identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EA6078453AFC10FBF97204FCAAFEE3.text	03EA6078453AFC10FBF97204FCAAFEE3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alloperla idei (Ricker 1935)	<div><p>Alloperla idei (Ricker)</p> <p>New Record: HUNTINGDON (S): at light, 2.5 km south of Jackson Corner, 25 June 2009, 1♂ (SJ). This collection, presumably from the nearby Standing Stone Creek in the central Pennsylvania Ridge and Valley Physiographic Province, fills in a gap between its known range of Georgia and Alabama north to Maine, Ontario, and Quebec (Surdick 2004). Baumann and Kondratieff (2009) noted that A. idei was more common in the northern part of its range and that individuals were often attracted to lights.</p> </div>	http://treatment.plazi.org/id/03EA6078453AFC10FBF97204FCAAFEE3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453AFC17FEB471D4FC2FF9E3.text	03EA6078453AFC17FEB471D4FC2FF9E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zealeuctra fraxina Ricker and Ross	<div><p>Zealeuctra fraxina Ricker and Ross</p> <p>New record: MONTGOMERY (D): on the PA Rte. 29/73 bridge over Mine Run, Schwenksville, PA, near the confluence of Perkiomen Creek, 4 March 2006, 1♂ (CL). The reported range of Z. fraxina is from West Virginia west to Illinois and south to Tennessee. Zealeuctra fraxina was collected in southeastern Pennsylvania and is the farthest eastern known record for this Nearctic genus. The closest reported occurrence is in the Potomac River basin in the eastern panhandle of West Virginia, the only West Virginia record (Tarter and Nelson 2006), over 250 km away. The genus Zealeuctra is most common in the midwestern United States south to Alabama and Texas, where several additional species are present and where the nymphs are most often associated with low order intermittent streams (Ricker and Ross 1969, Kondratieff and Zuellig 2004, Grubbs 2005). In the Ozarks, Zealeuctra are restricted to limestonebased intermittent streams (Poulton and Stewart 1991).</p> </div>	http://treatment.plazi.org/id/03EA6078453AFC17FEB471D4FC2FF9E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453DFC10FED97787FD41F9C7.text	03EA6078453DFC10FED97787FD41F9C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acroneuria kirchneri Stark and Kondratieff	<div><p>Acroneuria kirchneri Stark and Kondratieff</p> <p>New Record: CENTRE: (S) at light, approximately 2.5 km northeast of Martha Furnace, 2 July 1997, 1♀ (MH). The likely stream of origin is Bald Eagle Creek, about 2 km southeast of the collection site. Acroneuria adults are infrequently collected in PA except for the ubiquitous Acroneuria abnormis (Newman). This adult A. kirchneri was collected along with A. abnormis adults at a light trap set for terrestrial insects. This is a surprising addition to the Pennsylvania stonefly fauna; described in 2004, A. kirchneri was previously reported only from Kentucky, Virginia, and West Virginia. This new record is near the center of Pennsylvania, a distance of more than 250 km from the nearest recorded site, Preston County, in northeastern West Virginia.</p> </div>	http://treatment.plazi.org/id/03EA6078453DFC10FED97787FD41F9C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453DFC10FEB972FBFB3BFD35.text	03EA6078453DFC10FEB972FBFB3BFD35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allocapnia wrayi Ross 1964	<div><p>Allocapnia wrayi Ross</p> <p>Allocapnia wrayi was deleted from the Pennsylvania stonefly species list because previous records had been misidentified (Earle 2004a). Examination of new material, however, revealed several authentic records. This species reaches its northern limit in 1 st to 3 rd order creeks in the Potomac and lower Susquehanna River basins of Pennsylvania.</p> <p>New records: BEDFORD: (P) Blackberry Lick Run, PA Rte. 26, 20 January 2006, 10♂; Blues Gap Run, 0.8 km upstream of mouth, 20 January 2006, 2♂ 1♀; spring run to unnamed tributary to Blues Gap Run, 0.8 km north of Elbinsville, 20 January 2006, 1♂; Fifteenmile Creek, 1 st bridge upstream of the Maryland state line, 20 January 2006, 1m; CUMBERLAND: (S) Doubling Gap Creek, 10 m downstream of the lake at Colonel Denning State Park, 7 December 1997, 7♂; FRANKLIN: (P) Broad Run, 30 m downstream of water supply reservoir, 24 Mar. 2006, 4♀; FULTON: (P) Sideling Hill Creek, PA Rte. 484, 20 January 2006, 1♂.</p> </div>	http://treatment.plazi.org/id/03EA6078453DFC10FEB972FBFB3BFD35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453DFC10FCBE77EDFABBF9C7.text	03EA6078453DFC10FCBE77EDFABBF9C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capniidae	<div><p>Capniidae</p> <p>Capniidae is the largest family of Plecoptera in North America, consisting of 17 genera and approximately 250 mostly winter emerging species (Nations et al. 2007); however, only 2 genera, Allocapnia and Paracapnia, are found in the Mid- Atlantic/Appalachian region. The Pennsylvania fauna consists of 15 species of Allocapnia and one species of Paracapnia. Adults can be collected during the coldest months of the year along streams and on bridges, tree trunks, and buildings. Two to five species are normally collected at any one location in Pennsylvania. The most common species in Pennsylvania are Allocapnia granulata (Claassen), A. nivicola (Fitch), A. pygmaea (Burmeister), A. rickeri Frison, A. recta (Claassen), and Paracapnia angulata Hanson.