identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5457263E2A871D65CD7F0A578E7D33C1.text	5457263E2A871D65CD7F0A578E7D33C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus albanicus (A. J. Wagner 1914)	<div><p>Spelaeodiscus albanicus (A. J. Wagner, 1914)</p> <p>Diagnosis.</p> <p>A large species with usually widely spaced, strong ribs, and no or weak apertural teeth.</p> <p>Differential diagnosis.</p> <p>The most similar species in terms of shell size and shape is Spelaeodiscus densecostatus sp. n., for differences see under that species. Spelaeodiscus unidentatus is usually smaller, usually possesses denser ribs, and has stronger teeth and narrower aperture.</p> <p>Conservation status.</p> <p>Reischütz and Fehér (2017) assessed this species as Least Concern (LC). They claimed that there are at least three known locations and is likely that further field work reveals a larger range and more locations. Although that assessment was based partly on incorrect distribution records (Peuta Cave population is currently treated as S. unidentatus, whereas Raps-Starjë population as S. virpazarioides sp. n.), together with our new distribution records there are more than five locations. As we have no reason to suppose that the habitat quality, habitat extent, or population are deteriorating or extremely fluctuating, Least Concern (LC) seems to be a correct assessment.</p> </div>	http://treatment.plazi.org/id/5457263E2A871D65CD7F0A578E7D33C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
2AC1B5318AD5B49ADD8FACB5FD11896D.text	2AC1B5318AD5B49ADD8FACB5FD11896D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus albanicus subsp. albanicus (A. J. Wagner 1914) albanicus (A. J. Wagner 1914	<div><p>Spelaeodiscus albanicus albanicus (A. J. Wagner, 1914) Figure 3</p> <p>Aspasita albanica A. J. Wagner, 1914 in Sturany &amp; Wagner 1914: 67, plate 2, figs 10a-c.</p> <p>Spelaeodiscus albanicus - Bole 1965: 354, Plate 76, fig. B.</p> <p>Spelaeodiscus (Spelaeodiscus) albanicus - Gittenberger 1969: 294, fig. 2.</p> <p>Spel aeodiscus albanicus - Pilsbry 1926: 184, plate 22, figs 12-14.</p> <p>Spelaeodiscus albanicus - Welter-Schultes 2012: 213 (partim: locality data from Montenegro are of unknown origin).</p> <p>Spelaeodiscus albanicus - Reischütz et al. 2013: 62, fig. 3.</p> <p>Type material.</p> <p>Kiri-Brücke nächst Mesi b. Skutari, Albanien (im Genist), leg. Sturany, 27.04.1905, NHMW 43385 (lectotype, hereby selected, SW: 3.7 mm, SH: 2 mm, Fig. 3A-F); Drinasca-Ufer b. Skutari (angeschwemmt), leg. Sturany, 03.05.1905, NHMW 112351 (1 corroded, juvenile paralectotype).</p> <p>Other material.</p> <p>Vrelo Pronisicut (or Pronifkut), coll. Edlauer ex coll. Kuščer, NHMW 48236/1 shell (“photo”); Pronisicut, coll. Edlauer ex coll. Kuščer, NHMW48336/2 shells; Albania, Shkodër district, Drisht, right bank of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.609&amp;materialsCitation.latitude=42.1304" title="Search Plazi for locations around (long 19.609/lat 42.1304)">Kir river</a> opposite to the fortress hill, 90 m a.s.l. (roadside limestone rocks), 42°7.824'N, 19°36.540'E, (site code: 2016/32), leg. Z.P. Erőss, Z. Fehér, J. Grego &amp; M. Szekeres, 28.06.2016, HNHM 103195/1 (photographed shell, Fig. 3G-L), NHMW 112356/1 adult + 5 juvenile shells; Albania, Shkodër district, 1 km NE of Ura e Shtrenjtë, 160 m a.s.l., (roadside limestone rocks), 42°9.180'N, 19°40.000'E (site code: 2016/33), leg. Z.P. Erőss, Z. Fehér, J. Grego &amp; M. Szekeres, 28.06.2016, HNHM 103196/1 (photographed shell, Fig. 3M-R), NHMW 112357/9 juvenile/broken shells; Albania, Shkodër district, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.6889&amp;materialsCitation.latitude=42.1656" title="Search Plazi for locations around (long 19.6889/lat 42.1656)">4 km SW of Prekal</a>, 170 m a.s.l., (roadside limestone rocks), 42°9.936'N, 19°41.334'E (site code: 2016/35), leg. Z.P. Erőss, Z. Fehér, J. Grego &amp; M. Szekeres, 28.06.2016, JG/1 adult + 1 juvenile shell; Albania, Ura e Mesit, debris of the <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.574966&amp;materialsCitation.latitude=42.1145" title="Search Plazi for locations around (long 19.574966/lat 42.1145)">Kir river</a>, 50 m a.s.l., 42°6.870'N, 19°34.498'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Apr. 2012, REI/1; Albania, rocks southwest of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.4811&amp;materialsCitation.latitude=42.0386" title="Search Plazi for locations around (long 19.4811/lat 42.0386)">Zusi</a>, southwest of Skoder, 13 m a.s.l., 42°2.316'N, 19°28.866'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, May 2015, NHMW 112358/1 (photographed shell, Fig. 3S-X), REI/5; Albania, Felsspalten above Drisht at <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.613716&amp;materialsCitation.latitude=42.128918" title="Search Plazi for locations around (long 19.613716/lat 42.128918)">Mes</a>, 115 m a.s.l., 42°7.735'N, 19°36.823'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Jul. 2010, REI/2 juvenile shells; Albania, ruins 3 km above Drisht at <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.6141&amp;materialsCitation.latitude=42.124466" title="Search Plazi for locations around (long 19.6141/lat 42.124466)">Mes</a>, 195 m a.s.l., 42°7.468'N, 19°36.846'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Jul. 2010, REI/3; Albania, Periferi Shkodër, ca. 18 km upstream from dam at Koman, a left side-valley of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.905&amp;materialsCitation.latitude=42.226883" title="Search Plazi for locations around (long 19.905/lat 42.226883)">Liqeni</a> i <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.905&amp;materialsCitation.latitude=42.226883" title="Search Plazi for locations around (long 19.905/lat 42.226883)">Komanit</a>, 170 m a.s.l., limestone rocks, debris, 42°13.613'N, 19°54.300'E, leg. Z.P. Erőss, Z. Fehér, A. Hunyadi &amp; D. Murányi, 15 Apr. 2006, HNHM 102244/1.</p> <p>Diagnosis.</p> <p>Protoconch matte; aperture with a rather weak palatal and two weak basal teeth.</p> <p>Description.</p> <p>Shell rarely flat, usually spire somewhat elevated; protoconch consists of 1.5-1.75 whorls, very finely granulated, rather matte, not glossy; teleoconch with strong, equidistant ribs that are supported by fine periostracal filaments in fresh shells; rib density variable (43-93 ribs on body whorl), usually widely spaced; between main ribs some fine wrinkles discernible; entire shell with 4.25-4.5 whorls; aperture semilunar or due to the straight basal part triangular; peristome expanded and slightly reflected, especially on the palatal, basal and umbilical areas; palatal tooth of variable strength, usually weak, although present in all adult shells, palatal region of peristome without outer incision; basal portion of peristome usually straight, slightly thickened, with two low denticles that are visible in all adult shells; umbilicus funnel-shaped, wide (although width depends on spire height).</p> <p>Measurements. SW: 3.6-4.3 mm (median = 4.0 mm), SH: 1.9-2.2 mm (median = 2.0 mm), AW: 1.3-1.7 mm (median = 1.5 mm), AH: 1.3-1.6 mm (median = 1.5 mm), (n = 6; largest and smallest specimens of multiple populations measured).</p> <p>Differential diagnosis.</p> <p>See under S. albanicus edentatus ssp. n. Densely ribbed specimens of this subspecies might resemble large specimens of S. unidentatus. The latter species, however, usually has stronger teeth, and narrower aperture.</p> <p>Variation among specimens.</p> <p>Some variability was found between populations in terms of rib density, spire height, and strength of apertural teeth.</p> <p>Distribution.</p> <p>This subspecies is distributed in northwestern Albania. Most of the known distribution records are from the Kir River Valley. One shell found in the fluvial debris of the Drin River above the Koman Dam extends the range farther eastwards (Figure 4). Welter-Schultes (2012) erroneously reports this taxon from Montenegro.</p> <p>Conservation status.</p> <p>See under Spelaeodiscus albanicus.</p> </div>	http://treatment.plazi.org/id/2AC1B5318AD5B49ADD8FACB5FD11896D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
