identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038B87E9F21EED78FCB8FE4E6D02FA0E.text	038B87E9F21EED78FCB8FE4E6D02FA0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malenka diablo Verdone & Kondratieff 2018	<div><p>Malenka diablo Verdone &amp; Kondratieff, 2018</p> <p>(Figs. 2-13)</p> <p>http://lsid.speciesfile.org/urn:lsid: Plecoptera.speciesfile.org: TaxonName:503118</p> <p>Material examined. Holotype ♂ U.S.A. – California: Santa Clara Co., Colorado Creek, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.51095&amp;materialsCitation.latitude=37.43389" title="Search Plazi for locations around (long -121.51095/lat 37.43389)">Mines Rd.</a>, N 37.43389, W 121.51095, 27 February 2018, B.C. Kondratieff, C.J. Verdone, J.B. Sandberg (NMNH). Paratypes: same data as holotype, 19♂ 8♀, 8L (CSUIC); same data as holotype, 3♂, 3♀, 3L (BYU). Stanislaus Co., Del Puerto Creek, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.41609&amp;materialsCitation.latitude=37.40886" title="Search Plazi for locations around (long -121.41609/lat 37.40886)">Del Puerto Canyon Rd.</a>, N 37.40886, W 121.41609, 27 February 2018, B.C. Kondratieff, C.J. Verdone, J.B. Sandberg, 3♂, 1♀ (CSUIC).</p> <p>Additional material examined. Malenka biloba: California: Kern Co., Dougherty Creek, SR 178, N 35.47293, W 118.71338, 17 April 2018, W.K. Reeves, 2♂, 5♀ (CSUIC). <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.144&amp;materialsCitation.latitude=34.2638" title="Search Plazi for locations around (long -118.144/lat 34.2638)">Los Angeles Co.</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.144&amp;materialsCitation.latitude=34.2638" title="Search Plazi for locations around (long -118.144/lat 34.2638)">Arroyo Seco Creek</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.144&amp;materialsCitation.latitude=34.2638" title="Search Plazi for locations around (long -118.144/lat 34.2638)">Switzer Campground</a>, [N 34.26380, W 118.14400], 12 January 2012, B.C Kondratieff, D. Murányi, J.B. Sandberg, C. R. Nelson, 9♂, 8♀ (CSUIC); Coldbrook Creek, Coldbrook Campground, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.8395&amp;materialsCitation.latitude=34.2939" title="Search Plazi for locations around (long -117.8395/lat 34.2939)">Hwy</a> 39, N 34.29390, W 117.83950, 13 January 2012, B.C. Kondratieff, D. Murányi, J.B. Sandberg, C. R. Nelson, 2♂, 4♀ (CSUIC); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.838&amp;materialsCitation.latitude=34.301" title="Search Plazi for locations around (long -117.838/lat 34.301)">Elizabeth Lake</a> Canyon, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.838&amp;materialsCitation.latitude=34.301" title="Search Plazi for locations around (long -117.838/lat 34.301)">Cottonwood Campground</a>, [N 34.640264, W 118.504806], 18 January 1992, B.C. Kondratieff, 5♂, 2♀ 2L (CSUIC); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.838&amp;materialsCitation.latitude=34.301" title="Search Plazi for locations around (long -117.838/lat 34.301)">Pacoima Canyon Creek</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.838&amp;materialsCitation.latitude=34.301" title="Search Plazi for locations around (long -117.838/lat 34.301)">San Gabriel Mountains</a>, N 34.345958, W 118.36593, 19 January 1992, B.C Kondratieff 2♂ (CSUIC); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.838&amp;materialsCitation.latitude=34.301" title="Search Plazi for locations around (long -117.838/lat 34.301)">Soldier Creek</a>, Falling Springs, 2.3 mi. N Coldbrook Campground, N 34.30100, W 117.83800, 13 January 2012, B.C. Kondratieff, D. Murányi, J.B. Sandberg, C. R. Nelson, 3♂, 1♀ (CSUIC). Orange Co., <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.76941&amp;materialsCitation.latitude=33.71085" title="Search Plazi for locations around (long -116.76941/lat 33.71085)">Silverado Canyon</a>, end of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.76941&amp;materialsCitation.latitude=33.71085" title="Search Plazi for locations around (long -116.76941/lat 33.71085)">Silverado Canyon Rd.</a>, [N 33.75194, W 117.55495], 21 November 2005, L. Myers, 1♂, 2♀ (CSUIC). <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.76941&amp;materialsCitation.latitude=33.71085" title="Search Plazi for locations around (long -116.76941/lat 33.71085)">Riverside Co.