</p> <p>The following species are infrequently collected or have a limited range in Pennsylvania.</p></div> 	http://treatment.plazi.org/id/03EA6078453DFC10FCBE77EDFABBF9C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453DFC10FE837507FCBBFC63.text	03EA6078453DFC10FE837507FCBBFC63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sweltsa palearata Surdick 2004	<div><p>Sweltsa palearata Surdick</p> <p>New records: BEDFORD: (P) Fifteenmile Creek, Old Big Mountain Road, 14 May 1999, 7♂, 1♀; spring run to an unnamed tributary of Blues Gap Run, 0.8 km north of Elbinsville, 14 May 1999, 1♂. Both sites are 1 st order forested stream reaches. The spring run site is not delineated in blue on the 7.5 minute USGS topographic map and is likely dry in the summertime. One of several Sweltsa species whose ranges are limited to specific river basins, S. palearata reaches its northern limit in the main stem Potomac River basin of southcentral Pennsylvania. In West Virginia S. palearata is limited to the South Branch Potomac River tributaries, in Maryland to the main stem Potomac River tributaries, and in Virginia to the upper Potomac River basin and the North and South Forks of the Shenandoah River basin (Surdick 2004).</p> </div>	http://treatment.plazi.org/id/03EA6078453DFC10FE837507FCBBFC63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453DFC11FC94724EFDD2FC63.text	03EA6078453DFC11FC94724EFDD2FC63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allocapnia aurora Ricker 1952	<div><p>Allocapnia aurora Ricker</p> <p>is limited to small, mostly forested or rural coldwater creeks in the lower Susquehanna River basin and the extreme eastern portion of the Potomac River basin in southcentral Pennsylvania. Allocapnia aurora was previously confirmed from only 10 Pennsylvania creeks (Masteller 1996). Although not as rare as certain other Pennsylvania Allocapnia, it is included here to correct earlier misidentifications. Males can be difficult to separate from A. pygmaea and A. nivicola. Male Allocapnia from Indian Creek in Westmoreland County and French Creek in Chester County that were identified as A. aurora (Earle 1994 and 2003) were actually A. nivicola. The A. aurora records from Muddy Creek in York County (Masteller 1996) were also misidentified and were actually A. pygmaea.</p> <p>New records: CUMBERLAND: (S) Doubling Gap Creek, Elk Hill Rd. at PA Rte. 233, 31 January 2001, 7♂, 11♀; same location, 10 March 1997, 6♂, 4♀; Mountain Creek, Mount Holly Springs, 8 January 1997, 1♂; DAUPHIN: (S) Rattling Creek, in mountain gap at Lykens Borough Park, 28 December 2005, 3♂, 1♀; LANCASTER: (S) Trout Run, Pennsy Rd., 1 February 1997, 1♂; JUNIATA: (S) Delaware Creek, PA Rte. 333 near Thompsontown, 4 February 1994, 1♂; PERRY: (S) Fishing Creek, New Valley Rd. west of Marysville, 2 February 1999, 1♂; Laurel Run, tributary to Tuscarora Creek, PA Rte. 850, 3 January 2001, 6♂; same location, 3 March 1995, 2♀; McCabe Run, Barkley Rd., 11 April 1993, 3♀; Shaeffer Run, 2 nd Narrows Rd., 24 March 1994, 1♀ (LJ).</p> </div>	http://treatment.plazi.org/id/03EA6078453DFC11FC94724EFDD2FC63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453CFC11FF1E77A3FE00FA99.text	03EA6078453CFC11FF1E77A3FE00FA99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allocapnia frisoni Ross and Ricker	<div><p>Allocapnia frisoni Ross and Ricker</p> <p>is distributed west of the Allegheny Front from New York south to Tennessee and west to Wisconsin. In Pennsylvania it is most commonly collected in the Youghiogheny River drainage and in Lake Erie tributaries (Masteller 1996) where it often replaces the similar A. granulata.</p> <p>New records: FAYETTE: (O) Back Creek, 800 m upstream of mouth, 27 December 1996, 4♂; same location, 30 January 1997, 4♂, 1♀, same location, 1 March 1995, 1♂; Little Champion Creek, SR 1050, 20 December 1997, 1♂.</p> </div>	http://treatment.plazi.org/id/03EA6078453CFC11FF1E77A3FE00FA99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453CFC11FF1E7138FC7BFEBF.text	03EA6078453CFC11FF1E7138FC7BFEBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allocapnia vivipara (Claassen 1924)	<div><p>Allocapnia vivipara (Claassen)</p> <p>has been most commonly collected in the Lake Erie tributaries in northwestern PA, but has also occasionally been found in the lower Allegheny River and lower Delaware River basins (Masteller 1996). Allocapnia vivipara is the most common Allocapnia species in Illinois, where it lives in a variety of streams, from small pristine creeks to large clay bottomed rivers and streams with livestock waste (Webb 2002). The Pennsylvania records also indicate a tolerance to a variety of pollution sources including urban and agricultural runoff, alkaline coal mine drainage, and leaky septic systems.