 7 et) external and 9 – 10 (2 x 9, 15 x 10, 3 x 11) ventral trichobothria. Chela with 4 ventral trichobothria, of which V 4 is located on external surface near Eb 1. Femur granulated; patella rather smooth. Femur has granulated developed carinae; ventroexternal carina is incomplete. Patella with 5 complete carinae including irregular wide externomedian. DPS well developed. Manus dorsally with fine solitary granules, almost absent in females; median carina missing, other carinae are smooth. Male with pedipalp fingers type C, female with fingers undulate. Legs (Figs. 89 – 90). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules. Measurements. See Table 2. AFFINITIES. This species belongs to the subgenus Euscorpius Thorell, 1876, s. str.; it is morphologically similar to E. sicanus (C. L. Koch, 1837) from Italy (Sicily), which differs by having darker coloration. The presence of more than 4 (usually 5) trichobothria in eb series is a signature trait of a monophyletic ‘ Euscorpius sicanus complex’ (see Fet et al., 2003; Parmakelis et al., 2013). COMMENTS. Fet et al. (2018: 129) mentioned that “ ‘ Euscorpius sicanus complex’ representatives are found across Central Greece, including Mt. Parnassos ”. After E. sicanus was recently restricted to its type locality of Sicily (Tropea, 2017), the Greek populations of this diverse complex remained unnamed. DNA data are available for this species in Parmakelis et al. (2013) under FESP 5 (Locality 60; a specimen was collected by J. O. Rein in Lilaia, Mt. Parnassos). DISTRIBUTION. Greece, Mt. Parnassos (Fig. 184).	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB925FF89252FBAB8FB0A59B4.taxon	description	(Figures 115 – 149, 184, 186, Tables 3 – 4) http: // zoobank. org / urn: lsid: zoobank. org: act: 382 FE 1 BA- 8307 - 406 D-A 493 - 6 EC 930 BAA 309	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB925FF89252FBAB8FB0A59B4.taxon	materials_examined	TYPE LOCALITY AND TYPE REPOSITORY. Serbia, Sokobanja, 43.63 ° N 21.80 ° E; FKCP. TYPE MATERIAL. Serbia, Sokobanja, 43.63 ° N 21.80 ° E, 14. VI. 2014, leg. D. SadÍlek, 1 ♂ 1 ♀ (holotype and paratype, 543), FKCP; Nišava Province, Gadžin Han env., road between Donji Dušnik and Supotnica, Suva Planina foothills, 43.155 ºN 22.118 ºE, 5. VI. 2014, 1 ♀ (paratype), leg. P. ŠÍpek et al., FKCP.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB925FF89252FBAB8FB0A59B4.taxon	etymology	ETYMOLOGY. The species epithet is a patronym honoring David SadÍlek, the collector of holotype of the new species.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB925FF89252FBAB8FB0A59B4.taxon	diagnosis	DIAGNOSIS (♂ ♀). Total length 26 – 36 mm. Color reddish black, legs and telson yellow. Pedipalp patellar external trichobothria numbers: 4 eb, 4 eb a, 2 esb, 4 em, 4 est, 6 et, ventral aspect of patella with 8 – 9 trichobothria. Pectinal teeth number 9 in male, 6 – 8 in females. Chelicerae yellow, with reticulation. Male with pedipalp fingers type C, female with fingers undulate. Dorsal metasomal carinae on III – IV segments granulated. Dorsolateral metasomal carinae on II – IV segments absent. Ventrolateral metasomal carinae on II – IV segments present and smooth or granulate (mainly in male). Metasoma II – IV densely (roughly) granulated including lateral surface. Metasoma V ventrally granulated with median carina present. Chela length / width ratio, 2.2 in male and 2.4 – 2.5 in females. Metasoma IV length / width ratio, 1.68 in male and 1.8 in females. Telson length / depth ratio, 2.6 in male and 3 – 3.1 in females.