identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4D4A87AD8E3B5851FC6CF943FB6AFE71.text	4D4A87AD8E3B5851FC6CF943FB6AFE71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protonemura talboti : Rauser 1963	<div><p>Protonemura talboti (Nav{s, 1929)</p> <p>(Figs. 3, 6, 9-10, 16-27, 75)</p> <p>Nemoura talboti Nav {s, 1929:230. (original description from Algeria); Claassen, 1940:64. (catalog).</p> <p>Nemoura (Protonemura) talboti Aubert, 1952:239. (redescription of the types); Aubert, 1956:422. (complementary description of the imago, description of the larva, first records from Morocco); Aubert, 1961 (partim):219. (new records from Morocco).</p> <p>Protonemura talboti: Raušer, 1963:804. (comparative drawings, distribution map); Aubert, 1964a:72. (comparison); Aubert, 1964b:291. (initiation of the P. corsicana species group); Illies, 1966:243. (catalog); Meinander, 1967:45. (new records from Morocco); Berthélemy, 1973:1544. (distribution); Zwick, 1978:33. (composition of the P. corsicana group); Giudicelli &amp; Dakki, 1984:54 (distribution); Nicolai, 1985:249. (biogeography and composition of the P. corsicana group); Gagneur &amp; Aliane, 1991:312. (ecology, distribution); Zhiltzova, 2003:227. (composition of the P. corsicana group); Vinçon &amp; Zhiltzova, 2004:193. (composition of the P. corsicana group); Lounaci &amp; Vinçon, 2005:117. (new records from Algeria and Morocco, distribution); Mur{nyi, 2007:28. (comparison, distribution).</p> <p>Nemoura (Protonemura) algirica auctt., nec Aubert, 1956; Aubert, 1961 (partim):218. (new records from Morocco).</p> <p>Material examined. Morocco, High Atlas: M’Goun Massif, spring of Asif M’Goun, 2500 m, 17-VI-1954, 1♂ 1♀; Siroua Massif, Asif Siroua, above 2900 m, 29- VI-1954, 1♂ 4♀ (Vaillant leg, Aubert coll. ZML, published in Aubert 1956); Toufliht, 50 Km from Marrakech  Ouarzazate, 30-III-1988, 2♂; 68 Km from Marrakech  Ouarzazate, 1500 m, 30-III-1988, 1♂ (S{nchez- Ortega leg, GUC); tributary of Oued Ouarzazate, 116 Km from Marrakech  Ouarzazate, 2600 m, 9-III-1989, 4♂ 3♀ (Ropero &amp; Peña leg, GUC); Tizi n’Test pass, above Tjoukak, 24-II-1996, 3♂ (Vinçon leg, CGV); Tizi n’Tichka pass, 10-IV-1997, 7♂ 6♀ (Luzón leg, GUC); southern slope of Tizi n’Test pass, big spring and brook, 1900 m, 22-I-2006, 9♂ 5♀ - 4♂ 4♀, 1♀ larva (Vinçon leg, HNHM; 1♂ and 1♀ terminalia prepared for SEM); 21-I-2006, 1♂ 2♀ larvae, 1♀ exuviae (Vinçon leg, HNHM; 1♀ larva prepared for SEM); same brook, 1600 m, 22-I-2006, 2♂ 2♀ (Vinçon leg, CGV); above Oukaimeden ski station, 2800 m, 3-VI-2006, 5♂ 4♀; below Oukaimeden, Ait El Kake village, 2150 m, 3-VI-2006, 5♂ 10♀; below Oukaimeden, small torrent on the left side of the road, 2000 m, 3-VI-2006, 5♂ 9♀ (Vinçon leg, CGV). Morocco, Middle Atlas: Azrou, 3-V-1960, 4♀ macropterous (labeled P. algirica Aubert 1960 and Prot. spec Zwick 1982, published as N. (P.) algirica in Aubert 1961); Ifrane, Oued Ifrane, 1700 m, 3-V-1960, 26♂ 13♀ 3 larvae (Besuchet leg, Aubert coll. ZML). Morocco, Rif: R. Achraf, 4 Km from Ketama, 1400 m, 7-VI-1992, 1♀ (Azzouz leg, GUC); above Tétouan, above Sefliane, brook, 22-II-1996, 1♂ (Vinçon leg, CGV). Spain, Rif: Ceuta, Embalse del Renegado, 85 m, 14-III-1997, 1♂; 20-IV-1997, 1♀ (Moro &amp; Tierno de Figueroa leg, GUC). Algeria, Tlemcen Region: Oued Tlemcen, 31-V-1954, 5♂ 4♀ (Vaillant leg, Aubert coll. ZML, published in Aubert 1956).</p> <p>Male genitalia (Figs. 3, 9-10, 16-19). Paraprocts: Median lobe with a rounded base and a blade-shaped expansion with variable length (Aubert 1956: Figs. 9- 10); outer lobe sclerite widely enlarged at the tip, with a rounded dorsal expansion carrying few strong spines (1–7) and a rather long and thin ventral expansion, curved inwards, and ending in one or two spines (Fig. 19 and Aubert 1956: Fig. 9, Raušer 1963: Fig. 3a). Epiproct: Thin, with its sides slightly convex, ending progressively in an oval tip; terminal filament of moderate size and bifid at the tip (Figs. 10, 16). In side view the terminal filament is about parallel to the epiproct but it can be slightly rising due to the condition of the specimen (Figs. 3, 9). Lateral sclerites thin, visible by transparency, and slightly curved upwards on each side of the epiproct’s tip. Ventral sclerite with a prominent bulge partly covered with two rows of long spines forming a ‘V’ shape in ventral view (Fig. 3 and cf. P. pectinata Berthélemy &amp; Dia 1982: Fig. 23). Sternite IX: Vesicle racket-shaped and widening at the tip (Fig. 17 and Aubert 1956: Fig. 7), but this character is rather variable.</p> <p>Female genitalia (Figs. 20-21). Subgenital plate slightly concave, with rounded vaginal lobes. Sternite VII with a postero-median pigmented pregenital plate.