taxonID	type	description	language	source
A61087FCFFEEF5364FD9F951FB8A6439.taxon	materials_examined	Holotype ♂ (temporary repository American Museum of Natural History), Park Nacional Amborό, Caballero Province, Santa Cruz Dept., Bolivia Material examined. BOLIVIA: Santa Cruz Department, La Siberia, 17.820 ° S, 64.678 ° W, 2550 m, 20 April 2005, S. M. Clark, R. L. Johnson, 1 ♂ (BYUC). Comments. This species is known from more than 120 males all from Santa Cruz Department, Bolivia. The female and larva remain unknown (Stark 2004).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEEF5364FD9F951FB8A6439.taxon	discussion	Comments. This species is known from more than 120 males all from Santa Cruz Department, Bolivia. The female and larva remain unknown (Stark 2004).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEEF5364FD9F951FB8A6439.taxon	distribution	Anacroneuria amaru Stark, 2004 Santa Cruz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364D5BFE16FC5E64AF.taxon	description	(Figs. 1 - 5)	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364D5BFE16FC5E64AF.taxon	materials_examined	Material examined. Holotype ♂, BOLIVIA, La Paz Department, Province Nor Yungus, Huarinilla, Rio Selva Resort, 16.20198 ° S, 67.79363 ° W, 3821 ’, 28 April 2006, R. L. Johnson, V. J. Anderson, S. M. Clark (USNM).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364D5BFE16FC5E64AF.taxon	description	Adult habitus. General color brown, patterned with pale brown or yellow. Head mostly yellow but with a pair of dark brown curved bars extending from between ocelli and curved forward and laterad of ocelli; a pair of pale brown diagonal bars located on occiput behind ocelli and dark brown lappets and a pair of small dark triangles occur near anterior of frons (Fig. 1). Pronotum mostly brown but with pale area located near anterolateral margin. Wing membrane and veins brown except for pale costa and pale spot near cord. Fore and mid femora with dark brown dorsoapical band which is much reduced on hind femora; fore and mid tibiae dark brown along outer margin but pale along inner margin. Antennae dark brown. Male. Forewing length 9 mm. Hammer thimble shaped (Fig. 2). Aedeagal apex trilobed in ventral and dorsal aspect (Figs. 3, 5); pale mesal lobe projecting beyond lateral lobes and dark along lateral margins in basal half; lateral lobes strongly sclerotized and divergent from mesal lobe giving an ear-like appearance in lateral aspect (Fig. 4). Dorsal edge of aedeagal apex bearing a strong keel with short basal arms (Fig. 5). Hooks slender. Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364D5BFE16FC5E64AF.taxon	etymology	Etymology. The patronym honors Dr. Shawn Clark, Coleopterist of Brigham Young University. In addition to his expertise in the Coleoptera family Chrysomelidae, Shawn is well known to students of Plecoptera as an outstanding stonefly collector.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364D5BFE16FC5E64AF.taxon	diagnosis	Diagnosis. Anacroneuria clarki is part of a small Andean complex of species which includes A. cayapa Stark (Ecuador), A. marta Zúñiga & Stark (Colombia), A. tejon Baena & Stark (Colombia), A. x-nigrum Klap á lek (Peru) and A. zwicki Stark & Sivec (Bolivia, Peru) (Stark 2001; Stark & Sivec 1998; Stark et al. 1999; Zúñiga & Stark 2002). The aedeagus of the new species is similar to A. cayapa and A. x-nigrum in having the apex tri-lobed with a slender, mostly membranous mesoapical lobe projecting beyond the lateral lobes, and it further resembles A. x-nigrum in having a well developed dorsal keel along the entire length of the mesal lobe (Stark 2001; Stark & Sivec 1998). The two may be sister species within this complex and they are distinguished by the dark interocellar pigment, said to form an X-pattern in A. x-nigrum (Klap á lek 1921; Stark & Sivec 1998), and also by details of the aedeagal apex. In A. x-nigrum the mesoapical lobe is gradually narrowed from base to apex and does not have dark sclerotized edges whereas in A. clarki the mesoapical lobe is constricted near midlength and has dark lateral margins in the basal half.