</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.76941&amp;materialsCitation.latitude=33.71085" title="Search Plazi for locations around (long -116.76941/lat 33.71085)">Strawberry Creek</a>, Hwy 74, 3 mi. W <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.76941&amp;materialsCitation.latitude=33.71085" title="Search Plazi for locations around (long -116.76941/lat 33.71085)">Mountain Center</a>, N 33.71085, W 116.76941, 10 January 2012, B.C. Kondratieff, D. Murányi, J.B. Sandberg, C. R. Nelson, 1♀ (CSUIC). San Bernardino Co., Lytle Creek, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.496&amp;materialsCitation.latitude=34.26" title="Search Plazi for locations around (long -117.496/lat 34.26)">Applewhite Picnic Area</a>, [N 34.26000, W 117.49600], 9 January 1988, R. W. Baumann, B.J. Sargent, B.C. Kondratieff, C. R. Nelson, 4♂, 2♀ (CSUIC); same location, 8 January 2012, B.C. Kondratieff, D. Murányi, J.B. Sandberg, C. R. Nelson, 33♂, 35♀ (CSUIC). San Diego Co., Fry Creek, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.88&amp;materialsCitation.latitude=33.344" title="Search Plazi for locations around (long -116.88/lat 33.344)">Fry Creek</a> Campground, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.88&amp;materialsCitation.latitude=33.344" title="Search Plazi for locations around (long -116.88/lat 33.344)">Rd.</a> S-6, N 33.34400, W 116.88000, 11 January 2012, B.C. Kondratieff, D. Murányi, J.B. Sandberg, C. R. Nelson, 17♂, 5♀ (CSUIC); Iron Spring Creek, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.87142&amp;materialsCitation.latitude=33.33277" title="Search Plazi for locations around (long -116.87142/lat 33.33277)">Rd.</a> S-6, N 33.33277, W 116.87142, 11 January 2012, B.C. Kondratieff, D. Murányi, J.B. Sandberg, C. R. Nelson, 1♂, 1♀ (CSUIC). Santa Barbara Co., <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.30816&amp;materialsCitation.latitude=34.67949" title="Search Plazi for locations around (long -119.30816/lat 34.67949)">Cachuma Creek</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.30816&amp;materialsCitation.latitude=34.67949" title="Search Plazi for locations around (long -119.30816/lat 34.67949)">Sunset Valley Rd.</a>, [N 34.70248, W 119.91699], 21 January 1992, B.C. Kondratieff, 1♂ (CSUIC). <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.30816&amp;materialsCitation.latitude=34.67949" title="Search Plazi for locations around (long -119.30816/lat 34.67949)">Ventura Co.</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.30816&amp;materialsCitation.latitude=34.67949" title="Search Plazi for locations around (long -119.30816/lat 34.67949)">North Fork Matalija Creek</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.30816&amp;materialsCitation.latitude=34.67949" title="Search Plazi for locations around (long -119.30816/lat 34.67949)">Wheeler Gorge Campground</a>, Los Padres NF, [N 34.51076, W 119.27498], 20 January 1992, B.C. Kondratieff, 1♂, 1♀, 1L (CSUIC); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.30816&amp;materialsCitation.latitude=34.67949" title="Search Plazi for locations around (long -119.30816/lat 34.67949)">Reyes Creek</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.30816&amp;materialsCitation.latitude=34.67949" title="Search Plazi for locations around (long -119.30816/lat 34.67949)">Reyes Creek Rd.</a>, [N 34.67949, W 119.30816], 21 January 1992, B.C. Kondratieff, 4♂, 2♀, 2L (CSUIC).</p> <p>Malenka murvoshi: Nevada: Clark Co., Willow Creek, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.76391&amp;materialsCitation.latitude=36.41774" title="Search Plazi for locations around (long -115.76391/lat 36.41774)">Willow Creek Campground</a> , [N 36.41774, W 115.76391], 3 April 1981, R. W. Baumann, S.M. Clark, 10♂, 2♀ (CSUIC: Paratypes).</p> <p>Distribution. U.S.A. – CA (Fig. 1).</p> <p>Etymology. The new species is named after the region where it was collected, the Diablo Range, mountains that form the eastern flank of the Pacific Coast Ranges southeast of the San Francisco Bay area (Encyclopedia Britannica 2017). The proposed common name is the “Diablo Forestfly”.</p> <p>Male. Macropterous. Length of forewings 7.0– 7.7 mm (n = 10). Length of body 5.0– 5.7 mm (n = 10). General body color brown (Fig. 2). Head mostly brown; area between lateral ocelli and eyes yellow-gold. Antennae uniformly brown (Fig. 2). Occiput with rugose spots. Pronotum with scattered dark rugosities. Legs light brown (Fig. 2). Wings fumose (Fig. 2). Epiproct recurved over abdomen (Figs. 3– 4). Apical half of dorsal sclerite divided longitudinally (Figs. 3–5). Dorsal sclerite tightly appressed along division, except at the apex which opens into a deep V-shaped incision (Figs. 3, 5). Apex of dorsal sclerite expanded into lobes bearing large scales laterally (Fig. 4). Ventral sclerite tightly appressed to the dorsal sclerite (Fig. 4). Apex of ventral sclerite bifurcate, directed dorsally and situated within the apical incision of the dorsal sclerite (Figs. 3, 5). Paraprocts composed of three lobes; outer lobe short, blunt, darkly sclerotized, extending to the mesobasal sclerite encircling the cerci. Inner lobe triangular, apex bifurcate, forming a deep U-shaped