taxonID	type	description	language	source
03CD878C823DFFB10D3C5BDB0EB1FB2F.taxon	materials_examined	Lectotype ♀ (National Museum, Prague), Chiriqui, Panama, designation Stark (1998: 557)	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10D3C5BDB0EB1FB2F.taxon	materials_examined	Material examined. Panama, Chiriqui, Totumas Cloud Forest, lodge, 08 ° 53.060 ’ N, 82 ° 41.068 ’ W, 1920 m, 19 - 25 June 2011, E. Fuller, 11 ♂, 4 ♀ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10D3C5BDB0EB1FB2F.taxon	discussion	Comments. This species was previously reported from several Costa Rican provinces and from Chiriqui Province, Panama (Stark 1998).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10D0C5DED0D95F8D0.taxon	materials_examined	Holotype ♂ (Museo de Entomología de la Universidad del Valle, Cali), Rio Azul, Valle del Cauca, Colombia	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10D0C5DED0D95F8D0.taxon	materials_examined	Material examined. Panama, Darién, Rio Tuira, Bocos de Cupe, 18 February 1985, J. Louton, 1 ♂ (USNM). Darién, Rio Tuira between El Real and Rio Pucuro, 16 February 1985, J. Louton, 3 ♂, 3 ♀ (USNM).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10D0C5DED0D95F8D0.taxon	discussion	Comments. The specimens listed above were described by Harper (1992) as “ Anacroneuria sp. A ”. Subsequently, I determined them as A. azul, but these determinations were never published. The records are noteworthy as one of only four Anacroneuria species known from sites in Colombia (Stark et al. 1999; Zúñiga & Stark 2002; Zúñiga et al. 2006; Zúñiga et al. 2007) and also from Mesoamerican sites.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10F965828092FFD05.taxon	materials_examined	Holotype ♂ (USNM), Rio Los Ahogados, 11.3 km ENE Quebrada Grande, Guanacaste, Costa Rica	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10F965828092FFD05.taxon	materials_examined	Material examined. Costa Rica, Heredia, 9 km NE Vara Blanca, 10 ° 14 ’ N, 84 ° 06 ’ W, 1450 - 1550 m, 15 March 2005, S. M. Clark, D. R. Clark, 1 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10F965828092FFD05.taxon	discussion	Comments. This species is recorded from Honduras, Panama and five Costa Rican provinces (Alajuela, Guanacaste, Limón, Puntarenas and San Jose). This is the first report from Heredia Province.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10F4D5BFF0839FB2F.taxon	materials_examined	Holotype ♀ (Cornell University), Barro Colorado Island, Panama	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10F4D5BFF0839FB2F.taxon	materials_examined	Material examined. Panama, Chiriqui, Totumas Cloud Forest, lodge, 08 ° 53.060 ’ N, 82 ° 41.068 ’ W, 1920 m, 19 - 25 June 2011, E. Fuller, 2 ♂ (MLBM).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10F4D5BFF0839FB2F.taxon	discussion	Comments. This species is known from scattered sites in Costa Rica, Colombia and Panama (Stark 1998). This is apparently the first record from Chiriqui Province.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10FED5D910FF9F9F2.taxon	materials_examined	Holotype ♂ (USNM), Rio Zurqui, Parque Nacional Braulio Carrillo, San Jose, Costa Rica	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10FED5D910FF9F9F2.taxon	materials_examined	Material examined. Costa Rica, Heredia, 9 km NE Vara Blanca, 10 ° 14 ’ N, 84 ° 06 ’ W, 1450 - 1550 m, 15 March 2005, S. M. Clark, D. R. Clark, 1 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB10FED5D910FF9F9F2.taxon	discussion	Comments. This species was previously known from the holotype and four male paratypes, all from the type locality.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB70F955FAE0DACFAA1.taxon	description	(Figs. 1 - 5)	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB70F955FAE0DACFAA1.taxon	materials_examined	Material examined. Holotype ♂, Costa Rica, Guanacaste Province, Estación Biológica Maritza, Rio Tempisquito, 17 March 1989, light trap (USNM).