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364D5BFE16FC5E64AF.taxon	distribution	Anacroneuria clarki sp. n. La Paz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364FEBFC26FA8A6714.taxon	materials_examined	Holotype ♂ (temporary repository American Museum of Natural History), Park Nacional Amborό, Caballero Province, Santa Cruz Dept., Bolivia Material examined. BOLIVIA, Cochabamba Department, Province Chapare, Incachaca, 17 ° 14 ’ S, 65 ° 49 ’ W, 7340 ’, 8 December 2008, S. M. Clark, 1 ♂ (BYUC). La Paz Department, Province Nor Yungas, Rio Huarinilla, Huarinilla, 16 ° 12 ’ S, 67 ° 47 ’ W, 3750 ’, 11 November 2009, S. M. Clark, H. R. Hinkson, 1 ♂ (BYUC). Comments. All known previous records of this species are from, or near the type locality in Santa Cruz Department. (Stark 2004).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364FEBFC26FA8A6714.taxon	discussion	Comments. All known previous records of this species are from, or near the type locality in Santa Cruz Department. (Stark 2004).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364FEBFC26FA8A6714.taxon	distribution	Anacroneuria cusi Stark, 2004 Cochabamba, Santa Cruz, La Paz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364F95F995FBF867DD.taxon	materials_examined	Lectotype ♂ (Hofmuseum), Coroico, Bolivia, designated by Zwick (1973)	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364F95F995FBF867DD.taxon	materials_examined	Material examined. BOLIVIA: Cochabamba Department, Province Chapare: Rio Vinchuta, Shineota, 17.008 ° S, 65.309 ° W, 255 m, 22 April 2005, S. M. Clark, R. L. Johnson, 1 ♂ (BYUC). Rio Espiritu Santo, Villa Tunare, 22 April 2005, S. M. Clark, R. L. Johnson, 1 ♂ (BYUC). La Paz Department, Province Nor Yungus: Mun. Coroico, Vagante, 2 December 2004, R. L. Johnson, V. J. Anderson, 1 ♂ (BYUC). Vagante, nr Coroico, 16.17854 ° S, 67.69071 ° W, 1110 m, 30 April 2006, R. L. Johnson, V. J. Anderson, S. M. Clark, 1 ♂ (BYUC). Rio Vagante, Vagante, 16 ° 11 ’ S, 67 ° 41 ’ W, 3650 ’, 12 November 2009, S. M. Clark, H. R. Hinkson, 1 ♂, 2 ♀ (BYUC). Rio Huarinilla, Huarinilla, 16 ° 12 ’ S, 67 ° 48 ’ W, 3750 ’, 11 November 2009, S. M. Clark, H. R. Hinkson, 2 ♂ (BYUC).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364F95F995FBF867DD.taxon	discussion	Comments. Zwick (1973) selected the male lectotype from a surviving olivian syntype from Klap á lek’s (1922) type series. Stark & Sivec (1998) noted this species was quite common at Woytkowski’s “ Hacienda Maria ” site in Peru where it represented about half the catch of stonefly specimens. They also reported the species from two sites in Cochabamba which are the only previous records from Bolivia.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEFF5364F95F995FBF867DD.taxon	distribution	Anacroneuria handlirschi Klap á lek, 1922 Cochabamba, La Paz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364D13FD43FBA4679A.taxon	materials_examined	Holotype ♂ (United States National Museum), Manu, Pakitza, Madre de Dios, Peru Material examined. BOLIVIA: Cochabamba Department, Province Chapare, Rio Coni, Shineota, 17.012 ° S, 65.219 ° W, 240 m, 22 April 2005, S. M. Clark, R. L. Johnson, 1 ♂ (BYUC). Comments. This species was described from a single Peruvian male specimen (Stark & Sivec 1998) and has not been reported since the original description.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364D13FD43FBA4679A.taxon	discussion	Comments. This species was described from a single Peruvian male specimen (Stark & Sivec 1998) and has not been reported since the original description.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364D13FD43FBA4679A.taxon	distribution	Anacroneuria lupaca Stark & Sivec, 1998 Cochabamba	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364D0DFBBDFBF86643.taxon	materials_examined	Holotype ♂ (United States National Museum), pte. San Pedro, Paucartambo, Cuzco, Peru Material examined. BOLIVIA: La Paz Department, Nor Yungus Province: Mun. Coroico, Vagante, 21 December 2004, R. L. Johnson, V. J. Anderson, 1 ♂ (BYUC). Vagante, nr. Coroico, 16.17854 ° S, 67.69071 ° W, 1110 m, 30 April 2006, R. L. Johnson, V. J. Anderson, S. M. Clark, 1 ♂ (BYUC). Huarinilla, Rio Selva Resort, 16 ° 202 ’ S, 67 ° 494 ’ W, 29 October 2007, R. L. Johnson, V. J. Anderson, 1 ♂ (BYUC). Rio Huarinilla, Huarinilla, 16 ° 12 ’ S, 67 ° 48 ’ W, 3830 ’, 10 December 2008, R. L. Mower, 1 ♂ (BYUC). Same site but 3750 ’, 11 November 2009, S. M. Clark, H. R. Hinkson, 7 ♂ (BYUC). Comments. Bolivian and Peruvian records are available in Stark & Sivec (1998). The Bolivian records are from two sites including one from “ Yungas Las Paz ”.