</p> <p>New records: ARMSTRONG: (O) Long Run, PA Rte. 506, 9 February 2002, 2♂ (BS); Pine Run, SR 4071, 23 January 2002, 14♂ (BS); same location, 25 February 2002, 9♂, 2♀ (BS); Roaring Run, SR 2051, 9 February 2002, 1♀ (BS). WESTMORELAND: (O) Monastery Run, 15 m downstream of US Rte. 30, 30 January 1997, 3♂, 1♀.</p></div> 	http://treatment.plazi.org/id/03EA6078453CFC11FF1E7138FC7BFEBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453CFC11FC947507FB1FFBD9.text	03EA6078453CFC11FC947507FB1FFBD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allocapnia zola Ricker 1952	<div><p>Allocapnia zola Ricker</p> <p>ranges from New Brunswick south to Virginia and Kentucky. The only previous Pennsylvania records are from 3 small forested creeks in Carbon, Perry, and Westmoreland Counties (Masteller 1996). Allocapnia zola is found mainly in small clean cold water creeks; however, in Pennsylvania it also has been found in medium creeks and in streams receiving alkaline coal mine drainage. Most records are from Blue Mountain in southcentral Pennsylvania, the southern-most ridge in the Ridge and Valley Physiographic Province.</p> <p>New records: BEDFORD: (P) Fifteenmile Creek, 1 st bridge upstream of Maryland state line, 20 January 2006, 2♂; Piney Creek, PA Rte. 26, 6 February 2005, 2♂; FAYETTE: (O) Little Champion Creek, SR 1050, 24 February 1997, 1♂; Champion Creek, 50 m downstream of Little Champion Creek, 24 February 1997, 1♂, 1♀, FULTON: (P) Trough Run, PA Rte. 484, 24 March 2006, 1♂.</p></div> 	http://treatment.plazi.org/id/03EA6078453CFC11FC947507FB1FFBD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453CFC11FCBE705CFBD8FA51.text	03EA6078453CFC11FCBE705CFBD8FA51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leuctridae	<div><p>Leuctridae</p> <p>Leuctridae nymphs are important components of the benthos of small to medium creeks throughout Pennsylvania; 13 species in 4 genera are present. The most common Pennsylvania species is Leuctra ferruginea (Walker). Adult L. duplicata Claassen, L. sibleyi Claassen, L. tenuis (Pictet), and Paraleuctra sara (Claassen) are also commonly collected in Pennsylvania. Most of the Pennsylvania species are acid tolerant (Earle 2004b).</p> </div>	http://treatment.plazi.org/id/03EA6078453CFC11FCBE705CFBD8FA51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453CFC12FC9471F0FEDFFDC8.text	03EA6078453CFC12FC9471F0FEDFFDC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leuctra alexanderi Hanson 1941	<div><p>Leuctra alexanderi Hanson</p> <p>has been previously recorded from only 5 small cold forested creeks or spring runs in Pennsylvania (Earle 2004b, Masteller 1996). The Pennsylvania distribution of L. alexanderi is the Potomac River basin, west and north of the Allegheny Front through the Ohio and upper West Branch Susquehanna River basins, and east to the upper Delaware River basin in northeastern Pennsylvania. Leuctra alexanderi is not as widely distributed in North America as other Leuctra species, having been confirmed from only 5 states from Pennsylvania south to Tennessee. Leuctra alexanderi is common in small spring-fed creeks in western Maryland (Grubbs 1997).</p> <p>New Records: BEDFORD: (P) Fifteenmile Creek, Old Big Mountain Rd., 24 May 2002, 1♂, 1♀; spring run to an unnamed tributary to Blues Gap Run, 800 m north of Elbinsville, 6 June 2001, 2♂; FAYETTE: (O) Back Creek, 800 m upstream of mouth, 4 June 1996, 2♂; WESTMORELAND: (O) Grove Run, 75 m upstream of mouth, 13 June 2009, 2♂; spring run to Indian Creek, PA Rte. 381, 1 km north of PA Rte. 31, 4 June 1996, 1♂.</p> </div>	http://treatment.plazi.org/id/03EA6078453CFC12FC9471F0FEDFFDC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453FFC12FF1E7147FEF1F8CF.text	03EA6078453FFC12FF1E7147FEF1F8CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphinemura appalachia Baumann 1996	<div><p>Amphinemura appalachia Baumann</p> <p>is a relatively recent addition to the PA stonefly fauna, previously collected from only two small forested creeks in northeastern Pennsylvania, in the upper Delaware and North Branch Susquehanna River basins, one of which is the paratype location (Baumann 1996, Masteller 1996). Amphinemura appalachia as the name suggests is found in the Appalachians from Pennsylvania south to Georgia.</p> <p>New records: BERKS: (D) Northkill Creek, most downstream bridge in state game lands, 22 April 1994, 1♂; SCHUYLKILL: (S) Catawissa Creek, 10 m upstream of Audenreid Tunnel discharge, 20 April 2005, 5♂, 2♀.</p> </div>	http://treatment.plazi.org/id/03EA6078453FFC12FF1E7147FEF1F8CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453FFC13FC94726AFE06FEE3.text	03EA6078453FFC13FC94726AFE06FEE3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolotoperla rossi (Frison 1942)	<div><p>Bolotoperla rossi (Frison)</p> <p>, the rarest Pennsylvania species in the family, was previously known from only few adults collected from 3 medium sized creeks in the Susquehanna River basin (Masteller 1996). This species has apparently not been documented in the nymph stage in Pennsylvania. Emergence begins in early April and continues through mid-May, later than most other members of the family. Bolotoperla rossi ranges from Maine south to North Carolina.