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB70F955FAE0DACFAA1.taxon	description	Adult habitus. General color brown, patterned with pale brown. Head dark brown pigment extending from behind ocelli in a broad patch covering central frons; lappets and antennae dark brown, pale callosities lateral to ocelli distinct (Fig. 1). Median pronotal band yellow, most of disc brown. Wing membrane pale amber, veins brown. Hind femora with pale proximal band covering basal 2 / 3, apical third dark brown. Tibiae banded, pale in median field, dark brown at both ends. Male. Forewing length 12 mm. Hammer thimble shaped, height slightly greater than apical diameter (Fig. 2). Aedeagal apex simple, notched on apical margin; ventral membranous lobes conspicuous (Fig. 3). In lateral aspect, apex is somewhat foot-shaped, but with a slightly bulging process at ankle (Fig. 4). Dorsal keel consists of an obscure series of low chevron-like lines (Fig. 5), but median line absent. Hooks slender. Female. Unknown. Larva. Unknown.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB70F955FAE0DACFAA1.taxon	etymology	Etymology. The species name, based on Guanacaste Province, is used as a noun in apposition.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823DFFB70F955FAE0DACFAA1.taxon	diagnosis	Diagnosis. In the Stark (1998) key this species runs to couplet 18 where an impasse occurs due to the intermediate forewing length. If the impasse is resolved by choosing the greater length (13 mm), no resolution is gained, although a determination of A. marca or A. ventana Stark, 1998 can be forced; if the smaller length (10 mm) is chosen a determination of A. varilla Stark, 1998 is made. In some respects, A. guanacaste is intermediate between these species with a slender, thimble-shaped hammer like that of A. marca, and with an aedeagal apex more similar to that of A. varilla. The new species is distinguished from both by the apical notch in the aedeagal tip (Fig. 3), and by the chevron- like dorsal keel (Fig. 5). The species is probably a near relative of A. flavominuta Jewet, 1958 and other Mexican species (Stark & Kondratieff 2004.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70D025C1C0D47F94B.taxon	materials_examined	Holotype ♂ (USNM), Quebrada Zapilote, Parque Nacional Rincon de la Vieja, Guanacaste, Costa Rica	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70D025C1C0D47F94B.taxon	materials_examined	Material examined. Costa Rica, Cartago, near Tuis, 3000 ’, 16 - 22 June 1993, S. Keller, 1 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70D025C1C0D47F94B.taxon	discussion	Comments. This species has been reported from Honduras, Nicaragua and from Alajuela, Guanacaste and Heredia provinces in Costa Rica (Stark 1998; Stark & Kondratieff 2004). This is the first record from Cartago.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70D165E350913FB00.taxon	materials_examined	Holotype ♂ (Museum National d’Histoire Naturelle, Paris), Costa Rica	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70D165E350913FB00.taxon	materials_examined	Holotype ♀ (Cornell University Insect Collection), Río Santa Ana, British Honduras [Belize], synonymy Stark & Kondratieff, 2004: 26	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70D165E350913FB00.taxon	materials_examined	Material examined. Panama, Chiriqui Province, Rovira. Elvira Farm, 2200 ’, 11 July 1964, A. Broce, 1 ♂ (FSCA). Veraguas, Alto de Piedra, 08 ° 30.9 ’ N, 81 ° 07.0 ’ W, 16 - 18 June 2011, E. Fuller, 1 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70D165E350913FB00.taxon	discussion	Comments. Several Costa Rican provincial records of this species are given in Stark (1998) and additional records are given for Belize, Guatemala, Honduras and Mexico in Stark & Kondratieff (2004). Jewett (1958) lists a male and two female specimens from “ El Volcan, Chiriqui, Panama ” among the specimens he determined as A. sulana, a synonym of A. lineata (Stark & Kondratieff 2004). These records may be valid, however, to my knowledge they have not been examined since the 1950 ’ s. The specimens listed above indicate the species is a valid member of the Panamanian fauna. The specimen from Veraguas has a thimble-shaped hammer rather than nipple shaped and has other subtle features which suggest it may represent a species distinct from A. lineata.