taxonID	type	description	language	source
5457263E2A871D65CD7F0A578E7D33C1.taxon	diagnosis	Diagnosis. A large species with usually widely spaced, strong ribs, and no or weak apertural teeth.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
2AC1B5318AD5B49ADD8FACB5FD11896D.taxon	materials_examined	Other material. Vrelo Pronisicut (or Pronifkut), coll. Edlauer ex coll. Kuscer, NHMW 48236 / 1 shell (" photo "); Pronisicut, coll. Edlauer ex coll. Kuscer, NHMW 48336 / 2 shells; Albania, Shkoder district, Drisht, right bank of Kir river opposite to the fortress hill, 90 m a. s. l. (roadside limestone rocks), 42 ° 7.824 ' N, 19 ° 36.540 ' E, (site code: 2016 / 32), leg. Z. P. Eross, Z. Feher, J. Grego & M. Szekeres, 28.06.2016, HNHM 103195 / 1 (photographed shell, Fig. 3 G-L), NHMW 112356 / 1 adult + 5 juvenile shells; Albania, Shkoder district, 1 km NE of Ura e Shtrenjte, 160 m a. s. l., (roadside limestone rocks), 42 ° 9.180 ' N, 19 ° 40.000 ' E (site code: 2016 / 33), leg. Z. P. Eross, Z. Feher, J. Grego & M. Szekeres, 28.06.2016, HNHM 103196 / 1 (photographed shell, Fig. 3 M-R), NHMW 112357 / 9 juvenile / broken shells; Albania, Shkoder district, 4 km SW of Prekal, 170 m a. s. l., (roadside limestone rocks), 42 ° 9.936 ' N, 19 ° 41.334 ' E (site code: 2016 / 35), leg. Z. P. Eross, Z. Feher, J. Grego & M. Szekeres, 28.06.2016, JG / 1 adult + 1 juvenile shell; Albania, Ura e Mesit, debris of the Kir river, 50 m a. s. l., 42 ° 6.870 ' N, 19 ° 34.498 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Apr. 2012, REI / 1; Albania, rocks southwest of Zusi, southwest of Skoder, 13 m a. s. l., 42 ° 2.316 ' N, 19 ° 28.866 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, May 2015, NHMW 112358 / 1 (photographed shell, Fig. 3 S-X), REI / 5; Albania, Felsspalten above Drisht at Mes, 115 m a. s. l., 42 ° 7.735 ' N, 19 ° 36.823 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Jul. 2010, REI / 2 juvenile shells; Albania, ruins 3 km above Drisht at Mes, 195 m a. s. l., 42 ° 7.468 ' N, 19 ° 36.846 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Jul. 2010, REI / 3; Albania, Periferi Shkoder, ca. 18 km upstream from dam at Koman, a left side-valley of Liqeni i Komanit, 170 m a. s. l., limestone rocks, debris, 42 ° 13.613 ' N, 19 ° 54.300 ' E, leg. Z. P. Eross, Z. Feher, A. Hunyadi & D. Muranyi, 15 Apr. 2006, HNHM 102244 / 1.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
2AC1B5318AD5B49ADD8FACB5FD11896D.taxon	diagnosis	Diagnosis. Protoconch matte; aperture with a rather weak palatal and two weak basal teeth.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
2AC1B5318AD5B49ADD8FACB5FD11896D.taxon	description	Description. Shell rarely flat, usually spire somewhat elevated; protoconch consists of 1.5 - 1.75 whorls, very finely granulated, rather matte, not glossy; teleoconch with strong, equidistant ribs that are supported by fine periostracal filaments in fresh shells; rib density variable (43 - 93 ribs on body whorl), usually widely spaced; between main ribs some fine wrinkles discernible; entire shell with 4.25 - 4.5 whorls; aperture semilunar or due to the straight basal part triangular; peristome expanded and slightly reflected, especially on the palatal, basal and umbilical areas; palatal tooth of variable strength, usually weak, although present in all adult shells, palatal region of peristome without outer incision; basal portion of peristome usually straight, slightly thickened, with two low denticles that are visible in all adult shells; umbilicus funnel-shaped, wide (although width depends on spire height). Measurements. SW: 3.6 - 4.3 mm (median = 4.0 mm), SH: 1.9 - 2.2 mm (median = 2.0 mm), AW: 1.3 - 1.7 mm (median = 1.5 mm), AH: 1.3 - 1.6 mm (median = 1.5 mm), (n = 6; largest and smallest specimens of multiple populations measured).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
2AC1B5318AD5B49ADD8FACB5FD11896D.taxon	distribution	Distribution. This subspecies is distributed in northwestern Albania. Most of the known distribution records are from the Kir River Valley. One shell found in the fluvial debris of the Drin River above the Koman Dam extends the range farther eastwards (Figure 4). Welter-Schultes (2012) erroneously reports this taxon from Montenegro.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