</p> <p>Mature larva (Figs. 22-27, 75). Body relatively slender, body length 6.5–9.0 mm. General colour brown. Pilosity distinct. Scales present only in the dorsal midline of the femora and on the tarsi. Legs typical of the genus, tibiae slightly longer than femora; width of hind femora less than one third of its length. Head stout, brown with dark patches. The pronotum is subtrapezoidal, with distinct granules and rounded corners; it is slightly narrowing towards the posterior margin and its length is more than two thirds of its maximum width. Cervical gills simple, the longest one equivalent to the width of the fore coxa. Wing pads shortened or of typical length for the genus. Abdomen relatively slender, integument light and matt, first 6 abdominal segments divided by pleura. Posterior margin of sternite IX of the mature male larva triangular, tip rounded; paraprocts slightly elongated but not pointed (Fig. 75 and Aubert 1956: Figs. 13-14). Genital opening well visible on the mature female larva, and placed under the anterior half of sternite VIII; paraprocts not pointed (Aubert 1956: Fig. 15). Cerci long, with more than 30 segments; segment sides nearly parallel, the width of segments 13–17 is three fifths of their length.</p> <p>Pilosity: Head with dense, stout bristles, and a few sensilla; the eyes bear small setae between the ocelli. Antennal segments with short pilosity. Pronotum with dense, stout and blunt bristles, and a few sensilla. Margin of the pronotum bearing acute bristles, the length of the longest ones is less than 1/15 of the pronotum’s width (Fig. 22). The bristles on the anterior corners of the meso and metanotum are as long as the marginal bristles of the pronotum. The setae placed in lines on the wing pads are short and blunt. Legs with dense setation. All femora bear both short and long, acute bristles and a few thin hairs. Long bristles occur mostly on the outer surface; on hind femora they are placed on the apical half. Bristles not in a regular arrangement; the longest ones reach one fourth of the femur’s width on the first pair, one fifth on the hind legs (Figs. 23-24). A bald median line is conspicuous on the dorsal surface of all femora and it is covered with rounded scales. Tarsi relatively slender, covered with thin hairs and bristles, metatarsi with triangular scales on the dorsal surface; apical spike of tibiae short. Tergal segments with acute bristles and thin hairs. Paired spines on the posterior margin acute, not much longer than the other spines of the row; on tergite V they reach more than one fourth of the segment’s length (Fig. 25). Distal margin with tiny triangular spikes around the row of bristles. Cercal segments with acute bristles, blunt or thin ones occur only in the apical whorl (Figs. 26-27). The apical whorl on segments 13-15 is a set of 13–16 strong, acute spikes mixed with short, blunt and short, thin setae. Longest bristles reach one third of the segment’s length on segments 13–15.</p> <p>Affinities. In the male, the outer lobe of the paraprocts has a characteristic ventral finger-shaped expansion, and the epiproct is slender at the tip. In Protonemura algirica, P. berberica and P. dakkii sp. n., it is stronger at the tip, with wide rounded apex. The female is very close to that of P. dakkii sp. n. The larva is very similar to those of P. dakkii sp. n. and P. berberica, and only the pharate males can be identified with sure, on the basis of the male imago terminalia under the larval skin. However, it differs by its darker habitus and the frequency of stout, blunt bristles on the pronotum, tergites and femurs.</p> <p>Geographical distribution and ecology. P. talboti is a west Maghrebin species inhabiting the Rif, the Middle and High Atlas, and the Tlemcen Mountains in western Algeria (Fig. 6). It occurs in mountain springs and brooks at various altitudes (85–2900 m). The specimens collected in the highest localities of the High Atlas are brachypterous.</p> </div>	http://treatment.plazi.org/id/4D4A87AD8E3B5851FC6CF943FB6AFE71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Vinçon, Gilles;Mur, vid;nyi	Vinçon, Gilles, Mur, vid, nyi (2009): Contribution To The Knowledge Of The Protonemura Corsicana Species Group, With A Revision Of The North African Species Of The P. Talboti Subgroup (Plecoptera: Nemouridae). Illiesia 5 (7): 51-79, DOI: http://doi.org/10.5281/zenodo.4759484