</p> <p>New records: DAUPHIN: (S) Manada Creek, Furnace Road, 1 April 2000, 1♂ (CL); PIKE: (D) Dingmans Creek between falls and parking lot, 21 May 2000, 1♀ (CL).</p></div> 	http://treatment.plazi.org/id/03EA6078453FFC13FC94726AFE06FEE3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453FFC12FF1E7648FEC0FC3F.text	03EA6078453FFC12FF1E7648FEC0FC3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leuctra variabilis	<div><p>Leuctra variabilis</p> <p>is one of the few stoneflies reported from unusual habitats for stoneflies, bogs with imperceptible flow and pitcher plants (Nelson et al. 2002). Leuctra variabilis emerges rather late compared to most other Leuctra species, from late September to early December. Only 2 previous Pennsylvania records exist, from Susquehanna and Wyoming Counties in northeastern Pennsylvania (Masteller 1996). Leuctra variabilis has been reported from New England south to Maryland.</p> <p>New record: CARBON: (D) unnamed tributary to Lizard Creek, 20 m upstream of T-334, 16 October 2003, 1♂ (TD).</p></div> 	http://treatment.plazi.org/id/03EA6078453FFC12FF1E7648FEC0FC3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453FFC12FF1E7349FB78FE11.text	03EA6078453FFC12FF1E7349FB78FE11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nemoura trispinosa Claassen 1923	<div><p>Nemoura trispinosa Claassen</p> <p>was a recent addition to the Pennsylvania stonefly fauna based on one adult and numerous nymphs from one creek in Monroe County (Earle 2004a). In Pennsylvania, N. trispinosa appears to be limited to small, low gradient swampy creeks. Nemoura trispinosa is found from the Canadian Maritimes south to Pennsylvania and west to Manitoba and Wyoming.</p> <p>New record: CLINTON: (S) Swamp Branch of Big Run, Haynes Township, 17 May 2004, late instar nymph (CM).</p> </div>	http://treatment.plazi.org/id/03EA6078453FFC12FF1E7349FB78FE11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453FFC12FF407078FED4FAFE.text	03EA6078453FFC12FF407078FED4FAFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nemouridae	<div><p>Nemouridae</p> <p>Nemouridae are common stoneflies found mostly in small to medium creeks, seeps, springs, and spring runs. Seventeen species in 6 genera have been recorded in Pennsylvania. The most commonly collected Pennsylvania species are Amphinemura nigritta (Provancher), A. wui (Claassen), and Prostoia similis (Hagen).</p> </div>	http://treatment.plazi.org/id/03EA6078453FFC12FF407078FED4FAFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453FFC12FC9475B3FB74FD36.text	03EA6078453FFC12FC9475B3FB74FD36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ostrocera Ricker 1952	<div><p>Ostrocera nymphs</p> <p>are rarely collected during routine water quality monitoring; however, they can be common in small acidic, often intermittent creeks and spring runs (Earle 2004b). Adults can also be collected alongside small to medium cold water creeks, presumably emerging from nearby springs or seepage areas. The most common Pennsylvania species is O. albidipennis (Walker).</p> </div>	http://treatment.plazi.org/id/03EA6078453FFC12FC9475B3FB74FD36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453FFC12FC94769FFB8EFB07.text	03EA6078453FFC12FC94769FFB8EFB07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ostrocerca complexa (Claassen 1937)	<div><p>Ostrocerca complexa (Claassen)</p> <p>, which ranges from the Canadian Maritimes west to Ontario and south through the Appalachians to Virginia, was previously reported from only one small creek in Pennsylvania, Spring Brook in Lackawanna County, in northeastern Pennsylvania in 1945 (Masteller 1996). Three additional sites are now added in northcentral and northeastern Pennsylvania.</p> <p>New records: MONROE: (D) Two Mile Run, 20 m upstream of PA Rte. 115, 13 May 2008, 1♀; POTTER: (S) spring run to MacDonald Hollow, near mouth, 17 May 1976, 1♂; WAYNE: (D) third unnamed tributary downstream of Cobb Pond to Wagnum Creek, 23 May 2003, 1♂ (TD).</p> </div>	http://treatment.plazi.org/id/03EA6078453FFC12FC94769FFB8EFB07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453FFC12FCBE7163FB34FA2B.text	03EA6078453FFC12FCBE7163FB34FA2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniopterygidae Klapalek 1905	<div><p>Taeniopterygidae</p> <p>The Pennsylvania Taeniopterygidae fauna is represented by 11 species in 5 genera. The most commonly collected species are Strophopteryx fasciata (Burmeister), Taeniopteryx burksi Ricker and Ross, T. maura (Pictet), and T. nivalis (Fitch). Adults emerge in winter through early spring.</p> </div>	http://treatment.plazi.org/id/03EA6078453FFC12FCBE7163FB34FA2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453EFC13FC9475B0FC69FC3E.text	03EA6078453EFC13FC9475B0FC69FC3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alloperla aracoma Harper and Kirchner	<div><p>Alloperla aracoma Harper and Kirchner</p> <p>was previously reported from only one Pennsylvania stream, Powdermill Run, Westmoreland County, in the Conemaugh River portion of the Ohio River basin (Grubbs 1996). Maryland and West Virginia are the only other states with records of A. aracoma, where it is restricted to the Allegheny Plateau and the Ridge and Valley Physiographic Provinces (Surdick 2004).