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70F995DFC08ECF8B7.taxon	materials_examined	Holotype ♀ (American Museum of Natural History), El Volcan, Chiriqui, Panama	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70F995DFC08ECF8B7.taxon	materials_examined	Material examined. Costa Rica, Heredia, 9 km NE Vara Blancas, 10 ° 14 ’ N, 84 ° 06 ’ W, 1450 - 1550 m, 20 March 2005, S. M. Clark, D. R. Clark, 1 ♂ (BYUC). Same site, 14 March 2005, S. M. Clark, D. R. Clark, 1 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB70F995DFC08ECF8B7.taxon	discussion	Comments. This species was previously reported from Honduras, Panama and from three Costa Rican provinces (Alajuela, Guanacaste, Puntarenas) (Jewett 1958; Stark 1998). These are the first specimens recorded from Heredia Province.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB50FFF5E690DF6FEC2.taxon	materials_examined	Holotype ♂ (USNM), Rio Peje, Heredia, Parque Nacional Braulio Carillo, Costa Rica	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB50FFF5E690DF6FEC2.taxon	materials_examined	Material examined. Costa Rica, Heredia, Chilamate, 24 - 30 July 1993, W. J. Hanson, 3 ♂ (BYUC). Panama, Chiriqui, Totumas Cloud Forest, lodge, 08 ° 53.060 ’ N, 82 ° 41.068 ’ W, 1920 m, 10 - 25 June 2011, E. Fuller, 5 ♀ (BYUC). Chiriqui Province, Rio Caldera, Boquete, 1100 m, 18 June 1973, 1 ♂ (USNM).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C823BFFB50FFF5E690DF6FEC2.taxon	discussion	Comments. This species was previously reported from several Costa Rican provinces, including Heredia (Stark 1998). These are the first specimens recorded from Panama.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB50D6F58BF0E57F992.taxon	materials_examined	Holotype ♂ (USNM), Rio Tempisquito, Estacion Maritza, Guanacaste, Costa Rica	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB50D6F58BF0E57F992.taxon	materials_examined	Material examined. Costa Rica, Guanacaste, 3 km SE Rio Naranjo, 19 - 22 February 1993, F. D. Parker, 1 ♂ (BYUC). Same location, 21 - 30 April 1992, F. D. Parker, 1 ♂ (MLBM). Same location, 18 - 28 April 1993, F. D. Parker, 3 ♂ (BYUC). Same location, 15 - 30 April 1992, F. D. Parker, 1 ♂ (BYUC). Same location, 3 - 6 May 1993, F. D. Parker, 1 ♂ (BYUC). Same location, 12 - 14 May 1993, F. D. Parker, 2 ♂ (BYUC). Same location, 19 May 1993, F. D. Parker, 1 ♂ (BYUC). Same location, 24 - 26 May 1993, 2 ♂ (BYUC). Same location, 27 May 1992, F. D. Parker, 1 ♂ (BYUC). Same location, 28 - 31 May 1993, F. D. Parker, 2 ♂ (BYUC). Same location, 1 - 10 June 1992, F. D. Parker, 2 ♂ (BYUC). Same location, 11 - 20 June 1992, F. D. Parker, 1 ♂ (BYUC). Same location, 14 - 16 June 1993, F. D. Parker, 4 ♂ (BYUC). Same location, 18 - 23 June 1993, F. D. Parker, 5 ♂ (BYUC). Same location, 23 - 27 June 1993, F. D. Parker, 1 ♂ (BYUC). Same location, 21 - 30 June 1992, F. D. Parker, 3 ♂ (BYUC). Same location, 17 - 19 July 1993, F. D. Parker, 1 ♂ (BYUC). Same location, 21 - 31 July 1992, F. D. Parker, 1 ♂ (BYUC). Same location, 8 - 12 August 1993, F. D. Parker, 1 ♂ (BYUC). Same location 1 - 9 October 1992, F. D. Parker, 1 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB50D6F58BF0E57F992.taxon	discussion	Comments. This species was previously known from three sites in Guanacaste Province, Estacion Maritza, Monteverde and Rio Los Ahogados. The new records add another site, also in Guanacaste Province, from which this species is known to occur.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB50D625F4B08B0FA5B.taxon	description	(Figs. 6 - 10)	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB50D625F4B08B0FA5B.taxon	materials_examined	Material examined. Holotype ♂ (pinned), Costa Rica, Alajuela, Rio La Paz, Rt. 9, 7.6 km N Vara Blanca, 10.208 ° N, 84.166 ° W, 1340 m, 13 February 1986, J. Morse (USNM). Paratype: Costa Rica, Guanacaste, Rio Negro, Parque Nacional Rincón de la Vieja, 10.765 ° N, 85.313 ° W, 810 m, 3 March 1986, R. Holzenthal, Fasth, 1 ♂ (pinned) (USNM).