4D4A87AD8E375857FCAAFE0DFC31FD76.text	4D4A87AD8E375857FCAAFE0DFC31FD76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protonemura algirica subsp. algirica algirica (Aubert 1956	<div><p>Protonemura algirica algirica Aubert, 1956</p> <p>(Figs. 6, 13-14, 28-33)</p> <p>Nemoura (Protonemura) algirica Aubert, 1956:424. (original description from Algeria and Morocco); Aubert 1961 (partim):218. (new records from Morocco). Protonemura algirica: Raušer, 1963:804. (comparative drawings, distribution map); Aubert, 1964a:72. (comparison); Aubert, 1964b:291. (initiation of the P. corsicana species group); Illies, 1966:223. (catalog); Berthélemy, 1973:1544. (distribution); Zwick, 1978:33. (composition of the P. corsicana group); Nicolai, 1985:249. (biogeography and composition of the P. corsicana group); Gagneur &amp; Aliane, 1991:323. (distribution); S{nchez- Ortega &amp; Azzouz, 1998:452. (new records from Morocco, but probably belonging to P. berberica or P. dakkii sp. n.); Vinçon &amp; S{nchez- Ortega, 1999:233. (comparison); Zhiltzova, 2003:227. (composition of the P. corsicana group); Vinçon &amp; Zhiltzova, 2004:193. (composition of the P. corsicana group); Lounaci &amp; Vinçon, 2005:(new records from Algeria, ecology, distribution); Mur{nyi, 2007:28. (comparison, distribution).</p> <p>Material examined. Algeria, Atlas de Blida: Oued Chiffa, ‚ Ruisseau des Singes‛ brook, ‚ Rocher des Singes‛, 23-VI-1953, 2♂ 1♀ (written Paralectotypes on the label, Vaillant leg, Aubert coll. ZML). Algeria, Kabylie: Tala Guilef, 16-IV-1953, 1♀ (written Paralectotypes on the label, Vaillant leg, Aubert coll. ZML); Djurdjura Massif, Oued Aïssi, 480 m, 30-IV- 1985, 2♂ 1♀; 14-V-1985, 2♂ 1♀ - 1♂ (HNHM); 23-IV- 1986, 2♂ 1♀; 25-VI-1986, 3♂ 6♀; 20-IV-1993, 2♂ 2♀, 1♂ larva (larva pharate, but terminalia of the larval skin missing); Oued Aïssi, 140 m, 21-VIII-1985, 7♂ 8♀ (Lounaci leg, CGV); Central Djurdjura Massif, Tikjda, small torrent, 1470 m, 20-IV-1993, 4♂ 4♀, 2♂ nymph; Eastern Djurdjura Massif, Assif Sahel, 1000 m, 17-V- 1994, 1♂ nymph (Lounaci leg, CGV).</p> <p>Male genitalia (Figs. 13-14, 28-31). Paraprocts: Median lobe with a rather long sub-triangular base and a long blade-shaped expansion. Outer lobes sclerite widely enlarged at the tip; with about two spines on the dorsal expansion and one or two spines on the ventral expansion (Fig. 31 and Aubert 1956: Fig. 18, Raušer 1963: Fig. 3b); few scattered spines (1- 3) are sometimes located between the dorsal and ventral expansions. Epiproct: Strong, with its sides about parallel, ending abruptly in a rounded tip; well visible terminal filament rather short, bifid at the tip (Figs. 13, 28). In side view the terminal filament is about parallel to the epiproct (Fig. 14). Ventral sclerite with a prominent bulge partly covered with two rows of long spines forming a ‘V’ shape in ventral view. Sternite IX: Vesicle long and narrow (Fig. 29 and Aubert 1956: Fig. 16).</p> <p>Female genitalia (Figs. 32-33). Subgenital plate wide, bilobed. Vaginal lobes mostly hidden by the subgenital plate and partly visible on each side of the plate. Sternite VII without postero-median pigmented pregenital plate.</p> <p>Mature larva. Body shape, coloration and proportions similar to the larva of Protonemura algirica bejaiana ssp. n., as described below, and agreeing with the original description (Aubert 1956). Distribution and type of pilosity also similar, though the overall bluntness of the bristles appears not so pronounced.</p> <p>Affinities. In the males, Protonemura algirica algirica is mainly distinguished from P. talboti by the shape of the epiproct, ending abruptly in P. algirica and progressively in P. talboti. The apex of the outer lobe of the paraprocts is also a good distinctive character. P. dakkii sp. n. is distinguished from P. algirica by the terminal filament of the epiproct rising upwards and by the apex of the outer lobe of the paraprocts carrying a group of spines only on its posterior edge. In the females, the subgenital plate of P. algirica is thicker than that of P. talboti or P. dakkii sp. n. On sternite VII, the postero-median pigmented plate is lacking in P. algirica and present in P. talboti, P. dakkii sp. n. and P. berberica. Affinities of the larva are given below, for P. algirica bejaiana ssp. n.</p> <p>Geographical distribution and ecology. Protonemura algirica algirica is known with certainty only from the central part of Algeria (Atlas of Blida and Djurdjura) (Fig. 6). Its presence in eastern Algeria (Aubert 1956: Aurès Massif) should be confirmed by the capture of males. The Tunisian specimens belong to P. algirica bejaiana ssp. n. The Moroccan specimens from the Rif belong to P. berberica or P. talboti (Aubert, S{nchez- Ortega and Berthélemy collections), those from the Middle Atlas belong to P. dakkii sp. n. or P. talboti (Aubert, Berthélemy and Dakki collections) and those from the High Atlas remain problematic since the Moroccan paralectotypes (ZML) are presently lost and since all our specimens from the High Atlas belong to P. talboti. Therefore, the occurrence of P. algirica in Morocco is uncertain and needs to be confirmed. In the Djurdjura Massif (Kabylia), P. algirica algirica occurs in different kinds of mountain water courses (480–1300 m). The emergence period is from spring to summer (IV– VIII) (Lounaci &amp; Vinçon 2005).</p> </div>	http://treatment.plazi.org/id/4D4A87AD8E375857FCAAFE0DFC31FD76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Vinçon, Gilles;Mur, vid;nyi	Vinçon, Gilles, Mur, vid, nyi (2009): Contribution To The Knowledge Of The Protonemura Corsicana Species Group, With A Revision Of The North African Species Of The P. Talboti Subgroup (Plecoptera: Nemouridae). Illiesia 5 (7): 51-79, DOI: http://doi.org/10.5281/zenodo.4759484