9C12CEDA763A2D52A8FCACF407F76960.taxon	materials_examined	Additional material. Albania, Malesia district, Xhaje NE 0.5 km, 650 m a. s. l., 42 ° 19.838 ' N, 19 ° 36.341 ' E (site code: 2015 / 103), rocks, leg. Z. P. Eross, Z. Feher & J. Grego, 04.07.2015, HNHM 103493 / 5 juvenile shells (not paratypes); NHMW 110430 / MN / 0985 / 5 juvenile shells (not paratypes), JG / 6 juvenile shells (not paratypes).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
9C12CEDA763A2D52A8FCACF407F76960.taxon	diagnosis	Diagnosis. Protoconch glossy; aperture without teeth. Measurements. SW: 3.6 - 4.2 mm (median = 4.0 mm), SH: 1.9 - 2.2 mm (median = 2.1 mm), AW: 1.5 - 1.7 mm (median = 1.6 mm), AH: 1.3 - 1.6 mm (median = 1.4 mm), (n = 6; largest and smallest specimens of multiple populations measured).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
9C12CEDA763A2D52A8FCACF407F76960.taxon	etymology	Etymology. This new subspecies is named after its toothless aperture, which distinguishes it from the nominotypical subspecies.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
9C12CEDA763A2D52A8FCACF407F76960.taxon	distribution	Distribution. This taxon is found in the northeastern part of the Lake Shkoder Basin (Figure 4).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
54424E035AC3DAF2C040C4D72C1C1DF4.taxon	distribution	Distribution. The genus Spelaeodiscus has a disjunct distribution. One species (S. hauffeni) is only known from Slovenia, whereas the rest of the genus is distributed in the vicinity of the Skadar Lake Basin (also known as Shkoder Lake or Skutari Lake) in Montenegro and northern Albania (Figure 2).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
A8A5ACDDB663AE8CC5C192AE08235441.taxon	materials_examined	Other material. Vetajama bei Sokol, coll. Edlauer, NHMW 48071 / 2, (det. Gittenberger, 1974); Radetina pecina, Itijino brdo, 1300 m, leg. Dabovic, NHMW 48312 / 11 (large shells, similar to the type); Pecina Marka Vuksanovica, 1400 m, leg. Dabovic, coll. Edlauer ex coll. Kuscer 666 / 10, NHMW 49321 / 1 (det. Gittenberger, 1974); Montenegro, S of Virpazar, 1 km (in a straight line) ESE of Limljani, near the small road, 323 m a. s. l., 42 ° 11.414 ' N, 19 ° 06.277 ' E (site code: 20171019 B), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 19.10.2017, DT / 2, HA / 1, HNHM 103198 / 2, PGB / 1; Montenegro, S of Virpazar, 0.8 km (in a straight line) NE of Limljani, near the small road, 350 m a. s. l., 42 ° 12.068 ' N, 19 ° 05.969 ' E (site code: 20171019 C), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 19.10.2017, DT / 2, HA / 2. HNHM 103199 / 2, PGB / 1; Montenegro, 1.9 km (in a straight line) S of Virpazar, near the road, 160 m a. s. l., 42 ° 13.312 ' N, 19 ° 05.440 ' E (site code: 20171019 E), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 19.10.2017, DT / 24, EZP / 24, HA / 24, HNHM 103200 / 24 + 1 photographed shell (Fig. 6 K-O), PGB / 24; Montenegro, SE of Virpazar, 4.3 km (in a straight line) SSE of Ðuravci, near Besa / Bes near Krone i Besit, 330 m a. s. l., 42 ° 08.548 ' N, 19 ° 13.165 ' E (site code: 20171019 G), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 19.10.2017, DT / 60 - 70, EZP / 60 - 70, HA / 60 - 70, HNHM 103201 / 28, PGB / 60 - 70; Montenegro, SE of Virpazar, 7.3 km S (SE) of Ðuravci, 1.6 km (in a straight line) NNW of Tejani, 455 m a. s. l., 42 ° 06.803 ' N, 19 ° 13.384 ' E (site code: 20171019 I), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 19.10.2017, HNHM 103202 / 2; Montenegro, NNW of Virpazar, road between Rijeka Crnojevica and Virpazar, at the junction to Dupilo, 160 m a. s. l., 42 ° 15.085 ' N, 19 ° 04.987 ' E (site code: 20171020 A), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 20.10.2017, HNHM 103203 / 1; Montenegro, S of Virpazar, 1.1 km (in a straight line) E of Limljani, one of the tunnels on the old road, 400 m a. s. l., 42 ° 11.637 ' N, 19 ° 06.309 ' E (site code: 20171021 A), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 21.10.2017, DT / ca. 20, EZP / ca. 20, HA / ca. 20, HNHM 103204 / 6, PGB / ca. 20; Montenegro, S of Virpazar, 0.8 km (in a straight line) E of Limljani, above the village, 400 m a. s. l., 42 ° 11.698 ' N, 19 ° 06.217 ' E (site code: 20171021 B), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 