</p> <p>New record: SOMERSET: (S) Breastwork Run, first SR 1017 bridge south of US Rte. 30, 29 June 1994, 1♂. Breastwork Run is a small forested cold water creek in the upper Juniata River portion of the Susquehanna River basin, along the eastern flank of the Allegheny Front near the divide with the Ohio River basin.</p> </div>	http://treatment.plazi.org/id/03EA6078453EFC13FC9475B0FC69FC3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453EFC13FC947787FAD2F959.text	03EA6078453EFC13FC947787FAD2F959.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alloperla petasata Surdick 2004	<div><p>Alloperla petasata Surdick</p> <p>replaces Alloperla caudata Frison in its northern and eastern range in North America including Pennsylvania (Surdick 2004). Alloperla petasata ranges from the Canadian Maritimes south to Georgia; its sister species, A. caudata, is now restricted to Illinois, the Ozark- Ouachita Highlands, and the Lower Tennessee River in Alabama (Surdick 2004) and Mississippi (Willett and Stark 2009).</p> <p>New records: BEDFORD: (P) Blackberry Lick Run, PA Rte. 26, 9 June 2001, 1♂, 4♀; Blues Gap Run, 400 m east of Rte. T-323, 9 June 2001, 2♂, 3♀; INDIANA: (O) Little Yellow Creek, PA Rte. 259, 21 June 2000, 8♂, 3♀ (TWS), as Alloperla sp. in Yasik et al. (2002); PIKE: (D) Carley Brook, 1.5 km upstream of village of Carley Brook, 8 July 2003, 1♂ (TD). These sites are small forested creeks except for Little Yellow Creek, which is a 3rd order creek whose watershed has surface coal mines with alkaline mine drainage.</p> </div>	http://treatment.plazi.org/id/03EA6078453EFC13FC947787FAD2F959	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453EFC13FF407223FBC8FE12.text	03EA6078453EFC13FF407223FBC8FE12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chloroperlidae	<div><p>Chloroperlidae</p> <p>Most Chloroperlidae, except for Alloperla atlantica Baumann, Sweltsa lateralis (Banks), S. onkos, S. hoffmani, and Haploperla brevis (Ricker), have been little collected in the adult stage in Pennsylvania. Adults are bright yellow or pale green and can most often be seen soon after emergence flying over small to medium creeks or on instream rocks. Nymphs other than Sweltsa are uncommonly collected in Pennsylvania except just before emergence due to their largely hyporheic habitat. The Pennsylvania fauna contains 19 species in 6 genera; 10 species are in the genus Alloperla. Many North American species of Chloroperlidae are regional endemics that are restricted to specific river basins or physiographic provinces (Surdick 2004).</p> </div>	http://treatment.plazi.org/id/03EA6078453EFC13FF407223FBC8FE12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453EFC13FF1E7525FCB8FCAB.text	03EA6078453EFC13FF1E7525FCB8FCAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniopteryx metequi Ricker and Ross	<div><p>Taeniopteryx metequi Ricker and Ross</p> <p>is one of the rarest Pennsylvania members of the genus, having been reported from only four counties (Masteller 1996). Although a 1966 collection from one site in Snyder County in the Susquehanna River basin was included in Ricker and Ross (1968), all recent collections have been restricted to the Potomac and Monongahela River basins in PA. Interestingly, T. metequi has been reported in all river basins except the Potomac in West Virginia (Tarter and Nelson 2006) and only in the eastern Ridge and Valley portion of the Potomac River basin in Maryland (Grubbs 1997).</p> <p>New Records: FULTON: (P) Sideling Hill Creek, PA Route 484, 6 February 2005, 2♂; Tonoloway Creek, SR 2004 at Johnsons Mill, 6 February 2005, 2♂.</p> </div>	http://treatment.plazi.org/id/03EA6078453EFC13FF1E7525FCB8FCAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453EFC13FF1E77EDFE7DF9C6.text	03EA6078453EFC13FF1E77EDFE7DF9C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniopteryx ugola Ricker and Ross	<div><p>Taeniopteryx ugola Ricker and Ross,</p> <p>which ranges through the Appalachians from Pennsylvania south to Georgia, is generally restricted to the headwaters of cold small and medium sized creeks in Pennsylvania. Only 3 previous records exist, from Cambria and Cumberland Counties (Masteller 1996). Taeniopteryx ugola, which appears to be the only Taeniopteryx species collected in Pennsylvania creeks with pH less than 5.5 and with summer-fall stream dry periods, has been most frequently collected from upper Mountain Creek in Cumberland County, Pennsylvania.</p> <p>New records: LEBANON: (S) Indiantown Run, PA Rte. 443 at gap in mountain, 20 February 2004, 2♂; PERRY: (S) Shaeffer Run, 2 nd Narrows Road, 24 March 1995, 1♀ (LJ); SCHUYLKILL: (S) Catawissa Creek, Girard Manor Road, 13 January 2004, 3♂; unnamed tributary to Tomhicken Creek, PA Rte. 934, 19 March 2003, 1♂.