taxonID	type	description	language	source
038A8786765F0E40516FF9CAFDE4FDA9.taxon	description	(Figs. 1 - 4)	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765F0E40516FF9CAFDE4FDA9.taxon	materials_examined	Holotype ♀ (Museum of Comparative Zoology), Claremont [Los Angeles Co.], California	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765F0E40516FF9CAFDE4FDA9.taxon	materials_examined	Material examined. California: Butte Co., Heinz Creek, Hwy 70, 20 May 1982, B. Stark, D. Ziegler, 1 ♀ (BPS). Sierra Co., Little Truckee River, Little Truckee Campground, 4 July 1979, B. Stark, K. W. Stewart, 18 ♂, 3 ♀ (BPS). Montana: Lincoln Co., Yaak River, Yaak River Campground, 19 July 1979, B. Stark, K. W. Stewart, R. W. Baumann, 2 ♀ (BPS). Oregon: Union Co., Grande Ronde River, Hwy 244, 29 May 2000, B. Stark, I. Sivec, M. Zúñiga, 3 ♂, 4 ♀ (BPS).	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765F0E40516FF9CAFDE4FDA9.taxon	description	Egg. Outline oval with short stalked collar, ends broadly rounded (Fig. 1). Length including collar ca. 430 - 460 μm, equatorial width ca. 360 - 390 μm, collar length ca. 35 - 40 μm, collar width at midlength ca. 60 - 68 μm. Egg body at base of collar encircled by slightly raised ridge. Sides of collar with irregular, prominent struts (Fig. 2); rim flanged and irregularly incised (Figs. 1 - 2). Anchor biscuit-shaped, surface with scattered mushroom bodies (Fig. 3). Chorionic surface relatively smooth but with obscure hexagonal follicle cell impressions visible at 500 X (Figs. 2, 4). Micropylar row subequatorial.	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765C0E4051D6FDF4FC76FCE9.taxon	description	(Figs. 5 - 8)	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765C0E4051D6FDF4FC76FCE9.taxon	materials_examined	Material examined. California: Marin Co., Muir Woods, Redwood Creek, 21 June 1985, B. Stark, 1 ♀ (BPS). Same location, 18 June 1966, R. W. Baumann, 1 ♂, 1 ♀ (BYU).	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765C0E4051D6FDF4FC76FCE9.taxon	description	Egg. Outline oval, collar absent (Fig. 5). Length ca. 440 μm, equatorial width ca. 372 μm. Chorionic surface without punctations or obvious follicle cell impressions (Figs. 6, 8). Micropylar row subequatorial; micropylar orifices with raised rims (Fig. 7).	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765C0E4051D6FDF4FC76FCE9.taxon	discussion	Comments. Eggs of C. californica (Figs. 1 - 4) and the three species of Japanese Calineuria (Isobe 1988; 1997) have similar short stalked collars encircled by a low to well developed basal ridge. The chorionic surface is punctate in the Japanese species but smooth or with obscure follicle cell impressions in C. californica and C. sp. A (Figs. 2, 6). The male and female of C. sp. A appear indistinguishable from C. californica in color pattern, subgenital plate shape and aedeagal armature, however, the egg has no collar. These specimens may represent an undescribed species, but a larger sample and a comprehensive study of other Calineuria specimens is needed in order to evaluate its status. Drumming and molecular data may also be desirable.	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765D0E41515CFF7BFC09FD32.taxon	description	(Figs. 9 - 12)	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765D0E41515CFF7BFC09FD32.taxon	materials_examined	Holotype ♂ (United States National Museum), Burney Falls State Park, Shasta Co., California Material examined. California: Siskiyou Co., Big Springs, Mt. Shasta City Park, 7 July 1979, B. Stark, K. W. Stewart, 1 ♀ (BPS). Oregon: Benton Co., Kiser Creek, Hwy 39, 1 June 2000, B. Stark, I. Sivec, M. Zúñiga, 3 ♂, 1 ♀ (BPS). Multnomah Co., Wahkeena Falls, 25 June 1985, B. Stark, 24 ♂, 4 ♀ (BPS). Washington: Pierce Co., Carbon River, 16 August 1999, B. C. Kondratieff, 1 ♂, 3 ♀ (BPS).	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765D0E41515CFF7BFC09FD32.taxon	description	Egg. Outline oval with short, narrow collar (Fig. 9). Egg length ca. 500 μm, equatorial width ca. 400 μm, collar length ca. 20 μm, width ca. 50 μm. Chorionic surface coarsely punctate with shallow pits over middle half of egg (Figs 10, 12.); punctate zone ca. 260 μm wide; area around collar and a narrow opercular ring around lid lack pits; some pits on lid form obscure follicle cell impression walls. Micropylar row located on lid (Fig. 11); orifices raised, canals not clearly visible on surface.	