9C12CEDA763A2D52A8FCACF407F76960.text	9C12CEDA763A2D52A8FCACF407F76960.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus albanicus subsp. edentatus Pall-Gergely & P. L. Reischuetz 2018	<div><p>Spelaeodiscus albanicus edentatus Pall-Gergely &amp; P. L. Reischuetz ssp. n. Figure 5A-L</p> <p>Spelaeodiscus sp. (Spelaeodiscus aff. obodensis) - Reischütz et al. 2013: 62.</p> <p>Type material.</p> <p>Albania, rocks along the road Hani i Hotit to Vermosh, 7.2-7.8 km north of the junction, 370 m a.s.l., 42°22.451'N, 19°27.507'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Apr. 2014, NHMW 112360/1 (holotype, SW: 4.2 mm, SH: 2.2 mm, Fig. 5A-F), REI/4 paratypes; rocks above Hani i Hotit, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Apr. 2012, REI/1 paratypes; Montenegro, Podgorica Municipality, Izvor Vitoja S 1 km, near Shkodra Lake, 30 m, limestone rocks, 42°19.176'N, 19°22.158'E (site code: 2015/105), leg. Z.P. Erőss, Z. Fehér, J. Grego, 05 Jul. 2015, HNHM 103197/4 paratypes, JG/4 paratypes, NHMW 112359/4 paratypes; Montenegro, rocks across the road at Vitoja, Skadarsko Jezero, 16 m a.s.l., 42°19.554'N, 19°21.776'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Apr. 2012, NHMW 112361/1 (photographed paratype, Fig. 5G-L), REI/4 paratypes.</p> <p>Additional material.</p> <p>Albania, Malesia district, Xhajë NE 0.5 km, 650 m a.s.l., 42°19.838'N, 19°36.341'E (site code: 2015/103), rocks, leg. Z.P. Erőss, Z. Fehér &amp; J. Grego, 04.07.2015, HNHM 103493/5 juvenile shells (not paratypes); NHMW 110430/MN/0985/5 juvenile shells (not paratypes), JG/6 juvenile shells (not paratypes).</p> <p>Type locality.</p> <p>Albania, rocks along the road Hani i Hotit to Vermosh, 7.2-7.8 km north of the junction, 370 m a.s.l., 42°22.451'N, 19°27.507'E.</p> <p>Diagnosis.</p> <p>Protoconch glossy; aperture without teeth.</p> <p>Measurements. SW: 3.6-4.2 mm (median = 4.0 mm), SH: 1.9-2.2 mm (median = 2.1 mm), AW: 1.5-1.7 mm (median = 1.6 mm), AH: 1.3-1.6 mm (median = 1.4 mm), (n = 6; largest and smallest specimens of multiple populations measured).</p> <p>Differential diagnosis.</p> <p>This new subspecies differs from the nominotypical subspecies in the following traits: aperture relatively larger, and its basal area not straight; apertural barriers (teeth) absent; protoconch smooth, glossy; ribs somewhat less dense (rib density on body whorl: 35-54).</p> <p>Variation among specimens.</p> <p>This subspecies shows some variability in terms of shell size and rib density.</p> <p>Etymology.</p> <p>This new subspecies is named after its toothless aperture, which distinguishes it from the nominotypical subspecies.</p> <p>Distribution.</p> <p>This taxon is found in the northeastern part of the Lake Shkodër Basin (Figure 4).</p> <p>Conservation status.</p> <p>See under Spelaeodiscus albanicus.</p> </div>	http://treatment.plazi.org/id/9C12CEDA763A2D52A8FCACF407F76960	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
2F1634B5DD85C3BE36E277C50EF57316.text	2F1634B5DD85C3BE36E277C50EF57316.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus Brusina 1886	<div><p>Family Spelaeodiscidae Steenberg, 1925</p> <p>Remarks.</p> <p>Schileyko (1998) classified Spelaeodiscus into the family Spelaeodiscidae Steenberg, 1925, which has independently been introduced by Hudec (1970) as well. This taxon was also recognized as a separate family of the infraorder Orthurethra (and its only superfamily, Pupilloidea) by Bouchet et al. (2017). Although the genital anatomy of some species belonging to this family is known, its systematic position within Orthurethra is uncertain, because no molecular phylogenetic information is known (Harl et al. 2017).</p> </div>	http://treatment.plazi.org/id/2F1634B5DD85C3BE36E277C50EF57316	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
54424E035AC3DAF2C040C4D72C1C1DF4.text	54424E035AC3DAF2C040C4D72C1C1DF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus Brusina 1886	<div><p>Genus Spelaeodiscus Brusina, 1886</p> <p>Patula (Spelaeodiscus) Brusina, 1886: 37.</p> <p>Type species.</p> <p>Helix Hauffeni Schmidt, 1855.</p> <p>Distribution.</p> <p>The genus Spelaeodiscus has a disjunct distribution. One species (S. hauffeni) is only known from Slovenia, whereas the rest of the genus is distributed in the vicinity of the Skadar Lake Basin (also known as Shkodër Lake or Skutari Lake) in Montenegro and northern Albania (Figure 2).</p> <p>Included taxa.</p> <p>Spelaeodiscus albanicus albanicus (A. J. Wagner, 1914), S. albanicus edentatus Páll-Gergely &amp; P. L. Reischütz, ssp. n., S. dejongi Gittenberger, 1969, S. densecostatus Páll-Gergely &amp; A. Reischütz, sp. n., S. hauffeni (Schmidt, 1855) S. hunyadii Páll-Gergely &amp; Deli, sp. n., S. latecostatus Páll-Gergely &amp; Erőss, sp. n., S. obodensis Bole, 1965. S. unidentatus unidentatus Bole, 1961, S. unidentatus acutus Páll-Gergely &amp; Fehér, ssp. n., S. virpazarioides Páll-Gergely &amp; Fehér, sp. n. For key traits see Table 1.</p> <p>Delimitation of this genus.</p> <p>The reproductive anatomy of Spelaeodiscus and Aspasita is characterized by a short penial caecum, a well-developed penial appendix, sometimes an epiphallic caecum, and a bursa copulatrix without a diverticulum. The retractor muscle is divided into two bounds, one inserting on the penial appendix, whereas the other at the base of the penial caecum. Examining the anatomical descriptions and drawings of Spelaeodiscus (Bole 1965) and Aspasita (Hudec 1965, Gittenberger 1975, Schileyko 1998, Subai and Dedov 2008), we were unable to find characters that would constantly differ between the two groups. For example, the penial caecum was long and slender in A. tatrica (see Hudec 1965) and S. hauffeni (see Bole 1965), but was short and conical in A. triaria (see Subai and Dedov 2008) and S. unidentatus (see Bole 1965). Also, the shape of the bursa and the position of the starting point of the penial appendix was greatly variable across genera. Clear epiphallic caecum was only found in S. hauffeni, but some thickening was visible in S. unidentatus and A. triaria.</p> <p>As for shell characters, Spelaeodiscus is characterized by a mostly colourless shell that is smaller than 4.3 mm (majority of species are even smaller than 3.5 mm), the spire is relatively low (height of body whorl at least two third of the height of the entire shell), the body whorl is evenly rounded, the edge of the parietal callus is straight, and the peristome is only slightly expanded. In contrast, Aspasita shells are brownish, larger than 4.3 mm, they have higher spire (height of body whorl is approximately half of the height of the entire shell), the shell is shape reverse trapezoid from standard apertural view, the callus is heart-shaped, and the basal part of the peristome is strongly expanded.</p> <p>The habitat was the only “trait” mentioned by Gittenberger (1969) as difference between the two groups. Namely, Spelaeodiscus is subterranean, whereas Aspasita can be found on rock surfaces and among leaf litter at the base of limestone rocks. In the lack of sound molecular data it is difficult to infer their relationship, but based only on ecological, conchological, and biogeographical differences it seems reasonable to keep Aspasita and Spelaeodiscus as distinct genera.</p> </div>	http://treatment.plazi.org/id/54424E035AC3DAF2C040C4D72C1C1DF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
