taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764060/files/figure.png	http://doi.org/10.5281/zenodo.4764060	Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA & Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle.	Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA & Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764050/files/figure.png	http://doi.org/10.5281/zenodo.4764050	Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. & Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow & inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle.	Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. & Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow & inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764052/files/figure.png	http://doi.org/10.5281/zenodo.4764052	Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch & thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb & apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle	Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch & thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb & apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764058/files/figure.png	http://doi.org/10.5281/zenodo.4764058	Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle.	Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764064/files/figure.png	http://doi.org/10.5281/zenodo.4764064	Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow & inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle.	Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow & inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764066/files/figure.png	http://doi.org/10.5281/zenodo.4764066	Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch & thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow & insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle.	Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch & thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow & insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764054/files/figure.png	http://doi.org/10.5281/zenodo.4764054	Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO & Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow & inset-lateral view); spinule patch (right white arrow & inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle.	Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO & Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow & inset-lateral view); spinule patch (right white arrow & inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764056/files/figure.png	http://doi.org/10.5281/zenodo.4764056	Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow & inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch & sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle.	Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow & inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch & sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764062/files/figure.png	http://doi.org/10.5281/zenodo.4764062	Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA & Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle.	Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA & Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764597/files/figure.png	http://doi.org/10.5281/zenodo.4764597	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8314B27DF676A7E5FA3FA3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764593/files/figure.png	http://doi.org/10.5281/zenodo.4764593	Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua.	Figs. 19a-s. Isoperla subgenital plates, 50x. Horizontal arrows = subgenital plate width at base, vertical line (two lines when subgenital plate apex invaginated) = subgenital plate length, horizontal line = invagination width at apex. 19a. I.acula; 19b. I. adunca; 19c. I. baumanni; 19d. I. bifurcata; 19e. I. denningi; 19f. I. fulva, rounded; 19g. I. fulva, triangular; 19h. I. laucki; 19i. I. marmorata; 19j. I.miwok; 19k. I. mormona; 19l. I. pinta; 19m. I. quinquepunctata; 19n. I. rainiera; 19o. I. roguensis; 19p. I.sobria; 19q. I. sordida; 19r. I. tilasqua; 19s. I. umpqua.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8310B27FF45AA7E4FEE8A2B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764050/files/figure.png	http://doi.org/10.5281/zenodo.4764050	Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. & Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow & inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle.	Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. & Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow & inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8310B271F450A01EFE59A0C8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764050/files/figure.png	http://doi.org/10.5281/zenodo.4764050	Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. & Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow & inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle.	Figs. 1-2. Adult male characters of Isoperla acula, Deadman Cr., El Dorado Co., CA and I. adunca, Campbell Cr. & Trib., Butte Co., CA, 32-50x. Fig. 1a. I. acula aedeagus, lateral; fine spinule patch (arrow). Fig. 1b. Aedeagus, ventral; fine spinule patches (arrow & inset-posterior view). Fig. 1c. Paraprocts, (black arrow); abdominal spinules (white arrow). Fig. 1d. Vesicle; brush setae (inset). Fig. 2a. I. adunca aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A: posterobasal, B: posteromedian, C: anterior, D: dorsomedian pair. Fig. 2b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 2c. Paraprocts (arrow). Fig. 2d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A831EB271F436A3C3FBEEA323.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764052/files/figure.png	http://doi.org/10.5281/zenodo.4764052	Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch & thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb & apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle	Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch & thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb & apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A831EB271F436A3C3FBEEA323.