21.10.2017, DT / ca. 40, EZP / ca. 40, HA / ca. 40, HNHM 103205 / 8, PGB / ca. 40; Montenegro, NE of Bar, 1.2 km (in a straight line) NW of Tudjemili, 400 m a. s. l., 42 ° 08.489 ' N, 19 ° 08.144 ' E (site code: 20171021 F), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 21.10.2017, DT / ca. 40, EZP / ca. 40, HA / ca. 40, HNHM 103206 / 10, PGB / ca. 40; Montenegro, 1.6 km (in a straight line) NE Petrovac, 0.3 km S of Novoselje, 470 m a. s. l., 42 ° 13.032 ' N, 18 ° 57.336 ' E (site code: 20171021 G), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 21.10.2017, EZP / 3; Montenegro, 2.8 km (in a straight line) NE Petrovac, 1.2 km E of Novoselje, 630 m a. s. l., 42 ° 13.145 ' N, 18 ° 58.245 ' E (site code: 20171021 H), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 21.10.2017, DT / 3, EZP / 2, HA / 4, HNHM 103207 / 1, PGB / 2; Montenegro, Gradiste Monastery, 80 m a. s. l., 42 ° 12.212 ' N, 18 ° 57.772 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Jul. 2010, NHMW 112362 / 1 (photographed shell, Fig. 6 F-J); REI / 5; Montenegro, above Sv. Stefan south of Budva, 110 m a. s. l., 42 ° 14.926 ' N, 18 ° 54.131 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Mar. 2011, REI / 4; Montenegro, Tuđemili, 17 km towards Virpazar, Rumija Mts, 480 m a. s. l., 42 ° 10.974 ' N, 19 ° 6.546 ' E, leg. P. Subai, 15 Apr. 2009, NMBE 542070 / 56; Montenegro, Tuđemili, 1 km towards Virpazar, Rumija Mts, 395 m a. s. l., 42 ° 8.238 ' N, 19 ° 8.388 ' E, leg. P. Subai, 25 Sep. 2005, NMBE 542069 / 1; Montenegro, Tuđemili, 1 km towards Virpazar, Rumija Mts, 395 m a. s. l., 42 ° 8.238 ' N, 19 ° 8.388 ' E, leg. P. Subai, 25 Sep. 2005, NMBE 542068 / 3; Montenegro, Ðuravci, 10 Km E, southern side of Mount Kronistar, 500 m a. s. l., 42 ° 6.594 ' N, 19 ° 13.908 ' E, leg. P. Subai, 16 Sep. 2006, NMBE 542067 / 1; Montenegro, Zoganje N 5 km, on the Ulcinj-Shkoder road, 80 m a. s. l., 41 ° 58.762 ' N, 19 ° 15.547 ' E (site code: 2015 / 32), leg. T. Deli, Z. P. Eross & Z. Feher, 27 May 2016, DT / ca. 50, HNHM 103208 / 22, NHMW 112363 / ca. 50; Montenegro, Rumija Mts, Virpazar S 9 km (Virpazar-Bar road, between Boljevici and Tuđemili), old tunnel, 440 m, 42 ° 11.474 ' N, 19 ° 06.489 ' E (site code: 2008 / 182), leg. Z. Feher, J. Kontschan & D. Muranyi, 14.10.2008, HNHM 103209 / 23; Montenegro, scree (talus) ca. 5 km east of Dobra Voda in direction of Vladimir, 220 m a. s. l., 42 ° 1.508 ' N, 19 ° 11.160 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Jul. 2010, REI / 6; Montenegro, Donji Murici, above the village, at the junction of a minor road to Besa, 200 m a. s. l., 42 ° 9.233 ' N, 19 ° 12.930 ' E (site code 2017 / 007), leg. Z. P. Eross & Z. Feher 16.07.2017, HNHM 103492 / 1 (juvenile shell, identification uncertain).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
A8A5ACDDB663AE8CC5C192AE08235441.taxon	diagnosis	Diagnosis. A small to medium sized species with dense, low ribs, smooth protoconch, and toothless aperture.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
A8A5ACDDB663AE8CC5C192AE08235441.taxon	description	Description. Shell nearly flat, but spire somewhat always elevated; protoconch consists of 1.25 - 1.75 whorls, rather glossy; teleoconch with fine, equidistant, dense ribs; rib density variable (57 - 112 ribs on body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.25 - 3.75 whorls; aperture semilunar; peristome slightly thickened and expanded; aperture toothless; umbilicus regular funnel-shaped, relatively narrow (width depends on spire height). Measurements. SW: 1.9 - 3.4 mm (median = 2.3 mm), SH: 1.1 - 1.7 mm (median = 1.3 mm), AW: 0.7 - 1.3 mm (median = 0.9 mm), AH = 0.8 - 1.3 mm (median = 0.9 mm), AA = 56 - 68 ° (n = 15).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
A8A5ACDDB663AE8CC5C192AE08235441.taxon	distribution	Distribution. This species is distributed in the Rumija Mountain between the Shkoder Lake Basin and the Adriatic Sea. Northwards the range extends to the Cetinje area. According to the original labels, type material is of Slovenian origin, however, this species was never again found in Slovenia. It can be reasonably supposed that it is due to mislabelling and the ' type locality' is not the site where the type material actually came from (Gittenberger 1975). Welter-Schultes (2012) reports the species only from Slovenia, which is based on the originally incorrect type locality.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
