taxonID	type	description	language	source
6832D7BD71CAC4ADD1146410209988D5.taxon	materials_examined	Material examined. Holotype 1 ♂, United Arab Emirates, Wadi Maidaq, 460 m, 12. IV. 2006, leg. C. Gielis, GP. 5787 Asb, in Naturalis Biodiversity Center, Leiden; Paratypes, 1 ♂, United Arab Emirates, 10 km. NE. Huwaylat, 18. IV. 2006, leg. J. Buszko, GP 6234 Asb, 1 ♂ same data in Naturalis Biodiversity Center, Leiden. Holotype and Paratypes were examined by the second author. Diagnosis. Acteniopsis robustus was described by Asselbergs (2010) based on three male specimens. Examination of the types of A. robustus revealed that this species is almost similar to A. kurdistanella and the newly described species superficially; however, shares a variety of characters with other members of the genus Stemmatophora including: (1) very long maxillary palpi (more than three-fifth of the length of labial palpi and half times the horizontal diameter of compound eye in Paratypes) compared with very small and hardly recognizable maxillary palpi in A. kurdistanella and A. gambronensis; (2) long and relatively slender labial palpi in contrast to shorter and broadly scaled labial palpi of A. kurdistanella and A. gambronensis; (3) ciliae at the ventral surface antennae, consist of two pairs of plumose ciliae on each flagellum in A. robustus, while in A. kurdistanella and A. gambronensis they form a transverse row of at least ten ciliae (Fig. 4 C). There are some other differences between the male genitalia of A. robustus, A. kurdistanella and A. gambronensis: 1) Slender, rod-shaped cornutus of A. robustus (Fig. 2 F) compared with the stouter and anchor-shaped cornuti of A. kurdistanella and A. gambronensis (Figs 2 D, E). 2) In both A. kurdistanella and A. gambronensis the vinculum is basally pointed (Figs 2 A, B), while in A. robustus it is basally rounded. 3) In A. kurdistanellus and A. gambronensis the gnathos does not exceed the uncus (Fig. 2 A) or slightly exceeds it (Fig. 2 B), whereas in A. robustus it exceeds clearly. 4) The valva in A. robustus is slightly curved posteriorly, whereas in A. kurdistanella and A. gambronensis it is straight (Figs 2 A, B). It should be noted that Acteniopsis robustus has the largest wingspan (22 - 24 mm) compared with A. kurdistanella (17 - 18 mm) and A. gambronensis (13.7 - 17.0 mm). In view of the above, we hereby transfer the species A. robustus from Acteniopsis Amsel, 1959 to Stemmatophora Guenee, 1854.	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
4A2DE0FA3FF1D84810E0A940E2418793.taxon	materials_examined	Material examined. Holotype ♂, Iran, Hormozgan Prov.: Isin, Geno, 750 m a. s. l., 25. IV. 1986, Mirzayans, Borumand leg.; Paratypes 2 ♂♂ 3 ♀♀, same data as the Holotype. Deposited at HMIM, IRIPP, Tehran, Iran (GS: HA- 2151, HA- 2307, HA- 2309).	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
4A2DE0FA3FF1D84810E0A940E2418793.taxon	diagnosis	Diagnosis. Both Acteniopsis gambronensis and A. kurdistanella are similar to each other in wing venation and shape of the maxillary palpi and labial palpi; however their wing pattern is slightly different. In A. gambronensis, the antemedian and postmedian lines are clearly visible, while in A. kurdistanella the antemedian line is very weak and sometimes hardly distinguishable. In A. kurdistanella the two transverse lines are reddish-brown, but these lines are white in A. gambronensis. The forewing of A. kurdistanella is pale clay-coloured, while in A. gambronensis the ground colour is pale-ochreous to yellowish-ochre (Figs 1 C, D). Although these two species are similar in the shape of the male genitalia, there are some differences: 1) In A. gambronensis the phallus is straight and cleaver-shaped (its width in the proximal half is half length of its width in the distal part) (Fig. 2 A), while in A. kurdistanella it is slightly curved and is equally wide throughout its length (Fig. 2 B). 2) In A. gambronensis the gnathos does not extend to the distal margin of the uncus and is almost straight apically (Fig. 2 A), but in A. kurdistanella it reaches the distal margin or is slightly exceeding it with a small hook at the apex (Figs 2 B, C). 