</p> </div>	http://treatment.plazi.org/id/03EA6078453EFC13FF1E77EDFE7DF9C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA6078453EFC1CFC9472FBFDE6FE12.text	03EA6078453EFC1CFC9472FBFDE6FE12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rasvena terna (Frison 1942)	<div><p>Rasvena terna (Frison)</p> <p>has only two previous confirmed adult records in Pennsylvania, a 1994 record from Grays Run in Lycoming County and a 1958 record of unknown location. Surdick (2004) reported that R. terna is restricted to the Appalachian Highlands of eastern North America, from Maine south to North Carolina and Tennessee.</p> <p>New records: CLINTON: (S) North Branch Tangascootack Creek, near mouth, 13 May 1998, 1♀; FORREST: (O) Minister Creek, 100 m upstream of SR 2001, 14 April 2008, 5 late instar nymphs, (RR). These are the first confirmed collection of Rasvena nymphs in Pennsylvania. These records suggest a preference for small, clear coldwater creeks.</p> </div>	http://treatment.plazi.org/id/03EA6078453EFC1CFC9472FBFDE6FE12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784531FC1CFF1E71D5FEAEF8EB.text	03EA60784531FC1CFF1E71D5FEAEF8EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acroneuria Although	<div><p>Acroneuria:</p> <p>Although 11 species of Acroneuria have been reported from Pennsylvania, only 9 appear valid. Many records are from collections prior to the 1950’s before major species revisions, or are based on difficult to identify nymphs and need to be confirmed. The species most commonly collected as nymphs are A. abnormis and A. carolinensis (Banks), which are found in small to medium sized, usually forested creeks. Acroneuria abnormis, the most commonly collected adult Acroneuria, is also found in rivers.</p> </div>	http://treatment.plazi.org/id/03EA60784531FC1CFF1E71D5FEAEF8EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784531FC1CFF1E732AFB61FE2D.text	03EA60784531FC1CFF1E732AFB61FE2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acroneuria arenosa (Pictet)	<div><p>Acroneuria arenosa (Pictet)</p> <p>was originally described from Pennsylvania in 1841. The only valid Pennsylvania records were collected prior to 1950 and confirmed by Stark and Gaufin (1976a). Acroneuria arenosa ranges from Quebec south to Florida.</p> <p>New records: BERKS: (D) Schuylkill River near Douglassville, 17 May 1987, 1♀ (WE); JUNIATA: (S) Juniata River Thompsontown, 28 July 1989, 1♀ (LJ). Both sites are warm water rivers.</p> </div>	http://treatment.plazi.org/id/03EA60784531FC1CFF1E732AFB61FE2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784531FC1CFC057594FB18FC62.text	03EA60784531FC1CFC057594FB18FC62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acroneuria perplexa Frison 1937	<div><p>A female Acroneuria perplexa Frison</p> <p>collected at a light in Philadelphia (year unknown) and first reported in Frison (1937) has been of questionable status in Pennsylvania since this is the only Pennsylvania record and A. perplexa is a largely Southern and Midwestern U.S. species. The specimen was examined by Stark during his review of Acroneuria and determined to be A. arenosa; however, the corrected identification was not included in Stark and Gaufin (1976a) and the record was carried forward in some later publications including Stewart and Stark (2002) and Stark et al. (2009). This record should be changed to A. arenosa.</p> </div>	http://treatment.plazi.org/id/03EA60784531FC1CFC057594FB18FC62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784531FC1DFC9477A2FE9BFC62.text	03EA60784531FC1DFC9477A2FE9BFC62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Attaneuria ruralis (Hagen 1861)	<div><p>Attaneuria ruralis (Hagen)</p> <p>records in Pennsylvania are based on collections from 14 locations in 9 counties (Masteller 1996); however, most of these records are of adults collected prior to 1948 or nymphs collected and identified as part of stream bioassessments. Questions on the identification of A. ruralis nymphs by Pennsylvania aquatic biologists prompted me to examine nymph specimens of record at the Academy of Natural Sciences of Philadelphia and the Frost Entomology Museum of Pennsylvania State University, and additional material from the Pennsylvania Department of Environmental Protection, Harrisburg, PA, and United States Environmental Protection Agency, Wheeling, WV. Nymphs were also compared with A. ruralis nymphs from Iowa. All nymphs identified as A. ruralis in these collections and recorded in Masteller (1996) were determined to be Paragnetina media (Walker), a common stonefly found in small to large warm water creeks in Pennsylvania, except for one nymph from Pine Creek, Blackwell, Tioga County, which was an Acroneuria. Additionally, one adult from Fayette County in the Frost Museum collections identified as A. ruralis was an Acroneuria. The only valid Pennsylvania records for A. ruralis, therefore, appear to be the pre-1946 adult specimens from Cumberland, Dauphin, and Lebanon Counties noted in Surdick and Kim (1976), which were confirmed by Stark in his review of the Perlidae genera (Stark and Gaufin 1976b) or cited in Needham and Claassen (1925). All but one of these records is from the Harrisburg