A8A5ACDDB663AE8CC5C192AE08235441.text	A8A5ACDDB663AE8CC5C192AE08235441.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus dejongi Gittenberger 1969	<div><p>Spelaeodiscus dejongi Gittenberger, 1969 Figure 6</p> <p>Spelaeodiscus (Spelaeodiscus) dejongi Gittenberger, 1969: 295-296, fig. 3.</p> <p>Spelaeodiscus dejongi - Welter-Schultes 2012: 213.</p> <p>Type material.</p> <p>Jama Nadjama bei Gnezdu (Izitovice), Krain, leg. Kuščer, coll. Edlauer, NHMW 49517a (holotype, SW: 2.8 mm, SH: 1.6 mm, Fig. 6A-E); Same data, NHMW 49517b/17 paratypes. See remarks concerning the type locality.</p> <p>Other material.</p> <p>Vetajama bei Sokol, coll. Edlauer, NHMW 48071/2, (det. Gittenberger, 1974); Radetina pećina, Itijino brdo, 1300 m, leg. Dabović, NHMW 48312/11 (large shells, similar to the type); Pećina Marka Vuksanovića, 1400 m, leg. Dabović, coll. Edlauer ex coll. Kuščer 666/10, NHMW 49321/1 (det. Gittenberger, 1974); Montenegro, S of Virpazar, 1 km (in a straight line) ESE of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.104616&amp;materialsCitation.latitude=42.190235" title="Search Plazi for locations around (long 19.104616/lat 42.190235)">Limljani</a>, near the small road, 323 m a.s.l., 42°11.414'N, 19°06.277'E (site code: 20171019B), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 19.10.2017, DT/2, HA/1, HNHM 103198/2, PGB/1; Montenegro, S of Virpazar, 0.8 km (in a straight line) NE of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.099483&amp;materialsCitation.latitude=42.201134" title="Search Plazi for locations around (long 19.099483/lat 42.201134)">Limljani</a>, near the small road, 350 m a.s.l., 42°12.068'N, 19°05.969'E (site code: 20171019C), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 19.10.2017, DT/2, HA/2. HNHM 103199/2, PGB/1; Montenegro, 1.9 km (in a straight line) <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.090666&amp;materialsCitation.latitude=42.221867" title="Search Plazi for locations around (long 19.090666/lat 42.221867)">S of Virpazar</a>, near the road, 160 m a.s.l., 42°13.312'N, 19°05.440'E (site code: 20171019E), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 19.10.2017, DT/24, EZP/24, HA/24, HNHM 103200/24 + 1 photographed shell (Fig. 6K-O), PGB/24; Montenegro, SE of Virpazar, 4.3 km (in a straight line) SSE of Ðuravci, near Besa/Bes near <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.219418&amp;materialsCitation.latitude=42.142467" title="Search Plazi for locations around (long 19.219418/lat 42.142467)">Krone</a> i <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.219418&amp;materialsCitation.latitude=42.142467" title="Search Plazi for locations around (long 19.219418/lat 42.142467)">Besit</a>, 330 m a.s.l., 42°08.548'N, 19°13.165'E (site code: 20171019G), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 19.10.2017, DT/60-70, EZP/60-70, HA/60-70, HNHM 103201/28, PGB/60-70; Montenegro, SE of Virpazar, 7.3 km S(SE) of Ðuravci, 1.6 km (in a straight line) NNW of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.223066&amp;materialsCitation.latitude=42.113384" title="Search Plazi for locations around (long 19.223066/lat 42.113384)">Tejani</a>, 455 m a.s.l., 42°06.803'N, 19°13.384'E (site code: 20171019I), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 19.10.2017, HNHM 103202/2; Montenegro, NNW of Virpazar, road between Rijeka Crnojevića and Virpazar, at the junction to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.083117&amp;materialsCitation.latitude=42.251415" title="Search Plazi for locations around (long 19.083117/lat 42.251415)">Dupilo</a>, 160 m a.s.l., 42°15.085'N, 19°04.987'E (site code: 20171020A), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 20.10.2017, HNHM 103203/1; Montenegro, S of Virpazar, 1.1 km (in a straight line) E of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.10515&amp;materialsCitation.latitude=42.19395" title="Search Plazi for locations around (long 19.10515/lat 42.19395)">Limljani</a>, one of the tunnels on the old road, 400 m a.s.l., 42°11.637'N, 19°06.309'E (site code: 20171021A), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 21.10.2017, DT/ca. 20, EZP/ca. 20, HA/ca. 20, HNHM 103204/6, PGB/ca. 20; Montenegro, S of Virpazar, 0.8 km (in a straight line) E of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.103617&amp;materialsCitation.latitude=42.194965" title="Search Plazi for locations around (long 19.103617/lat 42.194965)">Limljani</a>, above the village, 400 m a.s.l., 42°11.698'N, 19°06.217'E (site code: 20171021B), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 21.10.2017, DT/ca. 40, EZP/ca. 40, HA/ca. 40, HNHM 103205/8, PGB/ca. 40; Montenegro, NE of Bar, 1.2 km (in a straight line) NW of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.135733&amp;materialsCitation.latitude=42.141483" title="Search Plazi for locations around (long 19.135733/lat 42.141483)">Tudjemili</a>, 400 m a.s.l., 42°08.489'N, 19°08.144'E (site code: 20171021F), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 21.10.2017, DT/ca. 40, EZP/ca. 40, HA/ca. 40, HNHM 103206/10, PGB/ca. 40; Montenegro, 1.6 km (in a straight line) NE Petrovac, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.9556&amp;materialsCitation.latitude=42.2172" title="Search Plazi for locations around (long 18.9556/lat 42.2172)">0.3 km S of Novoselje</a>, 470 m a.s.l., 42°13.032'N, 18°57.336'E (site code: 20171021G), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 21.10.2017, EZP/3; Montenegro, 2.8 km (in a straight line) NE Petrovac, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.97075&amp;materialsCitation.latitude=42.21908" title="Search Plazi for locations around (long 18.97075/lat 42.21908)">1.2 km E of Novoselje</a>, 630 m a.s.l., 42°13.145'N, 18°58.245'E (site code: 20171021H), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 21.10.2017, DT/3, EZP/2, HA/4, HNHM 103207/1, PGB/2; Montenegro, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.962866&amp;materialsCitation.latitude=42.203533" title="Search Plazi for locations around (long 18.962866/lat 42.203533)">Gradiste Monastery</a>, 80 m a.s.l., 42°12.212'N, 18°57.772'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Jul. 2010, NHMW 112362/1 (photographed shell, Fig. 6F-J); REI/5; Montenegro, above Sv. Stefan south of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.902184&amp;materialsCitation.latitude=42.248768" title="Search Plazi for locations around (long 18.902184/lat 42.248768)">Budva</a>, 110 m a.s.l., 42°14.926'N, 18°54.131'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Mar. 2011, REI/4; Montenegro, Tuđemili, 17 km towards Virpazar, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.1091&amp;materialsCitation.latitude=42.1829" title="Search Plazi for locations around (long 19.1091/lat 42.1829)">Rumija Mts</a>, 480 m a.s.l., 42°10.974'N, 19°6.546'E, leg. P. Subai, 15 Apr. 2009, NMBE 542070/56; Montenegro, Tuđemili, 1 km towards Virpazar, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.1398&amp;materialsCitation.latitude=42.1373" title="Search Plazi for locations around (long 19.1398/lat 42.1373)">Rumija Mts</a>, 395 m a.s.l., 42°8.238'N, 19°8.388'E, leg. P. Subai, 25 Sep. 2005, NMBE 542069/1; Montenegro, Tuđemili, 1 km towards Virpazar, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.1398&amp;materialsCitation.latitude=42.1373" title="Search Plazi for locations around (long 19.1398/lat 42.1373)">Rumija Mts</a>, 395 m a.s.l., 42°8.238'N, 19°8.388'E, leg. P. Subai, 25 Sep. 2005, NMBE 542068/3; Montenegro, Ðuravci, 10 Km E, southern side of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.2318&amp;materialsCitation.latitude=42.1099" title="Search Plazi for locations around (long 19.2318/lat 42.1099)">Mount Kronistar</a>, 500 m a.s.l., 42°6.594'N, 19°13.908'E, leg. P. Subai, 16 Sep. 2006, NMBE 542067/1; Montenegro, Zoganje N 5 km, on the Ulcinj-Shkodër road, 80 m a.s.l., 41°58.762'N, 19°15.547'E (site code: 2015/32), leg. T. Deli, Z.P. Erőss &amp; Z. Fehér, 27 May 2016, DT/ca. 50, HNHM 103208/22, NHMW 112363/ca. 50; Montenegro, Rumija Mts, Virpazar S 9 km (<a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.10815&amp;materialsCitation.latitude=42.191235" title="Search Plazi for locations around (long 19.10815/lat 42.191235)">Virpazar-Bar</a> road, between Boljevići and Tuđemili), old tunnel, 440 m, 42°11.474'N, 19°06.489'E (site code: 2008/182), leg. Z. Fehér, J. Kontschán &amp; D. Murányi, 14.10.2008, HNHM 103209/23; Montenegro, scree (talus) ca. 5 km east of Dobra Voda in direction of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.186&amp;materialsCitation.latitude=42.025135" title="Search Plazi for locations around (long 19.186/lat 42.025135)">Vladimir</a>, 220 m a.s.l., 42°1.508'N, 19°11.160'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Jul. 2010, REI/6; Montenegro, Donji Murići, above the village, at the junction of a minor road to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.2155&amp;materialsCitation.latitude=42.153885" title="Search Plazi for locations around (long 19.2155/lat 42.153885)">Besa</a>, 200 m a.s.l., 42°9.233'N, 19°12.930'E (site code 2017/007), leg. Z.P. Erőss &amp; Z. Fehér 16.07.2017, HNHM 103492/1 (juvenile shell, identification uncertain).