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB50D625F4B08B0FA5B.taxon	description	Adult habitus. General color yellow patterned with brown pigment. Head with dark pigment covering ocelli and extending forward and laterad of callosities; occiput dark behind eyes, lappets and an anteromedian triangular area of frons dark brown. Median pronotal band yellow, lateral areas of disc brown with darker rugosities (Fig. 6). Male. Forewing length 20 mm. Hammer a low mound (Fig. 7). Aedeagal apex weakly trilobed; large mesal lobe truncate or notched at tip, lateral lobes small and covered by conspicuous ventral membranous lobes (Figs. 8 - 9); hooks slender, dorsal keel absent (Fig. 10) Female. Unknown. Larva. Unknown.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB50D625F4B08B0FA5B.taxon	etymology	Etymology. The patronym honors Dr. John Morse, a distinguished trichopterist, colleague, friend, and collector of the holotype specimen.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB50D625F4B08B0FA5B.taxon	diagnosis	Diagnosis. The aedeagus of this species is generally similar to that of A. brailovskyi Stark & Kondratieff, 2004 a species known from Mexico (Stark & Kondratieff 2004), however in that species the aedeagal apex is expanded near the tip, the lateral lobes are less distinct and the apical section is not angled ventrad in lateral aspect as in the new species (Fig. 9). The aedeagus of a Costa Rican species, A. exquisita is also similar (Stark 1998) and the new species keys to couplet 11 in Stark (1998) which includes A. exquisita as one of the choices. However, the median lobe of the aedeagal apex is not notched and is more slender than in A. morsei. The new species is also distinct from A. brailovskyi and A. exquisita in head and pronotal pigment patterns (compare Fig. 6 with Fig. 41 in Stark 1998 and Fig. 29 in Stark & Kondratieff 2004).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB40F985F00089CF8B5.taxon	materials_examined	Lectotype ♂ (National Museum, Prague), Sierra du Naya, Mexico, designation Stark, 1998: 582	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB40F985F00089CF8B5.taxon	materials_examined	Material examined. Costa Rica, Heredia, 9 km NE Vara Blanca, 10 ° 14 ’ N, 84 ° 06 ’ W, 1450 - 1550 m, 14 March 2005, S. M. Clark, D. R. Clark, 1 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8239FFB40F985F00089CF8B5.taxon	discussion	Comments. This species is widely distributed from Mexico to Panama (Stark 1998; Stark & Kondratieff 2004) but has not previously been reported from Heredia Province in Costa Rica.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8238FFBB0F9D5E6F0E66FA13.taxon	description	(Figs. 11 - 15)	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8238FFBB0F9D5E6F0E66FA13.taxon	materials_examined	Material examined. Holotype ♂, Costa Rica, Guanacaste Province, Estación Biológica Maritza, Rio Tempisquito, 21 May 1990, light trap (USNM).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8238FFBB0F9D5E6F0E66FA13.taxon	description	Adult habitus. Head yellow except for diffuse brown area forward of ocelli and dark brown lappets and antennae (Fig. 11). Pronotum with pale mesal band; dark lateral bands cover most of disc. Wing membrane transparent with pale amber tint; veins darker except for pale costa. Femora pale except for narrow black apical band; tibiae dark in proximal fourth and with a small distal dark spot. Male. Forewing length 10 mm. Hammer reduced to a small, transparent, hairless disc (Fig. 12). Aedeagal apex pointed in ventral aspect and bearing a prominent pair of ventral, membranous lobes (Fig. 13); apex angled sharply ventrad in lateral aspect (Fig. 14), and bearing an obscure V-shaped dorsal keel composed of divergent, broken lines (Fig. 15). Hooks slender with long acute tips. Female. Unknown. Larva. Unknown.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8238FFBB0F9D5E6F0E66FA13.taxon	etymology	Etymology. The species name, based on Rio Tempisquito, is used as a noun in apposition.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8238FFBB0F9D5E6F0E66FA13.taxon	diagnosis	Diagnosis. This species does not appear to be closely related to others known from Mesoamerica, but is more similar to the Ecuadorian species, A. cajas Zúñiga & Vimos, 2006 (Zúñiga et al. 2006) and the Colombian species A. otun Stark & Zúñiga, 2007 (Zúñiga et al. 2007). The new species is distinguished from both on the basis of pigment pattern, the prominent ventral membranous lobes, the dorsal keel, and the obscure hammer. In Stark (1998) the new species will key to couplet 9 where a choice of A. varilla could be forced, despite conspicuous differences in aedeagal structure.