4D4A87AD8E31584BFCA7FD11FEA9FCC4.text	4D4A87AD8E31584BFCA7FD11FEA9FCC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protonemura algirica subsp. bejaiana Vincon & Mur	<div><p>Protonemura algirica bejaiana Vinçon &amp; Mur {nyi ssp. n.</p> <p>(Figs. 6, 15, 34-45, 76)</p> <p>Protonemura algirica ssp. nov.: Berthélemy, 1973:1544. (distribution and ecology).</p> <p>Material examined. Holotype male: Tunisia, Aïn Draham: Chabet El Khantra, Oued el Lil tributary, above Ben Metir dam, 540 m, 9-V-1969; Paratypes, same locality: 2-V-1969, 3♂ 2♀, 1♂ larva, 1 exuviae (larva pharate, terminalia of the larval skin missing; exuviae having thoracic parts only); 9-V-1969, 1♀; 10- V-1969, 1♂ 1♀; 16-XI-1969, 2♀; 10-I-1970, 2♂ 3♀ - 1♂ 1♀ larvae, 1♂ 1♀ exuviae (HNHM; ♀ larva prepared for SEM, ♂ larva pharate, terminalia of the larval skin missing); 12-II-1970, 2♂ 2♀; 18-IV-1970, 2♀; 7-VI- 1970, 1♂; 13-XI-1970, 1♀ nymph; 30-XII-1970, 1♂. Holotype and 2♂ 2♀ paratypes are deposited in the ZML, other paratypes held in the CGV.</p> <p>Other material: Tunisia, Aïn Draham: Oued ed Demène, near Ben Metir dam, 450 m, 19-IV-1970, 1♂ - 1♂ (HNHM); 9-V-1970, 4♀; 7-VI-1970, 1♀; Oued ed Demène at 630 m, 17-IV-1970, 1♀; Oued el Lebga, Oued el Lil tributary, above Ben Metir dam, 450 m, 19-IV-1970, 1♀ nymphae (Berthélemy coll. CGV).</p> <p>Description. Medium-sized Protonemura of the corsicana group. Body length, male: 6.9–7.5 mm; female: 7.1–8.3 mm; forewing length, male: 8.7–9.2 mm; female: 8.7–9.6 mm. Head brownish with two yellow triangular areas between eyes and ocelli; antennae brown. Pronotum brown with dark pattern, its lateral sides light brown. Legs yellowish with brown longitudinal marks. Abdomen light brown. Gills short, without subterminal constriction.</p> <p>Male (Figs. 15, 34-37). Tergites I-VI simple; tergite VII with few spines scattered medially on its posterior edge; tergite VIII similar but the spiny area deeply extends near the middle of the segment, in several specimens this group of spines is about separated in two spiny contiguous fields; tergite IX with two strong triangular groups of spines separated by a median smooth strip (Fig. 34); tergite X with several spines surrounding the tip of the epiproct. Paraprocts (Fig. 37): Inner lobe blade-shaped, partly hidden under the expansion of the hypoproct. Median lobe wide, with well developed globular membranous apex covered with thin setae; sclerotized base slightly longer than wide and ending in a gently curved, blade-shaped expansion which is more or less long, about reaching the tip of the outer lobe. Outer lobe sclerite enlarged at the tip, with a rounded, smooth and light posterior expansion without any spine; the anterior part of the lobe ends in a unique sharp spine, curved inwards. Epiproct: Rather strong and pale, slightly enlarging towards the tip in dorsal view and ending abruptly in a rounded tip (Fig. 15). A smooth bilobed expansion is visible at the tip. The epiproct terminal filament is rather short, bifid at the apex; in side view the filament is about parallel to the epiproct. Dorsal sclerite of the epiproct forked, well visible in dorsal view. The two lateral branches are getting nearer and narrowing toward the epiproct’s tip, where they are curving upwards and meeting close to the projection of the terminal filament (Fig. 15). Ventral sclerite with a prominent bulge partly covered with two rows of long spines. In ventral view, the rows of spines are getting nearer towards the apex, forming a ‘V’. Sternite IX: Hypoproct rounded distally, ending into a long tapering extension. Vesicle long, with its sides about parallel (Fig. 35).</p> <p>Female. (Figs. 38-39). Abdomen typical of the genus. Sternite VII: Pregenital plate not sclerotized and therefore hardly visible. Sternite VIII: Subgenital plate well sclerotized and clearly bilobed; vaginal lobes strong, extending on each side of the subgenital plate. Mature larva (Figs. 40-45, 76). Body relatively slender, body length 9.0–10.5 mm. General color brown. Pilosity distinct. Scales present only in the dorsal midline of the femora and on the tarsi. Legs typical of the genus, tibiae slightly longer than femora; width of hind femora less than one third of its length. Head stout, brown with dark patches. The pronotum is subtrapezoidal, with distinct granules and rounded corners; it is slightly narrowing towards the posterior margin and its length is two thirds of its maximum width. Cervical gills simple, the longest one equivalent to the width of the fore coxa. Wing pads of typical length for the genus. Abdomen relatively slender, integument light and matt, first 6 abdominal segments divided by pleura. Posterior margin of sternite IX of the mature male larva triangular, sharply pointed; paraprocts not pointed (Fig. 76). Genital opening well visible on the mature female larva, and placed under the anterior half of sternite VIII; paraprocts not pointed. Cerci long, with more than 35 segments; segment sides nearly parallel, the width of segments 13–17 is three fifths of their length.