</p> <p>Diagnosis.</p> <p>A small to medium sized species with dense, low ribs, smooth protoconch, and toothless aperture.</p> <p>Description.</p> <p>Shell nearly flat, but spire somewhat always elevated; protoconch consists of 1.25-1.75 whorls, rather glossy; teleoconch with fine, equidistant, dense ribs; rib density variable (57-112 ribs on body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.25-3.75 whorls; aperture semilunar; peristome slightly thickened and expanded; aperture toothless; umbilicus regular funnel-shaped, relatively narrow (width depends on spire height).</p> <p>Measurements. SW: 1.9-3.4 mm (median = 2.3 mm), SH: 1.1-1.7 mm (median = 1.3 mm), AW: 0.7-1.3 mm (median = 0.9 mm), AH = 0.8-1.3 mm (median = 0.9 mm), AA = 56-68°(n = 15).</p> <p>Differential diagnosis.</p> <p>See under S. obodensis and S. hunyadii sp. n.</p> <p>Variation among specimens.</p> <p>This is a widely distributed species with numerous known populations, most of them with unique character states of spire height, shell size, and rib density.</p> <p>Distribution.</p> <p>This species is distributed in the Rumija Mountain between the Shkodër Lake Basin and the Adriatic Sea. Northwards the range extends to the Cetinje area. According to the original labels, type material is of Slovenian origin, however, this species was never again found in Slovenia. It can be reasonably supposed that it is due to mislabelling and the 'type locality’ is not the site where the type material actually came from (Gittenberger 1975). Welter-Schultes (2012) reports the species only from Slovenia, which is based on the originally incorrect type locality.</p> <p>Conservation status.</p> <p>Assessed as Least Concern (LC) by Reischütz (2017a), because it is not an extremely rare species and there is no reason to suppose that the habitat quality, habitat extent or population are deteriorating or extremely fluctuating. Now, the number of known locations is more than 20, which confirms the LC status.</p> </div>	http://treatment.plazi.org/id/A8A5ACDDB663AE8CC5C192AE08235441	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
327E01A9D8F14601243E8A85C6851237.text	327E01A9D8F14601243E8A85C6851237.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus densecostatus Pall-Gergely & A. Reischuetz 2018	<div><p>Spelaeodiscus densecostatus Pall-Gergely &amp; A. Reischuetz sp. n. Figure 5M-R</p> <p>Type material.</p> <p>Montenegro, Hotel/Restaurant Izvor north of Sutomore, no GPS available, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, May 2015, NHMW 112364 (holotype: SW = 3.7 mm, SH = 1.8 mm, Figure. 5M-R), REI/1 juvenile/broken paratype.</p> <p>Type locality.</p> <p>Montenegro, Hotel/Restaurant Izvor north of Sutomore.</p> <p>Diagnosis.</p> <p>A large species with very low and dense ribs; aperture toothless.</p> <p>Description.</p> <p>Spire slightly elevated; protoconch consists of slightly more than 1.5 whorls, rather glossy; teleoconch with very fine, low, equidistant riblets (approx. 112 on the body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75 whorls; aperture semilunar, peristome; peristome expanded and slightly reflected on the basal and umbilical areas; aperture toothless; umbilicus regular funnel-shaped, relatively wide.</p> <p>Measurements. SW = 3.7 mm, SH = 1.8 mm, AW = 1.5 mm, AH = 1.4 mm (holotype).</p> <p>Differential diagnosis.</p> <p>Spelaeodiscus densecostatus sp. n. differs from S. albanicus by the smaller shell and the much denser and lower ribs on the teleoconch. Furthermore, S. albanicus albanicus has two basal and a palatal tooth in the aperture.</p> <p>Variation among specimens.</p> <p>The only known adult shell is the holotype. Therefore, the morphological diversity within population is unknown.</p> <p>Etymology.</p> <p>This new species is named for its dense ribs, which distinguishes it from the most similar S. albanicus.</p> <p>Distribution.</p> <p>This species is known from the type locality only (Figure 7).</p> <p>Conservation status.</p> <p>The number of known locations of this species is less than five (i.e. known from a single site) and AOO is smaller than 20 km2, but there is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number of mature individuals are declining or extremely fluctuating. Therefore, it should be assessed as Near Threatened (NT).</p></div> 	http://treatment.plazi.org/id/327E01A9D8F14601243E8A85C6851237	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
EE99D18B1BD685F0BF2418D5757617C6.text	EE99D18B1BD685F0BF2418D5757617C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus hauffeni (Schmidt 1855)	<div><p>Spelaeodiscus hauffeni (Schmidt, 1855) Figure 8A-O</p> <p>Helix Hauffeni Schmidt, 1855: 3-4.</p> <p>Patula Hauffeni - Clessin 1887: 104.</p> <p>Aspasita Hauffeni - Sturany and Wagner 1914: 67, plate 2, figs 11a-c.</p> <p>Spel aeodiscus hauffeni - Pilsbry 1926: 185, Plate 22, Figs 9 - 11.</p> <p>Spelaeodiscus hauffeni - Bole 1965: 351-352, fig. 1b, 2a, Plate 76, fig. A.</p> <p>Spelaeodiscus (Spelaeodiscus) hauffeni - Gittenberger 1969: 292-293, fig. 1.</p> <p>Spelaeodiscus hauffeni - Welter-Schultes 2012: 213.</p> <p>Type material.</p> <p>Krain: Krimberg-Grotte, ex coll. Schmidt, SMF 53902/3 syntypes; No locality, ("Orig. Ex."), NHMW 52776/2 syntypes (Figure 8A-E).</p> <p>Other material.</p> <p>Velikajama, Soko Kod, Sela Dopilo (geographic position unknown), coll. Edlauer ex coll. Dabović, NHMW 48473/2 shells (1 adult + 1 juvenile) (in brackets: neben 49.999, probably mistyped 48.999, because this was a mixed lot of S. hauffeni and S. dejongi; this is obviously incorrect locality for this species); Jama nadjama pri Gnezdu (Izilovice) (= Jama pri Gnezdu at 45.939°N, 14.271°E), coll. Edlauer ex coll. Kuščer, NHMW 49137(?)/1 (incorrect locality); Krain, coll. Oberwimmer, NHMW 71640/O/00161/2; Carn., coll. Schmidt, NHMW 112346/2; Krain, coll. Kobelt ex coll. Ullepitsch, SMF 10663/2 adult+2 juvenile shells; Krain: Stubič, coll. C.R. Boettger, 1905, SMF 112878/1; Krainer Höhlen, coll. Kaltenbach ex coll. Müller, SMF 259470/1; Carniolia (= Krain), coll. Knobbe ex coll. Hauffen, SMF 53904/1; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.293&amp;materialsCitation.latitude=45.925" title="Search Plazi for locations around (long 14.293/lat 45.925)">Hoehle in Innerkrain</a> , leg. Sever, coll. Ehrmann ex coll. Absolon, SMF 53905/1; Tekavičja jama (= Tkavčja jama, 45.824°N, 14.721°E), Dobrepolje, NHMW 112350/2 strongly corroded shells; Krain, Tekavcja jama, Debrepolje, coll. Edlauer ex coll. Kuščer, NHMW 48468/3; Tekarija jama b. Dobropolje, coll. S.H. Jaeckel ex coll. Kuščer, SMF 200961/3; Krain, Tekavčja jama, Dobrepolje, coll. Haas ex coll. Kuščer, SMF 53906/1; Krain, Berjakovo Brezno (= Malo Brezarjevo brezno, 46.080°N, 14.436°E), n. w. von Laibach, coll. Retaner (?), NHMW 12314/2; Berjakovo Brezno, nw v. Laibach, Krain, coll. Klemm ex coll. Edlauer, NHMW 79000/K/02938/2; Same data, coll. Schlickum, ex coll. Edlauer, SMF 275440/2; Berjakovo Brezno bei Dolnice n.w. von Ljubljana, coll. S.H. Jaeckel ex coll. Edlauer, SMF 200962/1; Radetina pećina, coll. Edlauer ex coll. Kuščer, NHMW 48751/6; Spodnja Skedevenica (at 45.854°N, 14.639°E), vel. Lašče, coll. Edlauer