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764597/files/figure.png	http://doi.org/10.5281/zenodo.4764597	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A831DB272F43DA7E4FBC8A3DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764052/files/figure.png	http://doi.org/10.5281/zenodo.4764052	Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch & thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb & apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle	Figs. 3-4. Adult male characters of Isoperla baumanni, Domingo Springs, Plumas Co., CA and I.bifurcata, Mosquito Spring, Plumas Co., CA, 25-50x. Fig. 3a. I. baumanni aedeagus, lateral; large posterobasal spinule patch & thin apical patch (arrows). Lobes A: posterobasal, B: posteromedian. Fig. 3b. Aedeagus, posterior; large basal spinule patch (arrow).Fig. 3c. Paraproct (white arrow); apical thin spinule patch (black arrow). Fig. 3d. Vesicle. Fig. 4a. I. bifurcata aedeagus, lateral. Lobes A: posterobasal, B: posteromedian, C: dorsal, D: anterior lobe.Sclerotized process characters 1: base, 2: median arm, 3: bifurcate arms, 4: subapical barb & apical tine. Fig. 4b. Aedeagus, ventral. Fig. 4c. Paraproct (arrow). Fig. 4d. Vesicle	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A831BB274F469A7E4FC39A51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764054/files/figure.png	http://doi.org/10.5281/zenodo.4764054	Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO & Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow & inset-lateral view); spinule patch (right white arrow & inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle.	Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO & Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow & inset-lateral view); spinule patch (right white arrow & inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A831BB276F6D3A66AFE68A262.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764054/files/figure.png	http://doi.org/10.5281/zenodo.4764054	Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO & Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow & inset-lateral view); spinule patch (right white arrow & inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle.	Figs. 5-6. Adult male characters of Isoperla denningi, San Mateo Cr., San Diego Co., CA, and I. fulva, Sand Cr., Larimer Co., CO & Pole Cr., Juab Co., UT, 32-50x. Fig. 5a. I. denningi aedeagus, lateral; sclerotized process (arrow & inset-lateral view). Lobes A-B: posterobasal, C: posteroapical, D: dorsolateral pair. Fig. 5b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 5c. Paraprocts (arrow). Fig. 5d. Vesicle. Fig. 6a. I.fulva aedeagus, lateral; sclerotized process (left white arrow & inset-lateral view); spinule patch (right white arrow & inset); paraprocts (inset-black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 6b. Aedeagus, ventral; sclerotized process (arrow & inset-posterior view). Fig. 6c. Aedeagus, dorsal; sclerotized process (black arrow); spinule patch (white arrow). Fig. 6d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A831BB276F6D3A66AFE68A262.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764597/files/figure.png	http://doi.org/10.5281/zenodo.4764597	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8319B276F41AA175FC22A322.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764056/files/figure.png	http://doi.org/10.5281/zenodo.4764056	Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow & inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch & sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle.	Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow & inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch & sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8318B277F5D3A600FB27A79D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764056/files/figure.png	http://doi.org/10.5281/zenodo.4764056	Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow & inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch & sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle.	Figs. 7-8. Adult male characters of Isoperla laucki, Dragsaw Spring, Humboldt Co., CA, and I. marmorata, Big Chico Cr., Tehama Co., CA, 32-40x. Fig. 7a. I. laucki aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); posteroapical and posterobasal spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterolateral pair. Fig.7b. Aedeagus, ventral; sclerotized process (white arrow & inset-dorsal view); posterobasal spinule patch (black arrow). Fig. 7c. Paraprocts (left white arrow); sclerotized process (right white arrow); posteroapical spinule patch (black arrow). Fig. 7d. Vesicle. Fig. 8a. I. marmorata aedeagus, lateral; sclerotized process (white arrow & inset-lateral view); paraprocts (black arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 8b. Aedeagus, posterior; spinule patch & sclerotized process (black arrow). Fig. 8c. Paraprocts (arrow). Fig. 8d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8318B277F5D3A600FB27A79D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764597/files/figure.png	http://doi.org/10.5281/zenodo.4764597	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8318B269F6AFA4F6FE79A76C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764058/files/figure.png	http://doi.org/10.5281/zenodo.4764058	Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle.	Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8318B269F6AFA4F6FE79A76C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764597/files/figure.png	http://doi.org/10.5281/zenodo.4764597	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8306B269F462A464FBC8A648.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764058/files/figure.png	http://doi.org/10.5281/zenodo.4764058	Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle.	Figs. 9-10. Adult male characters of Isoperla miwok, Campbell Cr., Butte Co., CA and I. mormona, Big Chico Cr., Butte Co., CA, 40-50x. Fig. 9a. I. miwok aedeagus, lateral; posteroventral pair and posteroapical spinule patches (arrows). Lobes A: posterior, B: dorsal. Fig. 9b. Aedeagus, ventral; posteroventral pair of spinule patches (black arrow), small pointed apicolateral lobe (white arrow). Fig. 9c. Paraprocts (white arrow); posteroapical spinule patch (black arrow). Fig. 9d. Vesicle. Fig. 10a. I. mormona aedeagus, lateral; paraproct (black arrow); fine dorsoapical spinulae (white arrow). Lobe Adorsoanterior. Fig. 10b. Aedeagus, ventral; paraproct base (arrow). Fig. 10c. Paraprocts (black arrow); fine dorsoapical spinulae (left white arrow); abdominal spinulae (right white arrow). Fig. 10d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8306B26BF6C3A540FDCCA013.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764060/files/figure.png	http://doi.org/10.5281/zenodo.4764060	Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA & Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle.	Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA & Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8304B26BF416A384FB1FA0E5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764060/files/figure.png	http://doi.org/10.5281/zenodo.4764060	Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA & Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle.	Figs. 11-12. Adult male characters of Isoperla pinta, Butte Cr., Butte Co., CA and I. quinquepunctata, Yuba R., Yuba Co., CA & Nehalem R., Clatsop Co., OR, 32-50x. Fig. 11a. I. pinta aedeagus, lateral. Lobes A: posterobasal, B: posteroapical lobe, C: anterolateral pair. Fig. 11b. Aedeagus, ventral. Fig. 11c. Paraprocts (arrow). Fig.11d. Vesicle; prothoracic pigment pattern (inset-dorsal view). Fig. 12a. I. quinquepunctata aedeagus, lateral. Lobes A: posterobasal, B: posteroapical, C: dorsal, D: anterior. Fig. 12b. Aedeagus, ventral. Fig. 12c. Paraprocts (black arrow); abdominal spinules (white arrows). Fig. 12d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8304B26DF6D3A3E3FD9CA6B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764062/files/figure.png	http://doi.org/10.5281/zenodo.4764062	Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA & Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle.	Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA & Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8302B26DF433A41FFC22A725.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764062/files/figure.png	http://doi.org/10.5281/zenodo.4764062	Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA & Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle.	Figs. 13-14. Adult male characters of Isoperla rainiera, Tyee Spring, Skamania Co., WA and I. roguensis, Butte Cr., Butte Co., CA & Mad R., Humboldt Co., CA, 32-50x. Fig. 13a. I. rainiera aedeagus, lateral; sclerotized process (arrow). Lobes A: posterior, B: dorsal, C: anterior. Fig. 13b. Aedeagus, ventral; sclerotized process (arrow).Fig. 13c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 13d. Vesicle. Fig. 14a. I. roguensis aedeagus, lateral; sclerotized process (white arrow); spinule patches (black arrows). Lobes A: posterior, B: dorsal, C: anterior. Fig. 14b. Aedeagus, ventral (Mad R.); sclerotized process (arrow); inset-posterior view (Butte Cr). Fig. 14c. Paraprocts (left arrow); sclerotized process (right arrow). Fig. 14d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8302B26DF433A41FFC22A725.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764597/files/figure.png	http://doi.org/10.5281/zenodo.4764597	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8302B26FF6D0A4A3FE8FA601.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764064/files/figure.png	http://doi.org/10.5281/zenodo.4764064	Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow & inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle.	Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow & inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8300B26FF456A58FFD94A2F9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764064/files/figure.png	http://doi.org/10.5281/zenodo.4764064	Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow & inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle.	Figs. 15-16. Adult male characters of Isoperla sobria, Tyee Spring, Skamania Co., WA and I. sordida, Cold Water Cr., Mono Co., CA, 25-40x. Fig. 15a. I. sobria aedeagus, lateral (Tyee Sp., in copula); spinule patch (arrow). Lobes A: anterior, B: dorsal pair, C: posteroapical (aberrant). Fig. 15b. Aedeagus, posterior; spinule patch (arrow). Fig. 15c. Paraprocts (black arrow), Lk. Davis-Corral Sp.; spinulae patch, lateral (white arrow); dorsal pair of lobes in typical position. Fig. 15d. Vesicle. Fig. 16a. I. sordida aedeagus, lateral; sclerotized process (white arrow); paraproct (black arrow). Lobes A: posteroventral, B: posteromedian, C: dorsal, D: anterior. Fig. 16b. Aedeagus ventral; sclerotized process (arrow & inset-posterior view).Fig. 16c. Paraprocts (black arrow); sclerotized process (white arrow). Fig. 16d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8300B26FF438A1D7FC0AA188.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764066/files/figure.png	http://doi.org/10.5281/zenodo.4764066	Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch & thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow & insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle.	Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch & thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow & insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8300B26FF438A1D7FC0AA188.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764597/files/figure.png	http://doi.org/10.5281/zenodo.4764597	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	Figs. 20a-f. The cleared aedeagus of six western Nearctic Isoperla species after live evertion; membranous opening of internal tube like process or apex (white arrows); tube like internal process (black arrows). 50x (20b, d & e, cropped and enlarged). 20a. I. baumanni, lateral view, partially everted; 20b. I. fulva, lateral view; 20c. I. marmorata, lateral view; 20d. I. miwok, dorsoapical view, tube like process openings along anteroapical margin; 20e. I. roguensis, ventral view, tube like process openings along anteroapical margin; 20f. I. tilasqua, lateral view.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
1B32387A8300B261F6A0A107FEBFA02C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4764066/files/figure.png	http://doi.org/10.5281/zenodo.4764066	Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch & thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow & insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle.	Figs. 17-18. Adult male characters of Isoperla tilasqua, Meacham Cr., Umatilla Co., OR and I. umpqua, Muir Cr., Douglas Co., OR, 25-40x. Fig. 17a. I. tilasqua aedeagus, lateral; large basal spinule patch & thin apical patch (arrows). Lobes A-B: posterior. Fig. 17b. Aedeagus, posterior; large basal spinule patch (arrow). Fig. 17c. Paraprocts (white arrow); apical thin spinule patch (right arrow); large basal patch (left arrow). Fig. 17d. Vesicle. Fig. 18a. I. umpqua aedeagus, lateral; sclerotized process (arrow). Lobe A: dorsoanterior. Fig. 18b. Aedeagus, ventral; sclerotized process (arrow & insetposterior view). Fig. 18c. Paraproct (left white arrow); sclerotized process (right white arrow); abdominal tergite 9 spinulae (black arrow). Fig. 18d. Vesicle.	2013-12-31	Sandberg, John B.;Kondratieff, Boris C.		Zenodo	biologists	Sandberg, John B.;Kondratieff, Boris C.			