area, likely from the Susquehanna River; the lone additional specimen is from an unspecified location in Lebanon County. Casual collections of adult stoneflies in the Harrisburg area during the past 20 years have not yielded any additional specimens of A. ruralis. Also, I have not been able to confirm the collection of any A. ruralis nymphs from Pennsylvania. Targeted collection efforts should be conducted in proper habitats to determine if this species is still present in Pennsylvania. Heimdal et al. (2004) indicated that A. ruralis nymphs could be found on woody debris or boulders in the slower current of rivers in interior Iowa. Poulton and Stewart (1991) stated that A. ruralis nymphs could be collected in Ozark streams by pulling debris from the river bottom or disturbing rocks or riprap upstream from a net.</p> </div>	http://treatment.plazi.org/id/03EA60784531FC1DFC9477A2FE9BFC62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784531FC1CFF40708EFDDCFA6C.text	03EA60784531FC1CFF40708EFDDCFA6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Perlidae	<div><p>Perlidae</p> <p>The Perlidae in Pennsylvania are represented by 30 species in 9 genera; however, many species are restricted to medium to large warm water creeks and rivers and are rarely collected. Adults can be most frequently collected at outdoor lights.</p> </div>	http://treatment.plazi.org/id/03EA60784531FC1CFF40708EFDDCFA6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784531FC1CFF1E75B0FEC4FB64.text	03EA60784531FC1CFF1E75B0FEC4FB64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Utaperla gaspesiana Harper and Roy	<div><p>Utaperla gaspesiana Harper and Roy</p> <p>is extremely rare in Pennsylvania. Adults were collected in the 1970’s by R. Surdick from Maul Spring and Powdermill Run, Westmoreland County, in southwestern Pennsylvania; however, four years of sampling in that watershed by Grubbs (1996) did not yield any additional specimens. Only one other previous Pennsylvania record exists, from Tancascootack Creek in Clinton County (Masteller 1996). Despite its rarity, benthic taxa lists for Pennsylvania streams assessments occasionally include Utaperla nymphs, which are most likely misidentifications of Alloperla usa Ricker, similar to the misidentifications discussed by Kondratieff et al. (1995) for North and South Carolina assessments. Utaperla gaspesiana ranges from Quebec and Maine south to West Virginia.</p> <p>New record: MONROE: (D) Pocono Creek, Camelback Road, 22 May 2002, 1♀ (CL). Pocono Creek is a small coldwater creek in northeastern Pennsylvania.</p> </div>	http://treatment.plazi.org/id/03EA60784531FC1CFF1E75B0FEC4FB64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784530FC1DFC947240FA02F95B.text	03EA60784530FC1DFC947240FA02F95B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diploperla duplicata (Banks 1920)	<div><p>Diploperla duplicata</p> <p>New record: CUMBERLAND: (S) Doubling Gap Creek, PA Rte. 233 and Elk Hill Rd., 22 June 1995, 1♀.</p> </div>	http://treatment.plazi.org/id/03EA60784530FC1DFC947240FA02F95B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784530FC1DFC9472FAFCD9F8E5.text	03EA60784530FC1DFC9472FAFCD9F8E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diploperla robusta	<div><p>Diploperla robusta</p> <p>New records: CAMBRIA: (O) Roaring Run, PA Rte. 160, 18 June 1998, 1♀; WESTMORELAND: (O) Powdermill Run, Bioblitz Site 1, 19 June 2003, 1♂ (JW).</p></div> 	http://treatment.plazi.org/id/03EA60784530FC1DFC9472FAFCD9F8E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784530FC1DFF1E77A3FD43F9C7.text	03EA60784530FC1DFF1E77A3FD43F9C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hansonoperla appalachia Nelson	<div><p>Hansonoperla appalachia Nelson</p> <p>is known from only a few small to medium forested cold water creeks in Pennsylvania and in the Appalachians from New Hampshire south to Virginia. Nymphs are rarely collected, and when present, usually only one or two individuals are found at any one site. Only one adult record from Pennsylvania exists (Masteller 1996). The most nymphs collected in Pennsylvania were from Babb Creek, Tioga County, which has been affected by abandoned coal mine discharges and has a pH of 5.0 and slightly elevated dissolved aluminum (Pennsylvania Department of Environmental Protection, unpublished data).</p> <p>New records: SUSQUEHANNA: (S) Nick Creek, Auburn Township, 23 June 2000, one nymph (TS); unnamed tributary to Choconut Creek, near outlet of Lake Sophia, 21 June 2000, one nymph (SRBC).</p></div> 	http://treatment.plazi.org/id/03EA60784530FC1DFF1E77A3FD43F9C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784530FC1DFF1E724FFACCFE2D.text	03EA60784530FC1DFF1E724FFACCFE2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Perlinella drymo (Newman 1839)	<div><p>Perlinella drymo (Newman)</p> <p>is the less commonly collected of the two Pennsylvania species of Perlinella. Both adults and nymphs are extremely elusive. Only two previous Pennsylvania adult records exist, one from Northampton County in 1931 and one from Cumberland County in 1949 (Masteller 1996). Perlinella drymo is a widespread North American species found from the Canadian Maritimes south to Florida and west to Minnesota and Texas.