ex coll. Kuščer, NHMW 48883/15; Spodnja Skedevnica, coll. Edlauer ex coll. Kuščer, NHMW 48422/2; Spodnja Kedenrica, coll. W. Klemm ex coll. Kuščer, NHMW 33672/1 (figured shell, Fig. 8K-O); Sponja Skednevica bei Preserje, coll. Edlauer ex coll. Kuščer, NHMW 48124/2; Am Eingang der Doline Jama bei Kompole (=Kompoljska jama at 45.800°N, 14.731°E), S. Oest. von Dobrepolje, coll. Klemm, NHMW 112347/1; In der Raska Skedebnza bei Ponikve südl. von Dobrepolje (= Skedenc at 45.89687418°N, 14.50100919°E), coll. Klemm, NHMW 112348/3; Reoteiner Grotte bei Poduzik (?) bei Ljubljana (Kalkofen) (Podutiško brezno at 46.076°N, 14.445°E), coll. Edlauer ex coll. Kuščer, NHMW 49111/1; Brezarjeva brezno pri Boduzik (?), coll. Edlauer ex coll. Kuščer, NHMW 48390/2; Brezno v. Leskovi dolini (at 45.941°N, 14.256°E), coll. Edlauer ex coll. Kuščer, NHMW 49200/1; Mašun b Gratenbrunn (probably Jama na Mašunu at 45.630°N, 14.373°E), Krain, leg. Schollmayer, NHMW 112349/2 juvenile shells; Duplica (= Dupliška jama at 45.963°N, 14.684°E), coll. Edlauer ex coll. Kuščer, NHMW 48576/16; Jama v Jurčelovih (?) (Jama 1 v Jurcetovih Percah at 46.09984722°N, 14.42093707°E or Jama 2 na Jurcetovih Percah at 46.101°N 14.418°E, the first is indicated on the map), percah Tosko čelo, coll. Edlauer ex coll. Kuščer, NHMW 48688/16; Zakrita jama odd B (at 45.925°N, 14.293°E), coll. Edlauer ex coll. Kuščer, NHMW 48748/1 juvenile shell; Unterkrain: Grotte Thauzhia jama, coll. C. v. Heyden ex coll. N. Hoffmann, SMF 53903/1; Same data, coll. Gysser ex coll. Heynemann ex coll. C. v. Heyden, SMF 53901/2; Same data, coll. Heynemann ex coll. C. v. Heyden ex coll. N. Hoffmann, SMF 259469/3; Krain: Jeliarc-Grotte (maybe Jeliare or Jelice), coll. Th. Krüper, SMF 112877/1; Höhle Malo Bukuje bei Dobrova, Krain, coll. Dr. Leo Rušnov, ex coll. Dr. A. Oberwimmer, NHMW 71770/R/19 (1 photographed shell, Fig. 8F-J).</p> <p>Diagnosis.</p> <p>A medium sized species with very widely spaced, strong ribs, rounded, toothless aperture, and finely granular protoconch.</p> <p>Description.</p> <p>Spire somewhat elevated; protoconch consists of 1.5-1.75 whorls, very finely granulated, rather matte, not glossy; teleoconch with strong ribs that are supported by fine periostracal filaments in fresh shells; ribbing less regular than in other congeneric species; ribs widely spaced (41-52 ribs on body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75-4.25 whorls; aperture toothless, semilunar/rounded; peristome slightly thickened, slightly reflected in direction of umbilicus; umbilicus regular funnel-shaped, relatively narrow.</p> <p>Measurements. SW: 2.8-3.5 mm (median = 3.0 mm), SH: 1.7-1.9 mm (median = 1.8 mm), AW: 1.1-1.4 mm (median = 1.2 mm), AH: 1.0-1.3 mm (median = 1.1 mm), (n = 6; largest and smallest specimens of multiple populations measured).</p> <p>Differential diagnosis.</p> <p>The most similar taxon to Spelaeodiscus hauffeni is S. albanicus edentatus ssp. n. in terms of shell size, shape, and rib density. However, the latter one is usually larger, has a glossier protoconch, a less rounded aperture caused by the straighter basal part, and the peristome edge on the palatal side is more strongly expanded (rather thickened only in S. hauffeni).</p> <p>Variation among specimens.</p> <p>Spelaeodiscus hauffeni shows some variability in terms of shell size and spire height, but the rib density and the formation of the aperture are stable characters.</p> <p>Distribution.</p> <p>This species is distributed in the southeastern Alps (Central Slovenia) relatively far from the ranges of its congeneric taxa. Slapnik (2005) found shells in the Škocjan Caves Regional Park, which are probably of Holocene age (Figure 9).</p> <p>Conservation status.</p> <p>As there are several known locations and no reason to suppose that the habitat quality, habitat extent or population are deteriorating or extremely fluctuating, it was assessed as Least Concern (LC) by Reischütz (2017b).</p> </div>	http://treatment.plazi.org/id/EE99D18B1BD685F0BF2418D5757617C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
AE80E259B9B1F01056A44B67B31E0FA1.text	AE80E259B9B1F01056A44B67B31E0FA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus hunyadii Pall-Gergely & Deli 2018	<div><p>Spelaeodiscus hunyadii Pall-Gergely &amp; Deli sp. n. Figure 10A-E</p> <p>Type material.</p> <p>Montenegro, SE of Virpazar, 4.3 km (in a straight line) SSE of Ðuravci, near Besa/Bes near Krone i Besit, 330 m a.s.l., 42°08.548'N, 19°13.165'E (site code: 20171019G), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 19.10.2017, HNHM 103210 (holotype, SW: 2.1 mm, SH: 1.2 mm, Fig. 10A-E), DT/40 paratypes, EZP/40 paratypes, HA/40 paratypes, HNHM 103211/3 paratypes, PGB/40 paratypes, NMBE 554178/3 paratypes, NHMW 112367/4 paratypes; Montenegro, Donji Murići junction S 2 km along the Virpazar-Ostros road, 320 m a.s.l., 42°8.544'N, 19°13.182'E (site code: 2017/009), leg. Z.P. Erőss &amp; Z. Fehér, 16.07.2017, HNHM 103212/3 paratypes, EZP/3 paratypes.</p> <p>Type locality.</p> <p>Montenegro, SE of Virpazar, 4.3 km (in a straight line) SSE of Ðuravci, near Besa/Bes near Krone i Besit, 330 m a.s.l., 42°08.548'N, 19°13.165'E.</p> <p>Diagnosis.</p> <p>A small, nearly flat species with strong, widely spaced ribs, glossy protoconch and strongly oblique, toothless aperture.</p> <p>Description.</p> <p>Spire somewhat elevated; protoconch consists of 1.25-1.5 whorls, smooth, glossy; teleoconch with very strong (thick), equidistant, widely spaced ribs; rib density: 42-48 ribs on body whorl; between main ribs some fine wrinkles discernible; entire shell with 3.5-3.75 whorls; aperture semilunar, toothless, strongly oblique to shell axis; peristome slightly thickened and expanded in direction of umbilicus; umbilicus funnel-shaped, relatively narrow.</p> <p>Measurements. SW: 2.1-2.2 mm (median = 2.2 mm), SH: 1.2-1.3 mm (median = 1.2 mm), AW: 0.8-0.9 mm (median = 0.9 mm), AH: 0.6-0.7 mm (median = 0.7 mm), AA = 47-52°(n = 4; largest and smallest specimens measured).</p> <p>Differential diagnosis.</p> <p>Spelaeodiscus obodensis has a more conical shell and roughly sculptured protoconch. The most similar species to Spelaeodiscus hunyadii sp. n. is S. dejongi, which lives sympatrically with the new species. It differs from Spelaeodiscus hunyadii sp. n. in the less oblique aperture and the denser ribs. Some other populations of S. dejongi also possess widely spaced ribs, but their aperture is less oblique, thus, they can be distinguished from this new species. See also under S. latecostatus sp. n.</p> <p>Variation among specimens.</p> <p>Specimens of the type sample show no notable conchological variability.</p> <p>Etymology.</p> <p>This new species is named after our colleague and friend, András Hunyadi, who is one of those who first collected this species.</p> <p>Distribution.</p> <p>This species is known from the type locality only (Figure 7).</p> <p>Conservation status.</p> <p>To our present knowledge this species is very rare (currently known from a single location) and thus AOO is smaller than 20 km2. However, there is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number of mature individuals are declining or extremely fluctuating. Therefore, it might be assessed as Near Threatened (NT).</p></div> 	http://treatment.plazi.org/id/AE80E259B9B1F01056A44B67B31E0FA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