3) Although as stated by Amsel (1959), in A. kurdistanella the uncus has parallel margins, in all the examined Iranian A. kurdistanella specimens, it was slightly constricted basally (Fig. 2 B); while in A. gambronensis the width of both proximal and distal ends were the same (Fig. 2 A). 4) The longer arm of anchor-shaped cornutus in A. gambronensis is almost finger-shaped (Figs 2 A, D), but in A. kurdistanella is tapering to a point (Figs 2 B, E). Examination of the female genitalia of A. gambronensis and the newly described genitalia of A. kurdistanella reveals that the latter species is very close to A. gambronensis in this regard. The only differences of these two species are as below: 1) In A. gambronensis the ductus bursae is relatively wide and bulbous near the junction with the antrum, with an almost knee-shaped bend (Figs 3 A, C), whilst in A. kurdistanella it is slightly narrowed posteriorly at the junction with the antrum, with an upturned bend (Figs 3 B, D). 2) In A. gambronensis the folded area of the antrum is clearly extended to the posterior end of ductus bursae (Figs 3 A, C), whereas in A. kurdistanella this area is slightly extended to the posterior end of the ductus bursae (Figs 3 B, D). Description. Female slightly larger than the male. Forewing length, males x ¯ = 6.5 mm + / - 0.5 (n = 3), females x ¯ = 7.1 mm + / - 0.56 (n = 3), Holotype, 6.5 mm. Head (Figs 1 A, B). Chaetosemata and ocelli present; frons smooth; frons and vertex covered with relatively wide, white smooth scales (sometimes tinged with pale yellow scales); occipital area roofed with relatively narrow and slightly appressed white scales; collar covered with relatively wide uniformly yellowish-cream scales; labial palpi porrect with slightly drooping apex, its length 1.45 - 1.70 times the horizontal diameter of compound eye (n = 6), second segment the longest, first segment almost two-thirds of the second one and apical segment very short, slightly more than one-fifth of the second segment. Labial palpi covered with light-ochreous scales tinged with slightly to much darker scales latero-ventrally, except for distal one-fifth (or slightly less than this) which is white dorso-laterally (Fig. 1 B), all the segments white internally except for darker ventro-marginal sides; maxillary palpi short (slightly more than one-fifth of the length of labial palpi), covered with pale-ochreous scales laterally; proboscis covered with light ochre to ochreous-creamy scales basally; antennae annulated with alternative light brown and creamy scales dorsally, and ciliated ventrally. In male, each flagellum with a transverse row of at least ten long ciliae ventrally, the lateral ciliae the longest, length of the longest cilia almost two times the width of shaft (Figs 4 C, E); in female each with a row of several short ventral ciliae (length of cilia less that the width of the shaft). Length of the antennae slightly more than half length of the forewing. Thorax, tegulae, and abdomen covered with yellowish-creamy and creamy scales, respectively. Forewing (Fig. 1 C) an elongated triangle, ground colour pale-ochreous to yellowish-ochre with two white antemedian and postmedian transverse lines, the antemedian line from the base at slightly beyond one-fifth of the forewing length, edged with darker scales externally, and straight to slightly convex medially, postmedian line from the base to slightly behind four-fifth of the forewing length, edged with darker scales internally, slightly convexed near to costal margin and then slightly concaved at two-thirds of its length from costal margin. The area between the two transverse lines slightly darker than the ground colour, and without discocellular stigma. Costal margin from the base to postmedian line with nine narrow, brown longitudinal stripes with different length at the upper as well as at the underside, alternating with short white stripes. Fringes creamy-white with