notch. Median lobe large, bifurcate, curved distally and upward, flanking the epiproct base (Figs. 6–8). Median lobe of subequal width throughout its length (Figs. 6–7). Proximal margin with a distally directed, sharply pointed apex (Figs. 6–7). Distal margin with a broadly triangular, distally directed, subapical tooth (Figs. 6–8). Cercus very lightly sclerotized, one segmented with a sclerotized nipple-like process on apex and a mesobasal lobe (Figs. 3, 9–10). Mesobasal lobe simple, lightly sclerotized and reduced to a low knob (Figs. 3, 9–10). Vesicle narrow and elongate, widest towards apex, tapering evenly to the base (Fig. 11). Vesicle surface covered with rounded pitlike indentations, basal and lateral margins with trichoid sensilla (Fig. 11). Hypoproct well developed, base covering medial ⅓ of sternum nine, apical portion sharply tapered to a narrow, triangular apex and nested between the inner paraproct lobes (Fig. 11).</p> <p>Female. Macropterous. Length of forewings 8.6– 10.0 mm (n = 10). Length of body 7.0– 8.4 mm (n = 10). General body coloration similar to male. Seventh sternum produced at the posteromedial margin, lightly sclerotized, bearing a nipple-like projection (Fig. 12). Eighth sternum with a narrow, medial V-shaped notch that extends to the base of segment (Fig. 12). Margins of notch lightly sclerotized (Fig. 12).</p> <p>Larva. General morphology as for genus (Fig. 13). Length of body 6.0– 7.3 mm (n = 10). Anterior cervical gill composed of 5–6 branches, posterior gill composed of 7–8 branches (Fig. 13 inset).</p> <p>Diagnosis. Malenka diablo is most similar to M. biloba and M. murvoshi. Males of these species share the following characters: (1) dorsal sclerite divided longitudinally, apex with a deep V-shaped excavation forming apical lobes bearing large scales (Figs. 3, 5, 14, 16; figs. 1–3, 6, 8–10 in Baumann &amp; Kondratieff 2010); (2) apex of ventral sclerite bifurcate, directed dorsally and situated within the apical excavation of the dorsal sclerite (Figs. 5, 16; figs. 3, 9 in Baumann &amp; Kondratieff 2010); (3) mesobasal lobes of cerci simple (Figs. 9–10, 19–20; figs. 1, 6 in Baumann &amp; Kondratieff 2010); and (4) vesicle surface covered with rounded pit-like indentations, basal and lateral margins with trichoid sensilla (Figs. 11, 21; fig. 12 in Baumann &amp; Kondratieff 2010).</p> <p>The new species is distinguished from M. biloba and M. murvoshi by details of the median paraproct lobes and cerci. In the new species, the median lobe of the paraproct is bifurcate apically, bearing well developed points on the proximal and distal margins, with the proximal prong positioned apically and the distal prong subapically (Figs. 6–8). In M. biloba, the median paraproct lobe is also bifurcate, but the proximal prong is finger-like, positioned medially and the distal prong is situated apically (Fig. 17–18). In M. murvoshi, the paraproct has a single point apically and a small spine on the basal proximal margin (figs. 1, 4, 7, 11 in Baumann &amp; Kondratieff 2010). The new species is further differentiated by the shape of the mesobasal lobes on the cerci. In M. diablo, the mesobasal lobe is reduced to a low knob (Fig. 9-10).</p> <p>In contrast, both M. biloba and M. murvoshi possess mesobasal lobes on the cerci that form swollen, nipple-like lobes (Figs. 19–20; figs. 1, 6 in Baumann &amp; Kondratieff 2010).</p> <p>Females of M. diablo (Fig. 12) cannot be separated from M. biloba (Fig. 22) and M. murvoshi (fig. 5, 13 in Baumann &amp; Kondratieff 2010) without associated adult males. Larvae cannot be separated from the other similar species by the number of cervical gills as hypothesized by Baumann and Kondratieff (2010). However, at present, these species appear to be geographically separated.</p> <p>Biological Notes. There is no information about the biology or life cycle of M. diablo. Based on the above records, the emergence period appears to be in late February and early March. The type locality (Fig. 23) and paratype locality (Fig. 24) are 2 nd and 3 rd order, seasonally intermittent streams in the</p> <p>Coastal Mountains level III ecoregion (USEPA 2018), at elevations of 806 m and 450 m, respectively. Other adult stoneflies collected with the type specimens were Taenionema californicum (Needham &amp; Claassen, 1925) and Capnia saratoga Nelson &amp; Baumann, 1987.</p> </div>	http://treatment.plazi.org/id/038B87E9F21EED78FCB8FE4E6D02FA0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Verdone, Chris J.;Kondratieff, Boris C.	Verdone, Chris J., Kondratieff, Boris C. (2018): Malenka Diablo, A New Species Of Stonefly From The Diablo Range Of California, U. S. A. (Plecoptera: Nemouridae). Illiesia 14 (6): 126-134, DOI: http://doi.org/10.5281/zenodo.4758021