5237C950927075D50D41CF76B3A5E789.text	5237C950927075D50D41CF76B3A5E789.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus latecostatus Pall-Gergely & Eross 2018	<div><p>Spelaeodiscus latecostatus Pall-Gergely &amp; Eross sp. n. Figure 10F-O</p> <p>Type material.</p> <p>Montenegro, S of Virpazar, 0.8 km (in a straight line) E of Limljani, above the village, 400 m a.s.l., 42°11.698'N, 19°06.217'E (site code: 20171021B), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 21.10.2017, HNHM 103214 (holotype, Fig. 10A-E).</p> <p>Other material.</p> <p>Montenegro, Seoća S 1 km, along the <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.15&amp;materialsCitation.latitude=42.2103" title="Search Plazi for locations around (long 19.15/lat 42.2103)">Virpazar-Ostros</a> road, 280 m a.s.l., 42°12.618'N, 19°9.000'E (site code: 2017/005), leg. Z.P. Erőss &amp; Z. Fehér, 16.07.2017, HNHM 103213/3 shells, not paratypes (Fig. 10K-O).</p> <p>Type locality.</p> <p>Montenegro, S of Virpazar, 0.8 km (in a straight line) E of Limljani, above the village, 400 m a.s.l., 42°11.698'N, 19°06.217'E (site code: 20171021B).</p> <p>Diagnosis.</p> <p>A small, nearly flat species with strong, very widely spaced ribs, glossy protoconch and a toothless aperture.</p> <p>Description of the holotype.</p> <p>Spire somewhat elevated; protoconch consists of ca 1.25-1.5 whorls (the holotype is corroded at the protoconch-teleoconch junction), rather smooth, moderately glossy; teleoconch with very strong, equidistant, widely spaced ribs (42 on the body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75 whorls; aperture semilunar, toothless; peristome slightly thickened and expanded; umbilicus funnel-shaped, relatively narrow.</p> <p>Measurements. SW: 2.2 mm, SH: 1.2 mm, AW: 0.8 mm, AH: ca. 0.8 mm, AA = 63°(holotype).</p> <p>Differential diagnosis.</p> <p>The widely spaced ribs are similar to S. hunyadii sp. n., but the less oblique aperture distinguishes S. latecostatus sp. n. from the other new species. Spelaeodiscus dejongi, which lives sympatrically with S. latecostatus sp. n., is similar in shell shape and size and the formation of the aperture, but has much denser ribs.</p> <p>Variation among specimens.</p> <p>See remarks.</p> <p>Etymology.</p> <p>This new species is named after its remarkably widely spaced ribs.</p> <p>Distribution.</p> <p>See under Remarks and Figure 7.</p> <p>Remarks.</p> <p>The holotype of this species was found in a large sample of S. dejongi. Therefore, even if the shell shape does not differ from that species, the widely spaced ribs indicate that S. latecostatus sp. n. differs from S. dejongi on species level. Three shells from 1 km S of Seoća possess denser ribs than other S. dejongi populations (47-54 ribs on the body whorl), but obviously the rib density is lower than that of the holotype of S. latecostatus sp. n. Since the rib density of that population is intermediate between S. dejongi and S. latecostatus sp. n., it is not possible to decide which species it belongs to. More populations around the sample from 1 km S of Seoća site are necessary in order to provide a reliable identification. Here we provisionally identify those shells as Spelaeodiscus cf. latecostatus sp. n.</p> <p>Conservation status.</p> <p>To our present knowledge this species is very rare (currently known from two locations) and thus AOO is smaller than 20 km2. However, there is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number of mature individuals are declining or extremely fluctuating. Therefore, it might be assessed as Near Threatened (NT).</p></div> 	http://treatment.plazi.org/id/5237C950927075D50D41CF76B3A5E789	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
4CCB7D685CD1F8B5B817DB16E619B447.text	4CCB7D685CD1F8B5B817DB16E619B447.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus obodensis Bole 1965	<div><p>Spelaeodiscus obodensis Bole, 1965 Figures 11, 12</p> <p>Spelaeodiscus obodensis Bole, 1965: 350, plate 76, fig. D.</p> <p>Spelaeodiscus (Spelaeodiscus) obodensis - Gittenberger 1969: 296-297, fig. 1.</p> <p>Spelaeodiscus obodensis - Welter-Schultes 2012: 213. (partim: the photos show S. unidentatus specimens)</p> <p>Type material.</p> <p>Obodska pećina, Rij. Crnojevića, Mtg., September 1956, MZBI 1018 (2 adult and 3 juvenile/broken syntypes, photos were examined, Figure 12A).</p> <p>Material examined.</p> <p>Vodna jama (Lovćen), coll. Edlauer ex coll. Kuščer, NHMW 48543/2 (det. Gittenberger, 1973 Sep.); Same data, NHMW 49765/2; Montenegro, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.029066&amp;materialsCitation.latitude=42.675568" title="Search Plazi for locations around (long 19.029066/lat 42.675568)">Ostrog</a>, rocks below the upper parking lot, 820 m a.s.l., 42°40.534'N, 19°1.744'E, leg. A. Reischütz, P. L. Reischütz &amp; N. Steiner-Reischütz, May 2017, REI/1; Montenegro, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.027718&amp;materialsCitation.latitude=42.675167" title="Search Plazi for locations around (long 19.027718/lat 42.675167)">Ostrog</a>, entrance of the penultimate parking lot, 770 m a.s.l., 42°40.510'N, 19°1.663'E, leg. A. Reischütz, P. L. Reischütz &amp; N. Steiner-Reischütz, May 2017, NHMW 112365 (photographed shell, Fig. 11F-J), REI/1; Montenegro, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.036467&amp;materialsCitation.latitude=42.6583" title="Search Plazi for locations around (long 19.036467/lat 42.6583)">Ostrog</a>, 1 km north from Hotel Sokoline on the new street to Podgorica, 630 m a.s.l., 42°39.498'N, 19°2.188'E, leg. A. Reischütz, P. L. Reischütz &amp; N. Steiner-Reischütz, May 2017, REI/2; Montenegro, 500 m south of the bridge over the <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.180416&amp;materialsCitation.latitude=42.457882" title="Search Plazi for locations around (long 19.180416/lat 42.457882)">Sitnica river</a>, north of Podgorica, 45 m a.s.l., 42°27.473'N, 19°10.825'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Apr. 2012, NHMW 112366/1 (photographed shell, Fig. 11K-O), REI/2; Yugoslavia: Pečina u Peckom Brdu bg., above Začir, leg. L. Pintér &amp; P. Subai, 20 Jul. 1972, HNHM 41127/2; Same data, HNHM 18062/3 (one of them is photographed, Fig. 11A-E); Montenegro, Pečina u peckom Brdu bei Začir (Brdu-cave at Začir), leg. H. Schütt, 06.06.1978, HNHM 42402/3.</p> <p>Diagnosis.</p> <p>A medium sized species with elevated spire, roughly sculptured protoconch, strong ribs on the teleoconch, and toothless aperture.</p> <p>Description.</p> <p>Spire elevated, shell low conical; protoconch consists of 1.25-1.5 whorls, roughly granulated/ ”hammered”, matte, not glossy; teleoconch with strong, equidistant ribs that are supported by fine periostracal filaments in fresh shells; rib density variable (43-76 ribs on body whorl), usually widely spaced; between main ribs some fine wrinkles discernible; entire shell with 3.75-4 whorls; aperture semilunar, toothless; peristome slightly thickened and expanded, especially in direction of the umbilicus; umbilicus funnel-shaped, relatively narrow.</p> <p>Measurements. SW: 2.6-3.0 mm (median = 2.8 mm), SH: 1.6-1.9 mm (median = 1.7 mm), AW: 0.9-1.2 mm (median = 1.1 mm), AH: 1.0-1.2 mm (median = 1.1 mm), AA = 64-70°(n = 10; largest and smallest specimens of multiple populations measured).</p> <p>Differential diagnosis.</p> <p>The most similar species is S. dejongi, which usually has a lower spire, weaker ribs, and a glossy protoconch.</p> <p>Variation among specimens.</p> <p>This species is the most variable in terms of shell size and rib density.</p> <p>Distribution.</p> <p>This species is found northwest of the Shkodër Lake Basin, as well as in the Zeta River Valley between Podgorica and the Ostrog Monastery (Figure 4). The Albanian record given by Reischütz et al. (2013) and referred by Reischütz (2017c) is actually S. albanicus edentatus ssp. n. So far, no Albanian occurrence is known.</p> <p>Conservation status.</p> <p>As there are at least seven known locations and no reason to suppose that the habitat quality, habitat extent or population are deteriorating or extremely fluctuating, it was assessed as Least Concern (LC) by Reischütz (2017c).</p> </div>	http://treatment.plazi.org/id/4CCB7D685CD1F8B5B817DB16E619B447	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