</p> <p>Pilosity: Head with dense, stout bristles and a few sensilla. Antennal segments with short pilosity. Pronotum with dense, stout bristles, a few sensilla and thin hairs. Margin of the pronotum bearing blunt bristles, the length of the longest ones is less than 1/15 of the pronotum’s width (Fig. 40). The bristles on the anterior corners of the meso and metanotum are as long as the marginal bristles of the pronotum. The setae placed in lines on the wing pads are short and blunt. Legs with dense setation. All femora bear both short and long, blunt bristles and thin hairs. Long bristles occur mostly on the outer surface; on hind femora they are placed on the apical half. Bristles not in a regular arrangement; the longest ones reach one fifth of the femur’s width on all legs (Figs. 41- 42). A bald median line is conspicuous on the dorsal surface of all femora and it is covered with rounded scales. Tarsi relatively stout, covered with thin hairs and bristles, metatarsi with triangular scales on the dorsal surface; apical spike of tibiae short. Tergal segments with short, blunt bristles and a few thin hairs. Paired spines on the posterior margin blunt or hardly acute, two times longer than the other spines of the row; on tergite V they reach one fourth of the segment’s length (Fig. 43). Distal margin lacks tiny triangular spikes around the row of bristles. Cercal segments with acute bristles, blunt or thin ones occur only in the apical whorl (Figs. 44-45; apical row scanty on Fig. 45). The apical whorl on segments 13-15 is a set of 10–12 strong, acute spikes mixed with short, blunt and short, thin setae. Longest bristles reach one fifth of the segment’s length on segments 13–15.</p> <p>Etymology. Protonemura algirica bejaiana ssp. n. is named after Beja, main town of the Khroumiry region, in Western Tunisia.</p> <p>Affinities. Protonemura algirica bejaiana s sp. n. mainly differs from P. algirica algirica by the shape of the outer lobe of the paraproct, without any spine on its dorsal expansion instead of carrying two spines or more in P. algirica algirica. The ventral part of the paraproct’s outer lobe always ends in a unique spine instead of one or two spines in P. algirica algirica. In the female, the vaginal lobes are more exposed on each side of the subgenital plate than in P. algirica algirica where they are often more hidden under the plate. P. algirica algirica and P. algirica bejaiana s sp. n. are also closely related to P. albanica from which they differ by the following features: in the male, the epiproct is not prolonged by a membranous expansion and its filament is much longer; in the female, the vaginal lobes are strongly extending in front of the plate. The overall shape and the terminalia of the larva are very similar to those of the other North African species, but the P. algirica larva can be separated by the blunt bristles on the margin of pronotum, tergites and femurs. In addition, it differs from P. dakkii sp. n. and P. berberica by its darker habitus. The nominal subspecies seems to have not so pronounced blunt bristles, but this feature should be confirmed with the study of additional material.</p> <p>Geographical distribution and ecology. Protonemura algirica bejaiana ssp. n. inhabits the north-western part of Tunisia (Khroumiry Mountains) (Fig. 6), where it occurs in brooks and brooklets at moderate altitudes (450–600 m). The adults emerge in winter and spring (XI– V).</p> </div>	http://treatment.plazi.org/id/4D4A87AD8E31584BFCA7FD11FEA9FCC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Vinçon, Gilles;Mur, vid;nyi	Vinçon, Gilles, Mur, vid, nyi (2009): Contribution To The Knowledge Of The Protonemura Corsicana Species Group, With A Revision Of The North African Species Of The P. Talboti Subgroup (Plecoptera: Nemouridae). Illiesia 5 (7): 51-79, DOI: http://doi.org/10.5281/zenodo.4759484