327E01A9D8F14601243E8A85C6851237.taxon	diagnosis	Diagnosis. A large species with very low and dense ribs; aperture toothless.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
327E01A9D8F14601243E8A85C6851237.taxon	description	Description. Spire slightly elevated; protoconch consists of slightly more than 1.5 whorls, rather glossy; teleoconch with very fine, low, equidistant riblets (approx. 112 on the body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75 whorls; aperture semilunar, peristome; peristome expanded and slightly reflected on the basal and umbilical areas; aperture toothless; umbilicus regular funnel-shaped, relatively wide. Measurements. SW = 3.7 mm, SH = 1.8 mm, AW = 1.5 mm, AH = 1.4 mm (holotype).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
327E01A9D8F14601243E8A85C6851237.taxon	etymology	Etymology. This new species is named for its dense ribs, which distinguishes it from the most similar S. albanicus.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
327E01A9D8F14601243E8A85C6851237.taxon	distribution	Distribution. This species is known from the type locality only (Figure 7).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
EE99D18B1BD685F0BF2418D5757617C6.taxon	materials_examined	Other material. Velikajama, Soko Kod, Sela Dopilo (geographic position unknown), coll. Edlauer ex coll. Dabovic, NHMW 48473 / 2 shells (1 adult + 1 juvenile) (in brackets: neben 49.999, probably mistyped 48.999, because this was a mixed lot of S. hauffeni and S. dejongi; this is obviously incorrect locality for this species); Jama nadjama pri Gnezdu (Izilovice) (= Jama pri Gnezdu at 45.939 ° N, 14.271 ° E), coll. Edlauer ex coll. Kuscer, NHMW 49137 (?) / 1 (incorrect locality); Krain, coll. Oberwimmer, NHMW 71640 / O / 00161 / 2; Carn., coll. Schmidt, NHMW 112346 / 2; Krain, coll. Kobelt ex coll. Ullepitsch, SMF 10663 / 2 adult + 2 juvenile shells; Krain: Stubic, coll. C. R. Boettger, 1905, SMF 112878 / 1; Krainer Hoehlen, coll. Kaltenbach ex coll. Mueller, SMF 259470 / 1; Carniolia (= Krain), coll. Knobbe ex coll. Hauffen, SMF 53904 / 1; Hoehle in Innerkrain, leg. Sever, coll. Ehrmann ex coll. Absolon, SMF 53905 / 1; Tekavicja jama (= Tkavcja jama, 45.824 ° N, 14.721 ° E), Dobrepolje, NHMW 112350 / 2 strongly corroded shells; Krain, Tekavcja jama, Debrepolje, coll. Edlauer ex coll. Kuscer, NHMW 48468 / 3; Tekarija jama b. Dobropolje, coll. S. H. Jaeckel ex coll. Kuscer, SMF 200961 / 3; Krain, Tekavcja jama, Dobrepolje, coll. Haas ex coll. Kuscer, SMF 53906 / 1; Krain, Berjakovo Brezno (= Malo Brezarjevo brezno, 46.080 ° N, 14.436 ° E), n. w. von Laibach, coll. Retaner (?), NHMW 12314 / 2; Berjakovo Brezno, nw v. Laibach, Krain, coll. Klemm ex coll. Edlauer, NHMW 79000 / K / 02938 / 2; Same data, coll. Schlickum, ex coll. Edlauer, SMF 275440 / 2; Berjakovo Brezno bei Dolnice n. w. von Ljubljana, coll. S. H. Jaeckel ex coll. Edlauer, SMF 200962 / 1; Radetina pecina, coll. Edlauer ex coll. Kuscer, NHMW 48751 / 6; Spodnja Skedevenica (at 45.854 ° N, 14.639 ° E), vel. Lasce, coll. Edlauer ex coll. Kuscer, NHMW 48883 / 15; Spodnja Skedevnica, coll. Edlauer ex coll. Kuscer, NHMW 48422 / 2; Spodnja Kedenrica, coll. W. Klemm ex coll. Kuscer, NHMW 33672 / 1 (figured shell, Fig. 8 K-O); Sponja Skednevica bei Preserje, coll. Edlauer ex coll. Kuscer, NHMW 48124 / 2; Am Eingang der Doline Jama bei Kompole (= Kompoljska jama at 45.800 ° N, 14.731 ° E), S. Oest. von Dobrepolje, coll. Klemm, NHMW 112347 / 1; In der Raska Skedebnza bei Ponikve suedl. von Dobrepolje (= Skedenc at 45.89687418 ° N, 14.50100919 ° E), coll. Klemm, NHMW 112348 / 3; Reoteiner Grotte bei Poduzik (?) bei Ljubljana (Kalkofen) (Podutisko brezno at 46.076 ° N, 14.445 ° E), coll. Edlauer ex coll. Kuscer, NHMW 49111 / 1; Brezarjeva brezno pri Boduzik (?), coll. Edlauer ex coll. Kuscer, NHMW 48390 / 2; Brezno v. Leskovi dolini (at 45.941 ° N, 14.256 ° E), coll. Edlauer ex coll. Kuscer, NHMW 49200 / 1; Masun b Gratenbrunn (probably Jama na Masunu at 45.630 ° N, 14.373 ° E), Krain, leg. Schollmayer, NHMW 112349 / 2 juvenile shells; Duplica (= Dupliska