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364D0DFBBDFBF86643.taxon	discussion	Comments. Bolivian and Peruvian records are available in Stark & Sivec (1998). The Bolivian records are from two sites including one from “ Yungas Las Paz ”.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364D0DFBBDFBF86643.taxon	distribution	Anacroneuria pachacuti Stark & Sivec, 1998 Cochabamba, La Paz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364FC2FF29FB8A660B.taxon	materials_examined	Holotype ♂ (American Museum of Natural History), Park Nacional Amborό, Caballero Prov., Santa Cruz Dept., olivia Material examined. BOLIVIA: Santa Cruz Department, La Siberia, 17.820 ° S, 64.678 ° W, 2550 m, 20 April 2005, S. M. Clark, R. L. Johnson, 1 ♂ (BYUC). Comments. This species was previously known from five specimens all collected in Santa Cruz Department, Bolivia.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364FC2FF29FB8A660B.taxon	discussion	Comments. This species was previously known from five specimens all collected in Santa Cruz Department, Bolivia.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5364FC2FF29FB8A660B.taxon	distribution	Anacroneuria spectori Stark, 2004 Santa Cruz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5384FC2FD68FC5E606A.taxon	description	(Figs. 6 - 10)	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5384FC2FD68FC5E606A.taxon	materials_examined	Material examined. Holotype ♂, BOLIVIA: La Paz Department, Province Nor Yungas, Rio Vagante, Vagante, 16 ° 11 ’ S, 67 ° 41 ’ W, 3650 ’, 12 November 2009, S. M. Clark, H. R. Hinkson (USNM).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5384FC2FD68FC5E606A.taxon	description	Adult habitus. General color brown, patterned with areas of dark brown and pale brown or yellow. Head with a large area of dark pigment covering ocellar region and extending forward and laterad over much of frons (Fig. 6); lappets dark brown and occiput with area of dark pigment outlining a pair of triangles with pale centers; antennae and palpi brown. Mid and fore femora banded with dark pigment in apical third, tibiae uniformly dark brown (hind legs missing). Wings membranes brown, veins dark brown. Male. Forewing length 8.5 mm. Hammer thimble shaped (Fig. 7). Aedeagal apex simple, scoop shaped (Figs. 8 - 10), but with heavily sclerotized lateral margins, a prominent dorsal keel which forms an Xpattern (Fig. 10), and a pair of prominent membranous ventral lobes; lateral margins in dorsal aspect somewhat scalloped. Hooks slightly wider than in most species. Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5384FC2FD68FC5E606A.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the type locality of Rio Vagante.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5384FC2FD68FC5E606A.taxon	diagnosis	Diagnosis. The aedeagus of A. vagante is most similar to that of A. karina Stark, known from Venezuela (Stark 1999), however that species lacks the distinctive head and pronotal pattern of A. vagante; in addition the bases of the aedeagal hooks in A. karina are more swollen, the dorsal keel is shorter, and the ventral membranous lobes smaller in that species than in A. vagante. The aedeagus of the new species is also generally similar to that of A. manauensis Ribeiro-Ferreira, described from Brazil (Ribeiro-Ferreira & Froehlich 2001), A. puna Stark, described from Ecuador (Stark 2001), and A. taylori Stark, described from Bolivia (Stark 2004). The new species differs from the latter two in having a much broader area of dark pigment on the frons, but also differs from the former two species in having a prominent pair of membranous lobes on the aedeagal apex. It differs from the latter species in lacking subchelate aedeagal hooks and in having larger membranous lobes on the aedeagal apex.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEDF5384FC2FD68FC5E606A.taxon	distribution	Anacroneuria vagante sp. n. La Paz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344FF5FD67FA2466D4.taxon	description	(Figs. 16 - 18)	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344FF5FD67FA2466D4.taxon	materials_examined	Material examined. PARAGUAY: Hernandarias, near Acaray River, 3 November 1956, C. J. D. Brown, 7 ♀ (BYUC).