some scattered pale-ochreous scales. Hindwing (Fig. 1 C) creamy-white with some scattered light-brown scales concentrated in the middle part and absent both at costal and internal margins, and some scattered light-brown scales at the internal margin of termen. In some species these scales are hardly visible. Fringes in the same colour as the hindwing. Underside of the both fore- and hindwing yellowish-creamy, but in the forewing with scattered pale-ochreous scales and some pale-ochreous ones at the area of postmedian line, close to the costal margin. Male genitalia (Figs 2 A, D) (n = 1). Uncus rectangular and apically rounded, with the same width throughout the length; free end of gnathos lanceolate, apically without clear hook; tegumen relatively wide, its width two times the width of uncus; valva nearly straight, slightly narrowed distally, proximal end of valva at costal margin with a small semicircular plate; sacculus prominent; width of juxta slightly decreasing distally; vinculum almost triangular with pointed tip; phallus cleaver-shaped, width of the basal half nearly half length of distal half, slightly shorter than the length of valva, with orifice of vesica positioned mid-ventrally, cornutus as an asymmetrical anchor-shaped sclerotized structure, and some minute spines arranged distally. Female genitalia (Figs 3 A, C) (n = 2). Papillae anales almost semi-ovoid with rounded tip, covered with both short and relatively long hairs; apophyses posteriores very long, 2.77 - 2.85 times the length of apophyses anteriores, and slightly narrowed posteriorly, apophyses anteriores slightly thicker than apophyses posteriores and slightly curved posteriorly; ostium bursae very small and rounded, positioned at the bottom of a cavity produced by membranous area between eighth and seventh abdominal sternite; antrum membranous, short and tube-shaped, surrounded by folded walls on lateral sides. The folded area clearly extended to the posterior end of ductus bursae; colliculum absent; ductus bursae relatively wide, bulbous near the junction with antrum, with an almost knee-shaped bend at this area; ductus seminalis originates from the bulbous area of ductus bursae in the back (Figs 3 A, C); border of ductus bursae and corpus bursae more or less indistinguishable; corpus bursae pear-shaped and without signum; eighth abdominal tergite with a distinct triangular plate in the middle; length of the seventh abdominal segment almost twice the eighth segment.	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
4A2DE0FA3FF1D84810E0A940E2418793.taxon	distribution	Distribution. Iran: Hormozgan Province (Isin).	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
4A2DE0FA3FF1D84810E0A940E2418793.taxon	etymology	Etymology. Isin (the type locality of the new species) is a rural district in the central district of Bandar Abbas County, Hormozgan Province, and this species is named according to the old name of Bandar Abbas, i. e. " Gambron ".	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
4A2DE0FA3FF1D84810E0A940E2418793.taxon	biology_ecology	Biology. Unknown.	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
36B296FBDC90160BC4C48BD7328027CE.taxon	materials_examined	Material examined. Iran, Ardebil Prov.: 1 ♂, Khalkhal, Hashtchin- Khalkhal Rd., N 37 ° 26 ' 54.5 ", E 048 ° 54 ' 50.6 ", 837 m, 16. VII. 2013, Afsarian, Sarafrazi leg.; Chaharmahal va Bakhtiari Prov.: 1 ♂ 1 ♀, Chelgerd, 2500 m, 17, 18. viii. 1995, Mirzayans, Badii leg., 2 ♂♂ 1 ♀, Lordegan (Oak forest), N 31 ° 32 ' 30 ", E 050 ° 57 ' 16.92 ", 2352 m, 15. VIII. 2010, Alipanah, Nematian leg.; Fars Prov.: 1 ♂, Sepidan, Margoon waterfall, 1910 m, 28. ix. 1996, Barari, Parchami-Araghi, Moghaddam leg., 1 ♀, Kamfiruz, Tang-e Bostanak, 1730 - 1900 m, 17. VIII. 2000, Badii, Moghaddam, Mofidi-Neyestanak leg.; Hamedan Prov.: 2 ♂♂ 1 ♀, Khan Gormaz protected area, N 34 ° 35 ' 17.8 ", E 048 ° 10 ' 39.5 ", 2047 m, 9. VII. 2014, Afsarian leg.; Hormozgan Prov.: 1 ♂, Bandar Abbas, Kahurestan, 17. iii. 1971, 15 m, Ayatollahi, Pazuki leg.; Kermanshah Prov.: 1 ♀, Chalabeh, Bisotun, N 34 ° 22 ' 57.2 ", E 47 ° 16 ' 38.7 ", 1652 m, 12. VII. 2014, Afsarian leg.; Kohgiluyeh va Boyerahmad Prov.: 2 ♂♂, Tang-e Meymand, 1650 m, 9. ix. 1971, Ebrahimi, Badii leg.; Kordestan Prov.: Marivan, 26. vi. 2004, Mansurghazi leg.; Markazi Prov.: 1 ♂, Salafchegan, Dizijan, 1550 m, 29. vii. 1997, Barari, Mofidi-Neyestanak leg.; Zanjan Prov.: 1 ♂, Zanjan, Abbar, N 36 ° 56 ' 19.2 ", E 48 ° 59 ' 32 89 ", 1156 m, 21. vii. 2010, Alipanah leg. (GS: HA- 2308, HA- 2310).	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