jama at 45.963 ° N, 14.684 ° E), coll. Edlauer ex coll. Kuscer, NHMW 48576 / 16; Jama v Jurcelovih (?) (Jama 1 v Jurcetovih Percah at 46.09984722 ° N, 14.42093707 ° E or Jama 2 na Jurcetovih Percah at 46.101 ° N 14.418 ° E, the first is indicated on the map), percah Tosko celo, coll. Edlauer ex coll. Kuscer, NHMW 48688 / 16; Zakrita jama odd B (at 45.925 ° N, 14.293 ° E), coll. Edlauer ex coll. Kuscer, NHMW 48748 / 1 juvenile shell; Unterkrain: Grotte Thauzhia jama, coll. C. v. Heyden ex coll. N. Hoffmann, SMF 53903 / 1; Same data, coll. Gysser ex coll. Heynemann ex coll. C. v. Heyden, SMF 53901 / 2; Same data, coll. Heynemann ex coll. C. v. Heyden ex coll. N. Hoffmann, SMF 259469 / 3; Krain: Jeliarc-Grotte (maybe Jeliare or Jelice), coll. Th. Krueper, SMF 112877 / 1; Hoehle Malo Bukuje bei Dobrova, Krain, coll. Dr. Leo Rusnov, ex coll. Dr. A. Oberwimmer, NHMW 71770 / R / 19 (1 photographed shell, Fig. 8 F-J).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
EE99D18B1BD685F0BF2418D5757617C6.taxon	diagnosis	Diagnosis. A medium sized species with very widely spaced, strong ribs, rounded, toothless aperture, and finely granular protoconch.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
EE99D18B1BD685F0BF2418D5757617C6.taxon	description	Description. Spire somewhat elevated; protoconch consists of 1.5 - 1.75 whorls, very finely granulated, rather matte, not glossy; teleoconch with strong ribs that are supported by fine periostracal filaments in fresh shells; ribbing less regular than in other congeneric species; ribs widely spaced (41 - 52 ribs on body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75 - 4.25 whorls; aperture toothless, semilunar / rounded; peristome slightly thickened, slightly reflected in direction of umbilicus; umbilicus regular funnel-shaped, relatively narrow. Measurements. SW: 2.8 - 3.5 mm (median = 3.0 mm), SH: 1.7 - 1.9 mm (median = 1.8 mm), AW: 1.1 - 1.4 mm (median = 1.2 mm), AH: 1.0 - 1.3 mm (median = 1.1 mm), (n = 6; largest and smallest specimens of multiple populations measured).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
EE99D18B1BD685F0BF2418D5757617C6.taxon	distribution	Distribution. This species is distributed in the southeastern Alps (Central Slovenia) relatively far from the ranges of its congeneric taxa. Slapnik (2005) found shells in the Skocjan Caves Regional Park, which are probably of Holocene age (Figure 9).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
AE80E259B9B1F01056A44B67B31E0FA1.taxon	diagnosis	Diagnosis. A small, nearly flat species with strong, widely spaced ribs, glossy protoconch and strongly oblique, toothless aperture.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
AE80E259B9B1F01056A44B67B31E0FA1.taxon	description	Description. Spire somewhat elevated; protoconch consists of 1.25 - 1.5 whorls, smooth, glossy; teleoconch with very strong (thick), equidistant, widely spaced ribs; rib density: 42 - 48 ribs on body whorl; between main ribs some fine wrinkles discernible; entire shell with 3.5 - 3.75 whorls; aperture semilunar, toothless, strongly oblique to shell axis; peristome slightly thickened and expanded in direction of umbilicus; umbilicus funnel-shaped, relatively narrow. Measurements. SW: 2.1 - 2.2 mm (median = 2.2 mm), SH: 1.2 - 1.3 mm (median = 1.2 mm), AW: 0.8 - 0.9 mm (median = 0.9 mm), AH: 0.6 - 0.7 mm (median = 0.7 mm), AA = 47 - 52 ° (n = 4; largest and smallest specimens measured).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
AE80E259B9B1F01056A44B67B31E0FA1.taxon	etymology	Etymology. This new species is named after our colleague and friend, Andras Hunyadi, who is one of those who first collected this species.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
AE80E259B9B1F01056A44B67B31E0FA1.taxon	distribution	Distribution. This species is known from the type locality only (Figure 7).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
5237C950927075D50D41CF76B3A5E789.taxon	materials_examined	Other material. Montenegro, Seoca S 1 km, along the Virpazar-Ostros road, 280 m a. s. l., 42 ° 12.618 ' N, 19 ° 9.000 ' E (site code: 2017 / 005), leg. Z. P. Eross & Z. Feher, 16.07.2017, HNHM 103213 / 3 shells, not paratypes (Fig. 10 K-O).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