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344FF5FD67FA2466D4.taxon	description	Adult habitus. General color pale brown, patterned with darker brown pigment. Head with a dark V connecting ocelli and extending toward bases of antennae (Fig. 16); lappets and antennae dark brown, palpi pale. Pronotum pale brown with darker, slender rugosities scattered over disk; median suture with a narrow dark band on either side. Hind femora darker in apical third, hind tibia uniformly brown. Wings tinted pale brown, veins darker. Male. Unknown. Female. Forewing length 10 - 11 mm. Subgenital plate weakly four lobed; inner lobes separated by a shallow V-shaped notch (Fig. 18) and outer lobes by a very shallow, wide notch. Sternum 9 posterior sclerite long, slender and sparsely setose; median sclerite covered basally and through median zone with fine, short setae, and bearing a few scattered longer, thicker setae near sclerite and laterally. Egg. Spindle shaped, long, slender and darker on anterior pole (Fig. 17). Larva. Unknown.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344FF5FD67FA2466D4.taxon	discussion	Comments. We are unable to associate these females with any of the species known from the region. Certainly they are distinct from A. debilis and A. trimacula (Froehlich 2002) and they are also distinct in egg morphology and subgenital plate shape from A. fuscicosta (Enderlein) and A. stanjewetti Froehlich, two regional species which have similar head patterns (Zwick 1973; Froehlich 2002).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344D44FDAFFADE63D5.taxon	description	(Figs. 11 - 15)	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344D44FDAFFADE63D5.taxon	materials_examined	Material examined. Holotype ♂, PARAGUAY: Amambay Department, Rio Ypane, cerca Pedro Juan Caballero, 25 November 1956, C. J. D. Brown (USNM).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344D44FDAFFADE63D5.taxon	description	Adult habitus. General color brown, patterned with areas of pale pigment. Head with a large area of dark pigment extending from ocellar region over central frons (Fig. 11); dark pigment area expanded laterally in central frons where a small oval pale area is located; lappets and antennae dark brown. Pronotum mostly dark but with a narrow pale median stripe and pale lateral areas. Hind femora dark brown in apical third but pale basally; hind tibia with small dark bands basally and apically but pale for most of length. Wing membrane brown, veins (particularly R vein) dark brown. Male. Forewing length 10 mm. Hammer thimble shaped, apical diameter less than height (Fig. 12). Aedeagal apex a short, simple, truncate scoop bearing a ventral pair of prominent membranous lobes (Fig. 13); dorsal margin of apical area expanded into a pair of small rounded lateral lobes near point where hooks cross (Fig. 15); dorsal keel lines absent, apex bent strongly dorsad, giving a foot-like shape in lateral aspect (Fig. 14). Hooks slender. Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344D44FDAFFADE63D5.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the type locality of Rio Ypane.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344D44FDAFFADE63D5.taxon	diagnosis	Diagnosis. This species is related to the Anacroneuria atrifrons Klap á lek group of species (Froehlich 2008) which includes at least A. azul Rojas & Baena (Stark et al. 1999), A. canchi Stark & Sivec (Stark & Sivec 1998) and A. pastaza Stark (Stark 2001). Of these species, the aedeagus of A. ypane is most similar to that of the Colombian species, A. azul, but the membranous lobes in that species are smaller and a small transverse dorsal keel is present. The latter species also differs from A. ypane in having a much smaller area of dark head pigment.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFEBF5344D44FDAFFADE63D5.taxon	discussion	Comments. Froehlich (2002) reported A. debilis (Pictet) from a single male collected in Parque Nacional Ybycui and A. trimacula Jewett from a single female collected at the same locality. Thus, Anacroneuria ypane becomes only the third member of the genus to be confirmed for Paraguay, and the first to be described from a Paraguayan specimen.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE8F5364D73F97DFEC5622F.taxon	description	(Figs. 19 - 21)	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE8F5364D73F97DFEC5622F.taxon	materials_examined	Material examined. PARAGUAY: Amambay Department, Rio Ypane, cerca Pedro Juan Caballero, 25 November 1956, C. J. D. Brown, 1 ♀ (BYUC).