36B296FBDC90160BC4C48BD7328027CE.taxon	diagnosis	Diagnosis. The female genitalia of this species is very close to that of A. gambronensis, and their main differences are in the width of ductus bursae and shape of the posterior end of ductus bursae (see diagnosis of A. gambronensis).	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
36B296FBDC90160BC4C48BD7328027CE.taxon	distribution	Distribution. Iraq (Haji Omran; the type locality); Iran (Saadatabad, Sine Sefid, Mian Kotal, Komehr, Karaj, Sultanabad) (Amsel 1961).	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
36B296FBDC90160BC4C48BD7328027CE.taxon	discussion	Discussion. As mentioned above, the two genera Acteniopsis and Stemmatophora have some differences in wing venations and ciliae on the ventral surface of the antennae; however our findings are somewhat different from those given by Amsel (1959). As we observed, in the forewing of males and females of both A. kurdistanella and A. gambronensis, like in the Stemmatophora species, vein R 5 is stalked with R 3 + R 4 (Figs 4 A, B). The length of the common stalk of R 5 with R 3 + R 4 in A. gambronensis is slightly shorter than in A. kurdistanella and in the latter species it is slightly shorter than in S. brunnealis. Moreover, based on Amsel (1959), in the hindwing of Acteniopsis the veins M 2 and M 3 originate from a common point, while in Stemmatophora these are on a short stalk. As we have shown, in the hindwing of A. gambronensis the veins M 2 and M 3 originate from one point, in S. brunnealis they are on a very short stalk. (Fig. 4 B), and only in A. kurdistanella they are clearly stalked (Fig. 4 A). Therefore, these characters cannot be considered as reliable diagnostic to distinguish Acteniopsis from Stemmatophora. Furthermore, based on the present study, the structure of the ventral ciliae of the male antennae in these two genera is completely different. In Stemmatophora each flagellum has two pairs of plumose ciliae, one pair at the proximal end of the flagellum which is larger and longer, and another pair beyond it, at the medial part of the segment and slightly positioned interiorly compared with the first pair which is smaller and shorter (Figs 4 D, E). In Acteniopsis, each flagellum has a transverse row of at least ten simple ciliae in which the lateral ciliae are the longest and slightly curved inward, whilst the medial ones are straight and about half as long as the lateral ciliae (Fig. 4 C). Based on the present study, the genus Acteniopsis is represented by two species worldwide that can easily be distinguished from each other based on their forewing pattern and shape of the phallus (see diagnosis of A. gambronensis). One of these two species has been described from Iraq and later form Iran, and another one is described from Iran. It seems that the genus is restricted to Saharo-Arabian and Irano-Turanian zoogeographical regions and it is most probable that these areas could be considered as its origin centre. Owing to the insufficient sampling in different parts of Iran, the presence of A. kurdistanella in some other parts of the country could be possible; this is also applicable for A. gambronensis along the southern borders of Iran.	en	Alipanah, Helen, Asselbergs, Jan (2018): Acteniopsis gambronensis sp. n. (Pyralidae: Pyralinae), a new species from Iran, with a revision of the genus Acteniopsis Amsel, 1959. Nota Lepidopterologica 41 (1): 87-97, DOI: http://dx.doi.org/10.3897/nl.41.22893, URL: http://dx.doi.org/10.3897/nl.41.22893