</p> <p>New records: LUZERNE: (S) Nescopeck Creek, old foot bridge at lower end of Nescopeck State Park, 20 April 2001, 1 nymph (MW); MONTGOMERY: (D) unnamed tributary to Schuylkill River near Spring City, 9 February 2005, 1 nymph (MB). Based on these records, P. drymo appears to prefer smaller cooler creeks than P. ephyre (Newman), which inhabits large warm water creeks and rivers in Pennsylvania.</p> </div>	http://treatment.plazi.org/id/03EA60784530FC1DFF1E724FFACCFE2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784530FC1DFCBE7649FB2BF9C1.text	03EA60784530FC1DFCBE7649FB2BF9C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Perlodidae	<div><p>Perlodidae</p> <p>The Pennsylvania Perlodidae fauna is represented by 27 species in 9 genera. The family is divided into 2 subfamilies, Isoperlinae which in Pennsylvania includes Isoperla and Clioperla, and Perlodinae, which includes the other genera. Isoperla has the most species of all the Pennsylvania stonefly genera, 17 described species and several undescribed species (Szczytko and Kondratieff unpublished). Perlodinae are most commonly collected in the forested mountainous regions of Pennsylvania. Although Perlodinae nymphs are often collected in the proper habitat, adults of most genera are rarely collected. Only new adult records are included for the following species.</p> <p>Two species of Diploperla have been reported from Pennsylvania: D. duplicata (Banks), which ranges from Pennsylvania south to Georgia, and D. robusta Stark and Gaufin, which was split from D. duplicata in 1974 and ranges from Connecticut south to Indiana and Kentucky. Recent collections suggest that D. duplicata is restricted to eastern Pennsylvania and D. robusta to western Pennsylvania; however, several records predating 1974 need to be examined and additional specimens collected. Both species of Diploperla are found in small to medium creeks. Diploperla nymphs are especially common in small creeks in the Allegheny National Forest in northwestern Pennsylvania.</p> </div>	http://treatment.plazi.org/id/03EA60784530FC1DFCBE7649FB2BF9C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784530FC1EFC94732CFE5CFD1A.text	03EA60784530FC1EFC94732CFE5CFD1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helopicus subvarians (Banks 1920)	<div><p>Helopicus subvarians (Banks)</p> <p>has been collected mostly from medium to large creeks and rivers in Pennsylvania. Previous adult records are from the Delaware River basin in Carbon County and the Susquehanna River basin in Huntingdon, Lycoming, and Union Counties. Helopicus subvarian s is found from Maine and Quebec south to Florida.</p> <p>New records: DAUPHIN: (S) Susquehanna River, on Market Street Bridge near downtown Harrisburg, 1 May 2000, 1♂, 1♀ (DB); Mifflin: (S) Penns Creek, near mouth of Swift Run, 20 May 1998, 1♂ (CL); same location, 22 May 2007, 1♀ (CL); WAYNE: (D) Delaware River, 1 km downstream of Stockport, 20 May 2007, 1♀ (TD); Delaware River, across from the New York State Forest Preserve at Bouchouxville Rd., 24 June 2007, 1♀ (TD).</p> </div>	http://treatment.plazi.org/id/03EA60784530FC1EFC94732CFE5CFD1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784533FC1EFF1E76BBFD6EFBAC.text	03EA60784533FC1EFF1E76BBFD6EFBAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isogenoides hansoni (Ricker 1952)	<div><p>Isogenoides hansoni (Ricker)</p> <p>nymphs have been most often collected in small to medium forested cold water creeks in northern and southwestern Pennsylvania. The type locality is Brodhead Creek, Monroe County, in northeastern Pennsylvania. Previous documentation of adults in Pennsylvania was limited to 6 creeks in Forest, McKean, Monroe, Potter, and Somerset Counties (Masteller 1996). Isogenides hansoni ranges from the Canadian Maritimes south to Virginia.</p> <p>New record: CLINTON: (S) Chatham Run, near village of Woolrich, 11 May 2003, 5♀ (GSm).</p></div> 	http://treatment.plazi.org/id/03EA60784533FC1EFF1E76BBFD6EFBAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
03EA60784533FC1EFF1E7014FDBAFA6C.text	03EA60784533FC1EFF1E7014FDBAFA6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yugus kirchneri Nelson	<div><p>Yugus kirchneri Nelson</p> <p>nymphs can be collected in low order forested cold water Pennsylvania creeks, mostly commonly in northern and southwestern Pennsylvania; previous adult records were limited to two creeks in Potter and Westmoreland Counties (Masteller 1996). Yugus kirchneri ranges in the Appalachians from Pennsylvania south to Virginia.</p> <p>New record: BEDFORD: (S) Pavia Run, near mouth, 10 June 2006, 1♀ (GS).</p> </div>	http://treatment.plazi.org/id/03EA60784533FC1EFF1E7014FDBAFA6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Earle, Jane I.	Earle, Jane I. (2009): New State Stonefly (Plecoptera) Records For Pennsylvania, With Additional Records And Information On Rare Species. Illiesia 5 (16): 169-181, DOI: http://doi.org/10.5281/zenodo.4759416