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB925FF89252FBAB8FB0A59B4.taxon	description	DESCRIPTION (♀ ♂). Total length about 26 – 36 mm in both sexes. The habitus is shown in Figs. 115 – 118. For position and distribution of trichobothria of pedipalps, see Figs. 141 – 147. Sexual dimorphism, see below in the description. Coloration (Figs. 115 – 118, 149). Base color uniformly reddish black, sternites lighter, yellowish brown, legs and telson yellow. Chelicerae yellow, reticulate. Carapace and mesosoma (Figs. 115 – 122). Carapace finely granulated with several smooth areas; carinae are absent. The anterior margin of the carapace is straight. Carapace with two lateral eyes. Tergites finely granulated, without developed carina. Tergite VII lacking lateral and median carinal pairs. Sternites III – VII rather smooth in females and sparsely finely granulated in male, both lacking lateral and median carinae. Stigmata small, narrow ellipical. Pectinal teeth number 9 in male and 6 – 8 (1 x 6, 1 x 7, 2 x 8) in females, fulcra are present. Pectines have three marginal lamellae and 4 – 6 middle lamellae. Metasoma and telson (Figs. 126 – 133). Metasoma very sparsely hirsute. Metasoma I – V granulated finely in females and roughly in male. Dorsal metasomal carinae on I – V segments distinctively irregularly granulated; dorsolateral carinae absent; ventrolateral carinae present, smooth in females and granulated in male. Metasoma V completely granulated in both sexes with median carina developed, on metasoma I – IV ventral median carina is absent. Anal arch with small pigmented granules. Telson finely granulated, elongate in females and swollen in males, with annular ring present in both sexes. Aculeus short more curved in male. Pedipalps (Figs. 134 – 148). Pedipalps very sparsely hirsute. Patella with 24 (4 eb, 4 eb a, 2 esb, 4 em, 4 est, 6 et) external and 8 – 9 (3 x 8, 1 x 9) ventral trichobothria. Chela with 4 ventral trichobothria, of which V 4 is located on external surface near Eb 1. Female paratype has additional trichobothrium eb a on chela external surface (Fig. 142), which is absent in male holotype. Femur and patella are granulated, femur roughly and patella finely. Femur has granulated developed carinae; ventroexternal carina is incomplete to missing. Patella with 5 complete carinae including irregular wide externomedian. DPS weak to moderate. Manus dorsally with fine, rounded granules, which do not form median carina; only five chelal carinae are developed. Male with pedipalp fingers type C, weakly undulate, similar ar females. Legs (Figs. 123 – 125). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules. Measurements. See Table 3. AFFINITIES. The combination of four characters (pedipalp patellar external trichobothria numbers: 4 eb, 4 eb a; metasoma IV length / width ratio 1.6 – 1.8; male with pedipalp fingers type C and female with fingers undulate; and metasoma II – IV densely (roughly) granulated including lateral surface) is unique in the entire genus Euscorpius. DISTRIBUTION. Serbia (Fig. 184).	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB92DFFB02504BA28FB0F58AF.taxon	description	(Figures 150 – 184, Table 3) http: // zoobank. org / urn: lsid: zoobank. org: act: 487753 D 8 - 3 E 57 - 45 C 6 - B 783 - 4 B 33969 A 4006	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB92DFFB02504BA28FB0F58AF.taxon	materials_examined	TYPE LOCALITY AND TYPE REPOSITORY. Greece, Attiki, Mt. Imittos, near Athens, ca 37.95 ° N 23.74 ° E; FKCP. TYPE MATERIAL. Greece, Attiki, Mt. Imittos, near Athens, ca. 37.95 ° N 23.74 ° E, 2013, 4 ♂ 10 ♀ (holotype and paratypes), leg. Michael Misch; FKCP.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB92DFFB02504BA28FB0F58AF.taxon	etymology	ETYMOLOGY. The species epithet is a patronym honoring our colleague Bernhard Scherabon, the author of important early studies on the genus Alpiscorpius (Euscorpiidae) in Austria.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB92DFFB02504BA28FB0F58AF.taxon	diagnosis	DIAGNOSIS (♂ ♀). Total length 24 – 31 mm. Color reddish brown, legs and telson yellow to orange. Pedipalp patellar external trichobothria numbers: 4 eb, 3 – 4 eb a, 2 esb, 4 em, 4 est, 6 et, ventral aspect of patella with 8 trichobothria. Pectinal teeth number 7 – 8 in males, 6 – 7 in females. Chelicerae yellow, without reticulation. Male with pedipalp fingers type C, female with fingers slightly undulate. Dorsal metasomal carinae on III – IV segments smooth. Dorsolateral metasomal carinae on II – IV segments absent. Ventrolateral metasomal carinae on II – IV segments present and smooth. Metasoma V ventrally with median carina absent. Chela length / width ratio, 2.3 in males and 2.5 in females. Metasoma IV length / width ratio, 2 – 2.15 in males and 1.9 – 2 in females. Telson length / depth ratio, 2.6 – 2.7 in males and 3 – 3.2 in females.	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