36EF9D32E0A8CA5100DA999403D10A99.text	36EF9D32E0A8CA5100DA999403D10A99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus unidentatus Bole 1961	<div><p>Spelaeodiscus unidentatus Bole, 1961</p> <p>Diagnosis.</p> <p>A small to medium sized species with dense riblets, and strong parietal and basal teeth/thickenings.</p> <p>Differential diagnosis.</p> <p>This species differs from most other Spelaeodiscus species by the presence of two well-developed apertural teeth (a palatal and a basal). See also under Spelaeodiscus albanicus.</p> <p>Conservation status.</p> <p>As there are more than five known locations and no reason to suppose that the habitat quality, habitat extent or population are deteriorating or extremely fluctuating, it was assessed as Least Concern (LC) by Reischütz (2017d).</p> </div>	http://treatment.plazi.org/id/36EF9D32E0A8CA5100DA999403D10A99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
F885B3E00F6762818BEFE50D0A1AFC3B.text	F885B3E00F6762818BEFE50D0A1AFC3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus unidentatus subsp. acutus Pall-Gergely & Feher 2018	<div><p>Spelaeodiscus unidentatus acutus Pall-Gergely &amp; Feher ssp. n. Figure 14A-L</p> <p>Spelaeodiscus unidentatus - Reischütz et al. 2013: 62, fig. 3.</p> <p>Type material.</p> <p>Albania, rocks north of Hajmel at the bridge, north of Lezhe, 40 m a.s.l., 41°58.426'N, 19°38.683'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Apr. 2012, NHMW 112371/1 (photographed paratype, Fig. 14G-L), REI/6 paratypes; Albania, rocks 1 km south of Hajmel, north of Lezhe, 50 m a.s.l., 41°56.848'N, 19°38.379'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Apr. 2012, REI/5 paratypes; Albania, Shkodër District, Vau i Dejës, Mjedë, near the dam, 45 m a.s.l., limestone rocks, 42°0.804'N, 19°37.188'E (site code: 2015/99), leg. Z.P. Erőss, Z. Fehér &amp; J. Grego, 04 Jul. 2015, HNHM 103252 (holotype, SW: 2.9 mm, SH: 1.5 mm, Figs 14A-F), NHMW 112372/1 paratype, JG/1 paratype.</p> <p>Type locality.</p> <p>Albania, Shkodër District, Vau i Dejës, Mjedë, near the dam, 45 m a.s.l., limestone rocks, 42°0.804'N, 19°37.188'E.</p> <p>Diagnosis.</p> <p>Basal tooth pointed; palatal part of peristome with slight incision at the position of the palatal tooth, palatal region not or slightly “pushed” from outside.</p> <p>Measurements. SW: 2.9-3.5 mm (median = 3.0 mm), SH: 1.4-1.8 mm (median = 1.5 mm), AW: 1.1-1.3 mm (1.2 mm), AH: 1.1-1.4 mm (median = 1.2 mm), (n = 9; largest and smallest specimens of multiple populations measured).</p> <p>Differential diagnosis.</p> <p>Spelaeodiscus unidentatus acutus ssp. n. differs from the nominotypical subspecies by the strong, pointed basal tooth, which is blunt, or represented as a thickening of the basal peristome in the nominotypical subspecies. Moreover, the palatal region is not or only slightly “pushed” from the outside in the position of the palatal tooth. Rib density: 64-91 ribs on the body whorl.</p> <p>Variation among specimens.</p> <p>This subspecies is variable in terms of shell size and spire height, i.e. one population has a nearly flat shell.</p> <p>Etymology.</p> <p>This new subspecies is named after its pointed (Latin: acutus) basal tooth, which distinguishes it from the nominotypical subspecies.</p> <p>Distribution.</p> <p>This subspecies is known from the southeastern side of the Shkodër Lake Basin in Albania (Figure 7).</p> </div>	http://treatment.plazi.org/id/F885B3E00F6762818BEFE50D0A1AFC3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
557123F26FE62D66CECD1A174B4C427A.text	557123F26FE62D66CECD1A174B4C427A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus unidentatus subsp. unidentatus unidentatus Bole 1961	<div><p>Spelaeodiscus unidentatus unidentatus Bole, 1961 Figure 13</p> <p>Spelaeodiscus unidentatus Bole, 1961: 205 fig. 1a-d.</p> <p>Spelaeodiscus unidentatus - Bole 1965: 350-351, fig. 1a, 2b, plate 76, fig. C.</p> <p>Spelaeodiscus (Spelaeodiscus) unidentatus - Gittenberger 1969: 297, fig. 1.</p> <p>Spelaeodiscus unidentatus - Welter-Schultes 2012: 214.</p> <p>Type material.</p> <p>Obodska pećina, Rij. Crnojevića, Mtg., 1956 September, MZBI 1019/6 adult syntypes and some juvenile syntypes (photos of the sample where examined, Figure 12B). See also Remarks.</p> <p>Other material.</p> <p>Bioče, N Titograd, Mtg. 23.09.1978, MZBI 15443 (photos of the sample where examined); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Megara</a>, Toluši, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Titograd</a>, Mtg. MZBI 2335 (photos of the sample where examined); Pećina od <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Zavora</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Peuta</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Titograd</a>, Mtg. 01.11.1963, MZBI 2338 (photos of the sample where examined); Höhle bei <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Virpazar</a>, leg. Dabović, coll. Edlauer, NHMW 48881/1; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Brosine cave</a> (33), leg. Kuscer 10266/67, coll. Edlauer, NHMW 38874/a (20 juvenile shells, "schlechte <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Expl</a> "); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Same</a> data, NHMW 58874/4 ("bessere <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Expl</a> "); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Virpazar</a>, mit "oculus mundi", leg. Dabović, coll. Edlauer, NHMW 78988/3 juvenile shells (det. Gittenberger, but identification not certain); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Smarjetna</a> gora + <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Grabocica cave</a> (drawing of a cave on the label), (<a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Trnovo</a>), NHMW 78989/7; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Same</a> data, NHMW 48086/5; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Vodna</a> jama (Lovćen), coll. Edlauer ex coll. Kuščer, NHMW 48544/1; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Same</a> locality, coll. Edlauer, NHMW 49764/1 (labelled as " Spelaeodiscus albanicus hadzii "); Montenegro, NNW of Virpazar, road between Rijeka Crnojevića and Virpazar, 0.4 km (in a straight line) W of Poseljani, 130 m a.s.l., 42°18.361'N, 19°02.898'E (site code: 20171020F), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 20.10.2017, DT/ca. 15, EZP/ca. 15, HA/ca. 15, HNHM 103248/5, PGB/ca. 15; Montenegro, W of Rijeka Crnojevića, 1 km (in a straight line) NNE of Zaćir, above Obodska pećina, near the road, 320 m a.s.l., 42°21.168'N, 19°00.125'E (site code: 2017.10.20J), leg. T. Deli, Z.P. Erőss, A. Hunyadi &amp; B. Páll-Gergely, 20.10.2017, DT/ca. 18, EZP/ca. 18, HA/ca. 18, HNHM 103249/5, PGB/ca. 18; Montenegro, 2 km north of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Velje Brdo</a> in the <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Zeta valley</a>, north of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Podgorica</a>, 70 m a.s.l., 42°28.932'N, 19°14.454'E, leg. A. Reischütz, P. L. Reischütz &amp; N. Steiner-Reischütz, May 2016, REI/5; Montenegro, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Moraca valley</a> , after the exit to the "China Road and Bridge" camp N of Bioće, 70 m a.s.l., 42°29.167'N, 19°18.646'E, leg. A. Reischütz, P. L. Reischütz &amp; N. Steiner-Reischütz, May 2016, NHMW 112368/1 (photographed shell, Fig. 13G-L), REI/10; Montenegro, road between Rijeka Crnojevića and Hrvasi, 180 m a.s.l., 42°21.491'N, 19°4.281'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Apr. 2012, REI/11; Montenegro, scree slope at the BMS-petrol station, south of the Morača gorge, 5 km south of Bioće, 90 m a.s.l., 42°29.064'N, 19°18.279'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Jul. 2008, REI/1; Montenegro, spring 1 km south of the junction towards Njive, south of Rijeka Crnojevića, 150 m a.s.l., 42°18.266'N, 19°2.900'E, leg. A. Reischütz, N. Reischütz &amp; P. L. Reischütz, Mar. 2011, NHMW 112369/1 (photographed shell, Fig. 13M-R); Montenegro, Žabljak Crnojevića, fortress, 50 m a.s.l., 42°19.030'N, 19°9.401'E (site code: 2015/007), leg. T. Deli, Z.P. Erőss &amp; Z. Fehér, 25.05.2015, HNHM 103250/1 adult and several juvenile shells; Montenegro, Začir, “Pečina u pecko brdo" (cave), 450 m