5237C950927075D50D41CF76B3A5E789.taxon	diagnosis	Diagnosis. A small, nearly flat species with strong, very widely spaced ribs, glossy protoconch and a toothless aperture.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
5237C950927075D50D41CF76B3A5E789.taxon	description	Description of the holotype. Spire somewhat elevated; protoconch consists of ca 1.25 - 1.5 whorls (the holotype is corroded at the protoconch-teleoconch junction), rather smooth, moderately glossy; teleoconch with very strong, equidistant, widely spaced ribs (42 on the body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75 whorls; aperture semilunar, toothless; peristome slightly thickened and expanded; umbilicus funnel-shaped, relatively narrow. Measurements. SW: 2.2 mm, SH: 1.2 mm, AW: 0.8 mm, AH: ca. 0.8 mm, AA = 63 ° (holotype).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
5237C950927075D50D41CF76B3A5E789.taxon	etymology	Etymology. This new species is named after its remarkably widely spaced ribs.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
5237C950927075D50D41CF76B3A5E789.taxon	distribution	Distribution. See under Remarks and Figure 7.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
4CCB7D685CD1F8B5B817DB16E619B447.taxon	materials_examined	Material examined. Vodna jama (Lovcen), coll. Edlauer ex coll. Kuscer, NHMW 48543 / 2 (det. Gittenberger, 1973 Sep.); Same data, NHMW 49765 / 2; Montenegro, Ostrog, rocks below the upper parking lot, 820 m a. s. l., 42 ° 40.534 ' N, 19 ° 1.744 ' E, leg. A. Reischuetz, P. L. Reischuetz & N. Steiner-Reischuetz, May 2017, REI / 1; Montenegro, Ostrog, entrance of the penultimate parking lot, 770 m a. s. l., 42 ° 40.510 ' N, 19 ° 1.663 ' E, leg. A. Reischuetz, P. L. Reischuetz & N. Steiner-Reischuetz, May 2017, NHMW 112365 (photographed shell, Fig. 11 F-J), REI / 1; Montenegro, Ostrog, 1 km north from Hotel Sokoline on the new street to Podgorica, 630 m a. s. l., 42 ° 39.498 ' N, 19 ° 2.188 ' E, leg. A. Reischuetz, P. L. Reischuetz & N. Steiner-Reischuetz, May 2017, REI / 2; Montenegro, 500 m south of the bridge over the Sitnica river, north of Podgorica, 45 m a. s. l., 42 ° 27.473 ' N, 19 ° 10.825 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Apr. 2012, NHMW 112366 / 1 (photographed shell, Fig. 11 K-O), REI / 2; Yugoslavia: Pecina u Peckom Brdu bg., above Zacir, leg. L. Pinter & P. Subai, 20 Jul. 1972, HNHM 41127 / 2; Same data, HNHM 18062 / 3 (one of them is photographed, Fig. 11 A-E); Montenegro, Pecina u peckom Brdu bei Zacir (Brdu-cave at Zacir), leg. H. Schuett, 06.06.1978, HNHM 42402 / 3.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
4CCB7D685CD1F8B5B817DB16E619B447.taxon	diagnosis	Diagnosis. A medium sized species with elevated spire, roughly sculptured protoconch, strong ribs on the teleoconch, and toothless aperture.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
4CCB7D685CD1F8B5B817DB16E619B447.taxon	description	Description. Spire elevated, shell low conical; protoconch consists of 1.25 - 1.5 whorls, roughly granulated / " hammered ", matte, not glossy; teleoconch with strong, equidistant ribs that are supported by fine periostracal filaments in fresh shells; rib density variable (43 - 76 ribs on body whorl), usually widely spaced; between main ribs some fine wrinkles discernible; entire shell with 3.75 - 4 whorls; aperture semilunar, toothless; peristome slightly thickened and expanded, especially in direction of the umbilicus; umbilicus funnel-shaped, relatively narrow. Measurements. SW: 2.6 - 3.0 mm (median = 2.8 mm), SH: 1.6 - 1.9 mm (median = 1.7 mm), AW: 0.9 - 1.2 mm (median = 1.1 mm), AH: 1.0 - 1.2 mm (median = 1.1 mm), AA = 64 - 70 ° (n = 10; largest and smallest specimens of multiple populations measured).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
4CCB7D685CD1F8B5B817DB16E619B447.taxon	distribution	Distribution. This species is found northwest of the Shkoder Lake Basin, as well as in the Zeta River Valley between Podgorica and the Ostrog Monastery (Figure 4). The Albanian record given by Reischuetz et al. (2013) and referred by Reischuetz (2017 c) is actually S. albanicus edentatus ssp. n. So far, no Albanian occurrence is known.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