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8237FFBB0D6D5CCE0915FAE9.taxon	description	(Figs. 16 - 20)	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8237FFBB0D6D5CCE0915FAE9.taxon	materials_examined	Material examined. Holotype ♂, Panama, Chiriqui, Totumas Cloud Forest, lodge, 08 ° 53.060 ’ N, 82 ° 41.068 ’ W, 1920 m, 19 - 25 June 2011, E. Fuller (USNM). Paratypes: Costa Rica: Puntarenas, Rio Bellavista, ca. 1.5 km NW Las Alturas, 1400 m, 10 - 11 August 1990, R. Holzenthal, R. Blahnik, F. Munoz, 1 ♂ (BPSC). San Jose, tributary Quebrada Caraiges, 3.6 km SW La Legua, 1650 m, R. Holzenthal, K. Kjer, F. Quesada, 2 ♂ (BPSC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8237FFBB0D6D5CCE0915FAE9.taxon	description	Adult habitus. General color brown patterned with yellow-brown. Head with dark lappets, antennae and a dark band extends from behind ocelli forward to M-line; dark band expanded laterally along M-line, and interrupted by a pale interocellar spot (Fig. 16). Pronotum dark over most of disc, but with a slender pale median band. Wings pale brown, veins brown except for C and apical half of SC. Femora pale in proximal half and dark brown in distal half; fore and mid tibiae brown, hind tibiae with pale transverse median band. Male. Forewing length 20 mm. Hammer a low, mound-like structure (Fig. 17). Aedeagal apex simple (Figs. 18 - 19); ventroapical aspect with a large pair of membranous lobes overlying darkly sclerotized lateral margins (Fig. 18); dorsal keel absent, lateral margins slightly convergent to truncate apex (Fig. 20). Hooks slender. Female. Unknown. Larva. Unknown.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8237FFBB0D6D5CCE0915FAE9.taxon	etymology	Etymology. The species name, based on the type locality at the Mount Totumas Cloud Forest, is used as a noun in apposition.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8237FFBB0D6D5CCE0915FAE9.taxon	diagnosis	Diagnosis. This species keys to couplet 11 in Stark (1998) where A. acutipennis Klapálek, 1923 and A. equisita, a pair of species with trilobed aedeagal apices are the options. Because the new species has a simple aedeagal apex without lateral lobes, neither of these choices are viable. The aedeagal apex of the new species is generally similar to that of A. varilla, a much smaller species (male FWL = 9 - 10 mm) known from Costa Rica and Panama (Stark 1998).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8237FFBB0FE45C5B083DF8D9.taxon	materials_examined	Holotype ♂ (USNM), Rio Peje, 1 km SE San Vicente, Alaquela, Costa Rica	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8237FFBB0FE45C5B083DF8D9.taxon	materials_examined	Material examined. Costa Rica, Heredia, 9 km NE Vara Blanca, 10 ° 14 ’ N, 84 ° 06 ’ W, 1450 - 1550 m, 20 March 2005, S. M. Clark, D. R. Clark, 2 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8237FFBB0FE45C5B083DF8D9.taxon	discussion	Comments. This species was previously known from the holotype and two paratypes, collected in the Costa Rican provinces of Alajuela, Cartago and San Jose (Stark 1998). These are the first specimens from Heredia Province.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8236FFB90CD85FF40E75FF58.taxon	materials_examined	Holotype ♂ (USNM), 4 miles W San Cristobal, Alta Verapaz, Guatemala	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8236FFB90CD85FF40E75FF58.taxon	materials_examined	Material examined. Guatemala: Baja Verapaz, 5 km S Purulha, cloud forest, 15 ° 12.95 ’ N, 90 ° 13.15 ’ W, 1680 m, 14 - 16 October 2011, E. Fuller, 2 ♂ (BYUC).	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
03CD878C8236FFB90CD85FF40E75FF58.taxon	discussion	Comments. This species was previously known from a pair of Guatemalan specimens from Alto Verapaz, and a single male from Honduras.	en	Stark, Bill P. (2014): Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species. Illiesia 10 (2): 6-16, DOI: http://doi.org/10.5281/zenodo.4760479