a.s.l., leg. L. Pintér, P. Subai, A. Szigeti, 20 Jul. 1972, NMBE 542072/76; Montenegro, Pečina u peckom, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Brdu-bg Zacirnal</a> (<a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Brdu-cave</a> at Začir), leg. L. Pintér &amp; P. Subai, 20 Jul. 1972, HNHM 65187/7 (one of them is photographed: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Figs</a> 13A-F); Montenegro, Pečina u peckom <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Brdu</a> bei Začir (<a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Brdu-cave</a> at Začir), leg. H. Schütt, 06.06.1978, coll. Gy. Kovács, HNHM 65188/1; <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Same</a> data, HNHM 42401/2; Montenegro, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Rijeka Crnojevica</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.048067&amp;materialsCitation.latitude=42.305782" title="Search Plazi for locations around (long 19.048067/lat 42.305782)">Obodska cave</a>, 110 m a.s.l., 42°21.120'N, 19°0.318'E, leg. P. Subai &amp; M. Szekeres, 23 Sep. 2006; NMBE 542071/1; Montenegro, Poseljani, on the Virpazar-Rijeka Crnojevića road, 150 m a.s.l., 42°18.347'N, 19°2.884'E (site code: 2015/016), leg. T. Deli, Z.P. Erőss &amp; Z. Fehér, 25.05.2015, HNHM 103251/8; NHMW 112370/8, DT/8.</p> <p>Diagnosis.</p> <p>Basal tooth/thickening low, not pointed; palatal part of peristome with strong incision at the position of the palatal tooth, palatal region strongly “pushed” from outside.</p> <p>Description.</p> <p>Spire elevated; protoconch consists of 1.5 whorls, roughly granulated/ ”hammered”, matte, not glossy; teleoconch with strong but dense, equidistant ribs that are supported by fine periostracal filaments in fresh shells; rib density variable (74-118 ribs on body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.75-4.5 whorls; aperture semilunar or triangular due to straight basal part; peristome thickened, slightly expanded and slightly reflected on the palatal, basal and umbilical areas; palatal tooth strong, pointed; palatal region of peristome with strong outer incision (i.e. the position of the palatal tooth is indicated with a groove on the outer side); basal portion of peristome straight, thickened; occasionally two small denticles visible, sometimes only the one situated closer to the palatal side is developed as a low denticle; umbilicus funnel-shaped, wide to relatively narrow (width depends on spire height).</p> <p>Measurements. SW: 2.4-3.2 mm (median = 2.7 mm), SH: 1.3-1.8 mm (median = 1.5 mm), AW: 0.9-1.1 mm (median = 1.0 mm), AH: 1.0-1.4 mm (median = 1.1 mm), (n = 11; largest and smallest specimens of multiple populations measured).</p> <p>Differential diagnosis.</p> <p>See under S. unidentatus acutus ssp. n.</p> <p>Variation among specimens.</p> <p>This subspecies is particularly variable in terms of shell size. The shape of the thickening of the basal part of the peristome is also slightly variable, although it never develops to a pointed tooth.</p> <p>Distribution.</p> <p>This taxon is found north and northwest of the Shkodër Lake Basin in Montenegro (Figure 7). Albanian records given by Reischütz et al. (2013) actually refer to S. unidentatus acutus ssp. n.</p> <p>Remarks.</p> <p>The only sample in the MZBI, which was collected before the original description is from Obodska pećina represent the type sample of this species (MZBI 1019). Bole collected both S. obodensis and S. unidentatus at Obodska pećina in September, 1956, but described only one (S. unidentatus) in 1961, and the other (S. obodensis) in 1965.</p> <p>Formerly, Peuta Cave population was incorrectly referred to as S. albanicus and was taken into consideration in that species’ Red List assessment (Reischütz and Fehér 2017).</p> </div>	http://treatment.plazi.org/id/557123F26FE62D66CECD1A174B4C427A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
3CE3B48E8A328FFAF621E885FDBE1638.text	3CE3B48E8A328FFAF621E885FDBE1638.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeodiscus virpazarioides Pall-Gergely & Feher 2018	<div><p>Spelaeodiscus virpazarioides Pall-Gergely &amp; Feher sp. n. Figure 14M-R</p> <p>Type material.</p> <p>Albania, Malësia district, a mountain pass 2 km N of Rraps-Starjë, 700 m a.s.l., 42°24.888'N, 19°30.240'E, leg. Erőss, Fehér, Szekeres, Grego, 27.06.2016, HNHM 103417 (holotype, SW = 3.45 mm, SH = 2.0 mm), HNHM 102765/4+1subadult+3fr (paratypes), NHMW 111672/3+2subadult+2fr; JG/3+2subadult+3fr (paratypes); same locality, leg. A. Reischütz, N. Reischütz &amp; P. Reischütz, Apr. 2006, REI/1 juvenile paratype.</p> <p>Type locality.</p> <p>Albania, Malësia district, a mountain pass 2 km N of Rraps-Starjë, 700 m a.s.l., 42°24.888'N, 19°30.240'E.</p> <p>Diagnosis.</p> <p>A medium sized to large species with elevated spire, strong, very widely spaced ribs, fine spiral lines consisting of series of minute tubercles, hammered protoconch, weak basal thickening, weak parietal tooth, and a thickened parietal callus.</p> <p>Description.</p> <p>Spire elevated; protoconch consists of ca. 1.25-1.75 whorls, roughly sculptured, “hammered”; teleoconch with very strong, equidistant, widely spaced ribs (40-70 on the body whorl; addition to the radial ribs very fine, dense spiral striation is visible on the entire teleoconch, consisting of minute tubercles; entire shell with 4-4.75 whorls; aperture subcircular/triangular; basal part with two low swellings (similar to those of S. albanicus albanicus), parietal wall with a weak tooth; parietal callus thickened, blunt; umbilicus funnel-shaped, relatively narrow.</p> <p>Measurements. SW: 3.3-3.6 mm (median = 3.5 mm), SH: 1.95-2.25 mm (median = 2.1 mm), AW: 1.3-1.6 mm (median = 1.4 mm), SW: 1.4-1.5 mm (median = 1.4 mm), (n = 11).</p> <p>Differential diagnosis.</p> <p>This new species can be distinguished from all congeners (especially the most similar S. albanicus albanicus) by the clearly visible spiral striation, roughly sculptured protoconch and thickened parietal callus. All Virpazaria species possess an elevated parietal callus, but it is sharp in most (all?) species, and the aperture of those species are more slender, crescent-shaped.</p> <p>Variation among specimens.</p> <p>The degree of the thickness of the parietal callus shows some recognisable variability within the single known population, but this trait might be due to the age (degree of development) of the examined shells.</p> <p>Etymology.</p> <p>This new species is named after its resemblance to Virpazaria species based on the thickened parietal callus.</p> <p>Distribution.</p> <p>Spelaeodiscus virpazarioides sp. n. is known from the type locality only (see also Figure 7).</p> <p>Remarks.</p> <p>Initially we considered placing this species to the genus Virpazaria due to the thickened parietal callus, which was mentioned in the original description of the genus Virpazaria. However, the subcircular/triangular aperture indicates that it is better to be placed in Spelaeodiscus.</p> <p>Formerly, this population was incorrectly referred to as S. albanicus and was taken into consideration in that species’ Red List assessment (Reischütz and Fehér 2017)</p> <p>Conservation status.</p> <p>To our present knowledge this species is very rare (currently known from a single location) and thus AOO is smaller than 20 km2. However, there is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number of mature individuals are declining or extremely fluctuating. Therefore, it might be assessed as Near Threatened (NT).</p></div> 	http://treatment.plazi.org/id/3CE3B48E8A328FFAF621E885FDBE1638	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pall-Gergely, Barna;Deli, Tamas;Eross, Zoltan Peter;Reischuetz, Peter L.;Reischuetz, Alexander;Feher, Zoltan	Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander, Feher, Zoltan (2018): Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13-48, DOI: http://dx.doi.org/10.3897/zookeys.769.25258, URL: http://dx.doi.org/10.3897/zookeys.769.25258