36EF9D32E0A8CA5100DA999403D10A99.taxon	diagnosis	Diagnosis. A small to medium sized species with dense riblets, and strong parietal and basal teeth / thickenings.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
F885B3E00F6762818BEFE50D0A1AFC3B.taxon	diagnosis	Diagnosis. Basal tooth pointed; palatal part of peristome with slight incision at the position of the palatal tooth, palatal region not or slightly " pushed " from outside. Measurements. SW: 2.9 - 3.5 mm (median = 3.0 mm), SH: 1.4 - 1.8 mm (median = 1.5 mm), AW: 1.1 - 1.3 mm (1.2 mm), AH: 1.1 - 1.4 mm (median = 1.2 mm), (n = 9; largest and smallest specimens of multiple populations measured).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
F885B3E00F6762818BEFE50D0A1AFC3B.taxon	etymology	Etymology. This new subspecies is named after its pointed (Latin: acutus) basal tooth, which distinguishes it from the nominotypical subspecies.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
F885B3E00F6762818BEFE50D0A1AFC3B.taxon	distribution	Distribution. This subspecies is known from the southeastern side of the Shkoder Lake Basin in Albania (Figure 7).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
557123F26FE62D66CECD1A174B4C427A.taxon	materials_examined	Other material. Bioce, N Titograd, Mtg. 23.09.1978, MZBI 15443 (photos of the sample where examined); Megara, Tolusi, Titograd, Mtg. MZBI 2335 (photos of the sample where examined); Pecina od Zavora, Peuta, Titograd, Mtg. 01.11.1963, MZBI 2338 (photos of the sample where examined); Hoehle bei Virpazar, leg. Dabovic, coll. Edlauer, NHMW 48881 / 1; Brosine cave (33), leg. Kuscer 10266 / 67, coll. Edlauer, NHMW 38874 / a (20 juvenile shells, " schlechte Expl "); Same data, NHMW 58874 / 4 (" bessere Expl "); Virpazar, mit " oculus mundi ", leg. Dabovic, coll. Edlauer, NHMW 78988 / 3 juvenile shells (det. Gittenberger, but identification not certain); Smarjetna gora + Grabocica cave (drawing of a cave on the label), (Trnovo), NHMW 78989 / 7; Same data, NHMW 48086 / 5; Vodna jama (Lovcen), coll. Edlauer ex coll. Kuscer, NHMW 48544 / 1; Same locality, coll. Edlauer, NHMW 49764 / 1 (labelled as " Spelaeodiscus albanicus hadzii "); Montenegro, NNW of Virpazar, road between Rijeka Crnojevica and Virpazar, 0.4 km (in a straight line) W of Poseljani, 130 m a. s. l., 42 ° 18.361 ' N, 19 ° 02.898 ' E (site code: 20171020 F), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 20.10.2017, DT / ca. 15, EZP / ca. 15, HA / ca. 15, HNHM 103248 / 5, PGB / ca. 15; Montenegro, W of Rijeka Crnojevica, 1 km (in a straight line) NNE of Zacir, above Obodska pecina, near the road, 320 m a. s. l., 42 ° 21.168 ' N, 19 ° 00.125 ' E (site code: 2017.10.20 J), leg. T. Deli, Z. P. Eross, A. Hunyadi & B. Pall-Gergely, 20.10.2017, DT / ca. 18, EZP / ca. 18, HA / ca. 18, HNHM 103249 / 5, PGB / ca. 18; Montenegro, 2 km north of Velje Brdo in the Zeta valley, north of Podgorica, 70 m a. s. l., 42 ° 28.932 ' N, 19 ° 14.454 ' E, leg. A. Reischuetz, P. L. Reischuetz & N. Steiner-Reischuetz, May 2016, REI / 5; Montenegro, Moraca valley, after the exit to the " China Road and Bridge " camp N of Bioce, 70 m a. s. l., 42 ° 29.167 ' N, 19 ° 18.646 ' E, leg. A. Reischuetz, P. L. Reischuetz & N. Steiner-Reischuetz, May 2016, NHMW 112368 / 1 (photographed shell, Fig. 13 G-L), REI / 10; Montenegro, road between Rijeka Crnojevica and Hrvasi, 180 m a. s. l., 42 ° 21.491 ' N, 19 ° 4.281 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Apr. 2012, REI / 11; Montenegro, scree slope at the BMS-petrol station, south of the Moraca gorge, 5 km south of Bioce, 90 m a. s. l., 42 ° 29.064 ' N, 19 ° 18.279 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Jul. 2008, REI / 1; Montenegro, spring 1 km south of the junction towards Njive, south of Rijeka Crnojevica, 150 m a. s. l., 42 ° 18.266 ' N, 19 ° 2.900 ' E, leg. A. Reischuetz, N. Reischuetz & P. L. Reischuetz, Mar. 2011, NHMW 112369 / 1 (photographed shell, Fig. 13 M-R); Montenegro, Zabljak Crnojevica, fortress, 50 m a. s. l., 42 ° 19.030 ' N, 19 ° 9.401 ' E (site code: 2015 / 007), leg. T. Deli, Z. P. Eross & Z. Feher, 25.05.2015, HNHM 103250 / 1 adult and several juvenile shells; Montenegro, Zacir, " Pecina u pecko brdo " (cave), 450 m a. s. l., leg. L. Pinter, P. Subai, A. Szigeti, 20 Jul. 1972, NMBE 542072 / 76; Montenegro, Pecina u