062F616DB92DFFB02504BA28FB0F58AF.taxon	description	DESCRIPTION (♀ ♂). Total length about 24 – 31 mm in both sexes. The habitus is shown in Figs. 150 – 153. For position and distribution of trichobothria of pedipalps, see Figs. 176 – 182. Sexual dimorphism, see below in the description. Coloration (Figs. 150 – 153). Base color uniformly reddish brown, legs and telson yellow to orange. Chelicerae yellow, without reticulation. Carapace and mesosoma (Figs. 150 – 157). Carapace finely granulated with several smooth areas; carinae are absent. The anterior margin of the carapace is straight. Carapace with two lateral eyes. Tergites finely granulated (more in males), without developed carina. Tergite VII lacking lateral and median carinal pairs. Sternites III – VII smooth, lacking lateral and median carinae. Stigmata small, narrow ellipical. Pectinal teeth number 7 – 8 (4 x 7, 4 x 8) in males and 6 – 7 (3 x 6, 17 x 7) in females, fulcra are reduced. Pectines have three marginal lamellae and 4 – 5 middle lamellae. Metasoma and telson (Figs. 160 – 167). Metasoma very sparsely hirsute. Metasoma I – IV smooth except several fine granules in dorsal surfaces, metasoma V very sparsely finely granulated, mainly in male. Dorsal metasomal carinae on I – V segments smooth; dorsolateral carinae absent; ventrolateral carinae smooth on segments I – IV and irregularly granulated on segment V. The entire metasoma without ventral median carina. Anal arch with small pigmented granules. Telson smooth, elongate in females and swollen in males, with annular ring indicated in females and developed in male. Aculeus short, more curved in male. Pedipalps (Figs. 168 – 183). Pedipalps very sparsely hirsute. Patella with 23 – 24 (4 eb, 3 – 4 eb a [4 x 3, 24 x 4] 2 esb, 4 em, 4 est, 6 et) external and 8 ventral trichobothria. Chela with 4 ventral trichobothria, of which V 4 is located on external surface near Eb 1. Femur is granulated in both sexes and patella in granulated in males and rather smooth in females. Femur has granulated developed carinae; ventroexternal carina is incomplete or missing. Patella with 5 complete carinae including irregular wide externomedian. DPS moderately developer. Manus smooth in females and finely granulated in males; dorsal median carina missing, other carinae smooth. Male with pedipalp fingers type C, female with fingers undulate. Legs (Figs. 158 – 159). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules. Measurements. See Table 3. AFFINITIES. The combination of five characters (pedipalp patellar external trichobothria numbers: 4 eb, 3 – 4 eb a; metasoma IV length / width ratio 1.9 – 2.15; male with pedipalp fingers type C; metasoma II – IV smooth; and metasoma V ventrally without median carina) is unique in the entire genus Euscorpius. COMMENTS. This species, found within the city limits of Athens, quite surprisingly, was never systematically collected. Fet et al. (2018: 129) recognized it as an undescribed species that “ inhabits the urban zone of Athens, including mountain ranges of Imittos, Pentelikon, and Parnitha ”. No DNA data are available for this species. An interesting reduction of trichobothrial number from 4 to 3 in eb a series is present in some specimens (14.3 % of pedipalps scored, n = 28), a trend that is independently observed in the closely related genus Alpiscorpius (see KovařÍk et al., 2019). DISTRIBUTION. Greece (Fig. 184).	en	Kovařík, František, Šťáhlavský, František (2020): Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia. Euscorpius 315: 1-37, DOI: http://doi.org/10.5281/zenodo.4648646