4D4A87AD8E2D584FFED8FCDFFCA6FCC4.text	4D4A87AD8E2D584FFED8FCDFFCA6FCC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protonemura berberica Vincon	<div><p>Protonemura berberica Vinçon &amp; S{nchez- Ortega, 1999</p> <p>(Figs. 1-2, 4, 6, 11-12, 46-58)</p> <p>Protonemura berberica Vinçon &amp; S{nchez- Ortega, 1999:231. (original description from Morocco); Vinçon &amp; Zhiltzova, 2004:193. (composition of the P. corsicana group); Lounaci &amp; Vinçon, 2005:(distribution); Mur{nyi, 2007:28. (distribution)</p> <p>Nemoura (Protonemura) algirica auctt., nec Aubert, 1956: Aubert, 1961 (partim):218. (records from the Moroccan Rif).</p> <p>Protonemura sp.: S{nchez- Ortega &amp; Azzouz, 1998:452. (records from the Moroccan Rif).</p> <p>Material examined. Morocco, Rif: Ketama, Djebel Tidirehine, 1800 m, 24-IV-1960, 1♂ nymph and 7 larvae - 2♀ 1♂ larvae (1♀ larva prepared for SEM, ♂ larva pharate, terminalia of the larval skin missing) (Besuchet leg, labeled P. algirica det. Aubert, Aubert coll. ZML, published as N. (P.) algirica in Aubert 1961); spring 14 Km South West Ketama, 1500 m, 21- VII-1980, 2♂ 1♀ (Berthélemy coll. CGV); 14 km from Boohivid, 3-VI-1992, 2♂ 1♀; Loukous brook, 300 m, 5-VI-1992, 1♂ 2♀ (dried specimens, labeled P. algirica Azzouz 1992 det and leg, GUC); Spring between Bab- Besen and Ketama, 1600 m, 22-II-1996, 2♂ 5♀; above Azila Ketama, Djebel Tidirehine, 1600-1800 m, 4-VI- 2006, 3♂ 9♀; Djebel Tidirehine, 2000-2100 m, 4-VI- 2006, 6♂ 16♀ 1 larva (Vinçon leg, CGV) - 3♂ 3♀ (HNHM; 1♂ and 1♀ terminalia prepared for SEM). Imago (Figs. 1-2, 4, 11-12, 46-51). A detailed description is already given in Vinçon &amp; S{nchez- Ortega (1999). The male is well characterized by the apex of the paraproct’s outer lobe, bordered by a regular comb-shaped fringe of spines (Fig. 49), and by the terminal filament of the epiproct risen upwards (Figs. 4, 11, 48). This filament is short and bifid at the tip (Figs. 1-2, 4).</p> <p>Mature larva (Figs. 52-58). Body relatively slender, body length 9.0–10.5 mm (Fig. 52). General color pale, yellowish brown. Pilosity pronounced but not distinct. Scales present only in the dorsal midline of the femora and on the tarsi. Legs typical of the genus, tibiae slightly longer than femora; width of hind femora more than one third of its length. Head stout, pale with indistinct, slightly darker patches. The pronotum is subtrapezoidal, with hardly visible granules and rounded corners; it is slightly narrowing towards the posterior margin and its length is two thirds of its maximum width. Cervical gills simple, the longest one equivalent to the width of the fore coxa. Wing pads of typical length for the genus. Abdomen relatively slender, integument light and matt, first 6 abdominal segments divided by pleura. Terminalia of the larval skin is missing on the single examined pharate male. Genital opening well visible on the mature female larva and placed under the anterior half of sternite VIII; paraprocts not pointed. Cerci long, with more than 30 segments; segment sides nearly parallel, the width of segments 13–17 is three fourths of their length.</p> <p>Pilosity: Head with dense, stout but acute bristles, and a few sensilla; eyes bearing small setae between the ocelli. Antennal segments with short pilosity. Pronotum with dense, stout but acute bristles, and a few sensilla. Margin of the pronotum bearing acute bristles, the length of the longest ones is less than 1/15 of the pronotum’s width (Fig. 53). The bristles on the anterior corners of the meso and metanotum are as long as the marginal bristles of the pronotum. The setae placed in lines on the wing pads are short but acute. Legs with dense setation. All femora bear both short and long, acute bristles and a few thin hairs. Long bristles occur mostly on the outer surface; on hind femora they are placed all along its length. Bristles not in a regular arrangement; the longest ones reach one fourth of the femur’s width on all legs (Figs. 54-55). A bald median line is conspicuous on the dorsal surface of all femora and it is covered with rounded scales. Tarsi relatively slender, covered with thin hairs and bristles, metatarsi with triangular scales on the dorsal surface; apical spike of tibiae short. Tergal segments with short bristles and a few thin hairs. Paired spines on the posterior margin acute, not much longer than the other spines of the row; on tergite V they reach one fourth of the segment’s length (Fig. 56). Distal margin lacking tiny triangular spikes around the row of bristles. Cercal segments with acute bristles, blunt or thin ones occur only in the apical whorl (Figs. 57-58). The apical whorl on segments 13-15 is a set of 9–11 strong, acute spikes mixed with short, blunt and short, thin setae. Longest bristles reach two fifths of the segment’s length on segments 13–15.</p> <p>Affinities. Protonemura berberica is closely related to P. dakkii sp. n. by the shape of the epiproct, with its terminal filament rising upwards in lateral view; this character is not observed in the other species of the P. talboti subgroup. The male clearly distinguishes by the shape of the paraproct’s outer lobe with its regular fringe of spines, and the female by its strongly developed vaginal lobes and the presence of a strongly pigmented pregenital plate on sternite VII. This plate is also present in P. talboti, but absent or hardly visible in the other mentioned species. The larva is very similar to those of P. dakkii sp. n. and P. talboti, and only the pharate males can be identified with sure, on the basis of the male imago terminalia under the larval skin. However, it differs from P. talboti by its pale habitus and the scarcity of stout, blunt bristles on the pronotum, tergites and femurs, from P. dakkii sp. n. by the longer bristles on the margin of the pronotum.