peckom, Brdu-bg Zacirnal (Brdu-cave at Zacir), leg. L. Pinter & P. Subai, 20 Jul. 1972, HNHM 65187 / 7 (one of them is photographed: Figs 13 A-F); Montenegro, Pecina u peckom Brdu bei Zacir (Brdu-cave at Zacir), leg. H. Schuett, 06.06.1978, coll. Gy. Kovacs, HNHM 65188 / 1; Same data, HNHM 42401 / 2; Montenegro, Rijeka Crnojevica, Obodska cave, 110 m a. s. l., 42 ° 21.120 ' N, 19 ° 0.318 ' E, leg. P. Subai & M. Szekeres, 23 Sep. 2006; NMBE 542071 / 1; Montenegro, Poseljani, on the Virpazar-Rijeka Crnojevica road, 150 m a. s. l., 42 ° 18.347 ' N, 19 ° 2.884 ' E (site code: 2015 / 016), leg. T. Deli, Z. P. Eross & Z. Feher, 25.05.2015, HNHM 103251 / 8; NHMW 112370 / 8, DT / 8.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
557123F26FE62D66CECD1A174B4C427A.taxon	diagnosis	Diagnosis. Basal tooth / thickening low, not pointed; palatal part of peristome with strong incision at the position of the palatal tooth, palatal region strongly " pushed " from outside.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
557123F26FE62D66CECD1A174B4C427A.taxon	description	Description. Spire elevated; protoconch consists of 1.5 whorls, roughly granulated / " hammered ", matte, not glossy; teleoconch with strong but dense, equidistant ribs that are supported by fine periostracal filaments in fresh shells; rib density variable (74 - 118 ribs on body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75 - 4.5 whorls; aperture semilunar or triangular due to straight basal part; peristome thickened, slightly expanded and slightly reflected on the palatal, basal and umbilical areas; palatal tooth strong, pointed; palatal region of peristome with strong outer incision (i. e. the position of the palatal tooth is indicated with a groove on the outer side); basal portion of peristome straight, thickened; occasionally two small denticles visible, sometimes only the one situated closer to the palatal side is developed as a low denticle; umbilicus funnel-shaped, wide to relatively narrow (width depends on spire height). Measurements. SW: 2.4 - 3.2 mm (median = 2.7 mm), SH: 1.3 - 1.8 mm (median = 1.5 mm), AW: 0.9 - 1.1 mm (median = 1.0 mm), AH: 1.0 - 1.4 mm (median = 1.1 mm), (n = 11; largest and smallest specimens of multiple populations measured).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
557123F26FE62D66CECD1A174B4C427A.taxon	distribution	Distribution. This taxon is found north and northwest of the Shkoder Lake Basin in Montenegro (Figure 7). Albanian records given by Reischuetz et al. (2013) actually refer to S. unidentatus acutus ssp. n.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
3CE3B48E8A328FFAF621E885FDBE1638.taxon	diagnosis	Diagnosis. A medium sized to large species with elevated spire, strong, very widely spaced ribs, fine spiral lines consisting of series of minute tubercles, hammered protoconch, weak basal thickening, weak parietal tooth, and a thickened parietal callus.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
3CE3B48E8A328FFAF621E885FDBE1638.taxon	description	Description. Spire elevated; protoconch consists of ca. 1.25 - 1.75 whorls, roughly sculptured, " hammered "; teleoconch with very strong, equidistant, widely spaced ribs (40 - 70 on the body whorl; addition to the radial ribs very fine, dense spiral striation is visible on the entire teleoconch, consisting of minute tubercles; entire shell with 4 - 4.75 whorls; aperture subcircular / triangular; basal part with two low swellings (similar to those of S. albanicus albanicus), parietal wall with a weak tooth; parietal callus thickened, blunt; umbilicus funnel-shaped, relatively narrow. Measurements. SW: 3.3 - 3.6 mm (median = 3.5 mm), SH: 1.95 - 2.25 mm (median = 2.1 mm), AW: 1.3 - 1.6 mm (median = 1.4 mm), SW: 1.4 - 1.5 mm (median = 1.4 mm), (n = 11).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
3CE3B48E8A328FFAF621E885FDBE1638.taxon	etymology	Etymology. This new species is named after its resemblance to Virpazaria species based on the thickened parietal callus.	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
3CE3B48E8A328FFAF621E885FDBE1638.taxon	distribution	Distribution. Spelaeodiscus virpazarioides sp. n. is known from the type locality only (see also Figure 7).	en	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