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE8F5364D73F97DFEC5622F.taxon	description	Adult habitus. General color pale brown, patterned with brown pigment. Head with a brown V-shaped pattern over ocelli and curving outwards toward anterior margins of eyes (Fig. 19). Lappets and antennae pale brown, palpi paler. Hind femora entirely pale, tibiae pale except for small brown areas on distal and proximal ends. Wings transparent, veins pale brown. Male. Unknown. Female. Forewing length 15.5 mm. Subgenital plate four lobed; inner lobes separated by a wide, shallow notch (Fig. 21), lateral lobes separated by shallow notches. Sternum 9 with obscure posterior sclerite; median sclerite covered laterally and along posterior margin with long setae, basal, median area covered with fine, short setae. Egg. Spindle shaped, long, slender and bearing a small, membranous cap (Fig. 20). Larva. Unknown.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE8F5364D73F97DFEC5622F.taxon	discussion	Comments. This female was collected with the holotype of A. ypane and appears distinct from it on the basis of color pattern, and from others known from Paraguay (Froehlich 2002). We are also not able to match it with others known from the region.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D73FC61FC5B651A.taxon	description	(Figs. 22 - 24)	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D73FC61FC5B651A.taxon	materials_examined	Material examined. PARAGUAY: cerca Hernandarias, Boca de Rio Acaray, 4 November 1956, C. J. D. Brown, 2 ♀ (BYUC).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D73FC61FC5B651A.taxon	description	Adult habitus. General color brown, patterned with pale areas. Head with a dark quadrate area covering ocelli and extending to M-line; diffuse brown pigment occurs forward of M-line and extending anterolaterally from dark pigment (Fig. 22); lappets and antennae brown, palpi pale brown. Pronotum pale brown with darker rugosities and a pale median band. Hind femora and tibia brown. Wings transparent, veins brown. Male. Unknown. Female. Forewing length 16 mm. Subgenital plate four lobed with lobes approximately equal in size (Fig. 24). Sternum 9 with a wide, well developed posterior sclerite which bears prominent setae over most of length but bare mesally; mesal sclerite bearing long setae posterolaterally and fine setae basally and mesally. Egg. Spindle shaped moderate in length and broadly rounded at anterior pole. Collar bears a small membranous cap (Fig. 23).	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D73FC61FC5B651A.taxon	discussion	Comments. The subgenital plate of this species is generally similar to that of A. debilis (Froehlich 2002) but the egg and color pattern are distinct.	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71FAB0FB8A64C5.taxon	distribution	Santa Cruz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71FA53FB8A64E6.taxon	distribution	Santa Cruz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71FA77FBA46482.taxon	distribution	Cochabamba	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71F9CEFB8A674B.taxon	distribution	Santa Cruz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71F985FBEF6730.taxon	distribution	La Paz, Santa Cruz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71F95CFB1667F9.taxon	distribution	Department unknown	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71F913FC5E67A7.taxon	distribution	La Paz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71F8EAFB16666F.taxon	distribution	Department unknown	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE9F5364D71F8A1FB8A66D4.taxon	distribution	Santa Cruz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE7F5384D71FF29FC5E61AD.taxon	distribution	La Paz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE7F5384D71FECDFBA46049.taxon	distribution	Cochabamba	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE7F5384D71FE84FB8A6036.taxon	distribution	Santa Cruz	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE7F5384D71FE84FB8A6036.taxon	distribution	Cochabamba	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
A61087FCFFE7F5384D71FEA7FBA460D2.taxon	distribution	Cochabamba	en	Stark, Bill P., Baumann, Richard W. (2011): Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species. Illiesia 7 (19): 182-191, DOI: http://doi.org/10.5281/zenodo.4760267