</p> <p>Geographical distribution. This species is probably endemic of the Moroccan Rif (Fig. 6). The previous citations of P. algirica from the Rif all concern P. berberica, as it was verified by the study of adults and male nymphs from the collections of Aubert, Berthélemy, S{nchez- Ortega and Tierno de Figueroa.</p> </div>	http://treatment.plazi.org/id/4D4A87AD8E2D584FFED8FCDFFCA6FCC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Vinçon, Gilles;Mur, vid;nyi	Vinçon, Gilles, Mur, vid, nyi (2009): Contribution To The Knowledge Of The Protonemura Corsicana Species Group, With A Revision Of The North African Species Of The P. Talboti Subgroup (Plecoptera: Nemouridae). Illiesia 5 (7): 51-79, DOI: http://doi.org/10.5281/zenodo.4759484
4D4A87AD8E295840FF03FCDBFD51FAF6.text	4D4A87AD8E295840FF03FCDBFD51FAF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protonemura dakkii Vincon & Mur	<div><p>Protonemura dakkii Vinçon &amp; Mur {nyi sp. n.</p> <p>(Figs. 6-8, 59-74, 77-79)</p> <p>Nemoura (Protonemura) algirica auctt., nec Aubert, 1956: Aubert 1961 (partim):218. (records from Morocco). Nemoura (Protonemura) talboti auctt., nec Nav{s, 1929: Aubert 1961 (partim):219. (records from Morocco).</p> <p>Material examined: Holotype male: Morocco, Middle Atlas: Ifrane, Ifrane Oued, 1700 m, 3-V-1960, micropterous (Besuchet leg, labeled P. algirica by Aubert, Aubert coll. ZML, published as N. (P.) algirica in Aubert 1961); Paratypes: same date and locality, 5♂ 5♀ micropterous, 2 larvae - 2♂ 2♀, 1 non matured larva. Holotype and 2♂ 2♀ paratypes are deposited in the ZML, other paratypes held in the CGV.</p> <p>Other material: Morocco, Middle Atlas: Ain Leuh, 4-V-1960, 1♀ macropterous (Besuchet leg, labeled P. talboti Aubert 1960, Aubert coll. ZML, published as N. (P.) talboti in Aubert 1961); Azrou, 3-V-1960, 2♀ macropterous (leg. Besuchet, labeled P. talboti Aubert 1960 and Protonemura spec. Zwick 1982, Aubert coll. ZML, published as N. (P.) algirica in Aubert 1961); Oriental Middle Atlas, Imouzzer des Marmoucha, high Oued Al Mahçar, karstic permanent spring, labeled P. algirica, SM (JI), 20-VII-1981, 3♂ 5♀ macropterous, 5 larvae (Dakki leg, Berthélemy coll., CGV); from them: 1♂ 1♀, 1♂ 1♀ larvae (HNHM; female larva prepared for SEM); karstic temporary spring of the same hydrographic basin, ‚Aghbalou Abekhbakh‛, labeled P. algirica, AA (JN 83), 2♂ macropterous, 20-VI-1983 (Dakki leg, Berthélemy coll., CGV).</p> <p>Description (Fig. 65). Medium-sized Protonemura of the corsicana group. Body length, male: 5.6–6.1 mm; female: 7.5–8.4 mm; forewing length, male: 0.2 (micropterous specimens) – 6.2 mm; female: 0.2 (micropterous specimens) – 9.0 mm. Head dark brown with a median rounded yellow spot between the ocelli; antennae brown. Pronotum brown with dark pattern, its anterior corners light brown. Legs and abdomen light brown. Gills short, without subterminal constriction.</p> <p>Male (Figs. 7-8, 59-62). Tergites I-VII simple; tergite VIII with a median rounded group of spines on its posterior edge; tergite IX with two oval groups of spines widely separated by a space about as wide as the width of each group of spines (Fig. 59); tergite X with two groups of spines placed on each side of the epiproct’s tip. Paraprocts (Fig. 62): Inner lobe bladeshaped, partly hidden under the expansion of the hypoproct. Median lobe wide, with well developed globular membranous apex covered with thin setae; sclerotized base slightly longer than wide and ending in a gently curved, blade-shaped expansion which is more or less long, about reaching the tip of the outer lobe. Outer lobe sclerite enlarged at the tip, with a rounded posterior expansion with many (more than 5) spines; the anterior part of the lobe weakly produced and bearing no spine. Epiproct: Rather strong and pale, slightly enlarging towards the tip in dorsal view and ending abruptly in a rounded tip (Figs. 8, 59). A smooth bilobed expansion is visible at the tip. The epiproct terminal filament is of moderate size, bifid at the apex; in side view the filament is risen upwards (Figs. 7, 61). Dorsal sclerite of the epiproct forked, well visible in dorsal view. The two lateral branches are getting nearer and narrowing toward the epiproct’s apex; they are curving upwards on each side of the epiproct’s tip where they meet close to the terminal filament (Fig. 8). Ventral sclerite with a prominent bulge partly covered with two rows of long spines (Fig. 7). In ventral view, the rows of spines are getting nearer towards the apex, forming a ‘V’. Sternite IX: Hypoproct rounded distally ending in a short, slightly tapering extension. Vesicle short, slightly widening at the tip (Fig. 60).</p> <p>Female (Figs. 63-64). Abdomen typical of the genus. Sternite VII: Pregenital plate very slightly sclerotized and therefore hardly visible. Sternite VIII: Subgenital plate well sclerotized, posterior margin rectilinear, very slightly indented; vaginal lobes strong, extending on each side of the subgenital plate. Sternite IX simple.</p> <p>Mature larva (Figs. 66-74, 77). Body relatively stout, body length 7.0–9.0 mm (Fig. 66). General color pale, yellowish brown. Pilosity not distinct. Scales present only in the dorsal midline of the femora and on the tarsi. Legs typical of the genus, tibiae slightly longer than femora; width of hind femora equivalent to one third of its length. Head stout, brown with dark patches. The pronotum is subtrapezoidal, with distinct granules and rounded corners; it is slightly narrowing towards the posterior margin and its length is two thirds of its maximum width. Cervical gills simple, the longest one is equivalent to the width of the fore coxa (Fig. 77). Wing pads shortened or completely absent. Abdomen relatively stout, integument light and matt, first 6 abdominal segments divided by pleura. Posterior margin of sternite IX of the mature male larva triangular, weakly pointed; paraprocts not pointed (Fig. 74). Genital opening well visible on the mature female larva and placed under the anterior half of sternite VIII; paraprocts not pointed (Fig. 73). Cerci of the matured larva with about 20 segments; they are longer, with more than 30 segments, in the not matured larva.</p> <p>Pilosity: Head with dense, stout but acute bristles and a few hairs and sensilla; the eyes bear small setae between the ocelli (Fig. 72). Antennal segments with short pilosity. Pronotum with stout bristles, a few sensilla and thin hairs. Margin of the pronotum bearing acute bristles, the length of the longest ones is less than 1/20 of the pronotum’s width (Fig. 67). The bristles on the anterior corners of the meso and metanotum are as long as the marginal bristles of the pronotum. The setae placed in lines on the wing pads are short and blunt. Legs with dense setation. All femora bear both short and long, blunt bristles and thin hairs. Long bristles occur mostly on the outer surface; on hind femora they are placed all along its length. Bristles not in a regular arrangement; the longest ones reach one fifth of the femur’s width on all legs (Figs. 68-69, 71). A bald median line is conspicuous on the dorsal surface of all femora and it is covered with rounded scales. Tarsi relatively stout, covered with thin hairs and bristles, metatarsi with triangular scales on the dorsal surface; apical spike of tibiae short. Tergal segments with acute bristles and thin hairs. Paired spines on the posterior margin acute, hardly longer than the other spines of the row; on tergite V they reach one fourth of the segment’s length (Fig. 70). Distal margin with scarce tiny triangular spikes around the row of bristles. Cercal pilosity of the larva similar to that of the closely related species.</p> <p>Etymology. This species is named in honor of Mohamed Dakki (Rabat Scientific Institute), who has collected this species in several places of the Middle Atlas.</p> <p>Affinities. Protonemura dakkii sp. n. is distinguished from P. algirica and P. berberica by the shape of the epiproct in lateral view and by the shape of the exterior lobe of the paraprocts. The females resemble those of P. algirica and P. talboti and therefore the presence of P. dakkii sp. n. in new localities should be confirmed by the capture of males. The larva is very similar to P. berberica and P. talboti, and only the pharate males can be identified with sure, on the basis of the male imago terminalia under the larval skin. However, it differs from P. talboti by its pale habitus and the scarcity of stout, blunt bristles on the pronotum, tergites and femurs, from P. berberica by the shorter bristles on the margin of the pronotum. Larvae of the micropterous form can be separated by the absence of wing pads, as micropterous form are hitherto not known in the other North African species. The diminution of the imaginal ventral sclerites from the normal winged form towards the micropterous form (Figs. 78-79) shows a similar, but lesser degree than it was shown by Berthélemy (1969) in the case of the genus Capnioneura Ris, 1905, and by Vinçon &amp; Pardo (1994) in the case of the genus Leuctra Stephens, 1835.</p> <p>Geographical distribution. This species is probably endemic of the Moroccan Middle Atlas (Fig. 6).</p></div> 	http://treatment.plazi.org/id/4D4A87AD8E295840FF03FCDBFD51FAF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Vinçon, Gilles;Mur, vid;nyi	Vinçon, Gilles, Mur, vid, nyi (2009): Contribution To The Knowledge Of The Protonemura Corsicana Species Group, With A Revision Of The North African Species Of The P. Talboti Subgroup (Plecoptera: Nemouridae). Illiesia 5 (7): 51-79, DOI: http://doi.org/10.5281/zenodo.4759484