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765A0E465084FF7BFBFDFC8C.taxon	description	(Figs. 13 - 16)	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765A0E465084FF7BFBFDFC8C.taxon	materials_examined	Holotype ♀ (Cornell University), Yellowstone National Park, Wyoming Material examined. Montana: Gallatin Co., Hyalite Creek, above Window Rock, 20 August 2004, B. Stark, I. Sivec, 1 ♀ (BPS). Wyoming: Johnson Co., Middle Fork Clear Creek, 17 August 2004, B. Stark, I. Sivec, 2 ♂, 2 ♀ (BPS). Park Co., Lake Creek, Hwy 296, 18 August 2004, B. Stark, I. Sivec, 3 ♂, 3 ♀ (BPS).	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765A0E465084FF7BFBFDFC8C.taxon	description	Egg. Outline oval with short, narrow collar (Fig. 13). Egg length ca. 530 μm, equatorial width ca. 400 μm, collar length ca. 30 μm, width ca. 60 μm. Chorionic surface coarsely punctate in a band ca. 325 μm wide, extending from near collar (Figs. 14, 16) to smooth opercular band surrounding lid. Lid with obscure, irregularly shaped follicle cell impressions, but punctations absent (Fig.). Micropylar row located on lid; orifices raised, canals obscure (Fig. 15).	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765A0E465084FF7BFBFDFC8C.taxon	discussion	Comments. Eggs of Doroneuria are large, oval, and have small collars. The chorionic surface has a broad median zone of shallow pits, a narrow smooth opercular band and micropyles located on the lid near the anterior pole (Figs. 9, 13). The eggs of the two species are quite similar, but differ subtly among specimens examined in two respects. The punctate zone for D. theodora eggs extends nearer the collar and is, therefore, considerably broader in that species than in D. baumanni (Figs. 10, 14). In addition, the lid of D. theodora eggs lacks punctations, whereas in D. baumanni this area has obscure punctations which outline the irregular margins of follicle cell impressions (Fig. 12).	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765B0E47515BFF7BFA8AFDD6.taxon	description	(Figs. 17 - 20)	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765B0E47515BFF7BFA8AFDD6.taxon	materials_examined	Holotype ♂ (Natural History Museum, Los Angeles County), Gibson Creek, Mendocino Co., California Material examined. California: Plumas Co., Domingo Springs, 8 miles NW Chester, Old Red Bluff Road, 21 September 2007, J. Sandberg, 2 ♂, 5 ♀ (BPS). Sierra Co., Big Spring, Hwy 49, 2 miles N Bassetts, 3 July 2010, P. A. Opler, 1 ♀ (CSU).	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765B0E47515BFF7BFA8AFDD6.taxon	description	Egg. Outline oval with short, wide collar (Fig. 17); collar base encircled by prominent raised ridge, sides of egg body adjacent to ridge with a series of vertical struts. Egg length ca. 400 μm, equatorial width ca. 360 μm, collar length ca. 50 μm, median width ca. 75 μm. Sides of collar with several prominent longitudinal ridges (Fig. 19); collar rim flanged and irregularly incised (Figs. 18, 20). Anchor biscuit shaped with moderately long pedicel; surface covered with mushroom bodies. Chorionic surface covered with shallow, hexagonal follicle cell impressions. Micropylar row subequatorial.	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765B0E4A516DF899FD24FC5E.taxon	description	(Figs. 21 - 30)	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
038A8786765B0E4A516DF899FD24FC5E.taxon	materials_examined	Holotype ♀ (Museum of Comparative Zoology), Olympia [Thurston Co.], Washington Material examined. Alberta: Battle Creek at Reservoir Lake, 23 June 1975, L. Dosdall, 1 ♀ (BPS). California: Humboldt Co., East Fork Willow Creek, Hwy 299, 24 April 1987, B. Stark, R. W. Baumann, C. R. Nelson, 2 ♀ (BPS). Shasta Co., North Fork Battle Creek, south of Shingletown, 8 June 2004, B. Stark, R. W. Baumann, 1 ♀ (BPS). Tehama Co., North Fork Battle Creek, Wildcat Road, 26 April 1987, B. Stark, R. W. Baumann, C. R. Nelson, 2 ♀ (BPS). Colorado: Gunnison Co., Gunnison River, Hwy 135, 8 June 2003, B. Stark, 3 ♂, 2 ♀ (BPS). Idaho: Bonneville Co., Snake River, 4 July 1972, G. Tourtlotte, 1 ♂, 1 ♀ (BPS). Fremont Co., Big Springs, 5 miles east Macks Inn, 25 July 1979, B. Stark, K. W. Stewart, R. W. Baumann, 4 ♀ (BPS). Montana: Carbon Co., East Rosebud River, Roscoe, 23 July 1989, B. Stark, 5 ♂, 3 ♀ (BPS). Lewis & Clark Co., Trout Creek, Vigilante Campground, 21 August 2004, B. Stark, I. Sivec, 1 ♂, 1 ♀ (BPS). New Mexico: San Miguel Co., Pecos River, 6 miles north Pecos, 12 July 1973, B. Stark, 2 ♂, 3 ♀ (BPS). Nevada: Elko Co., Secret Creek, Secret Pass, 15 June 1974, B. Stark, 1 ♂, 1 ♀ (BPS). Oregon: Jefferson Co., Metolius River at headwater spring source, 11 July 1979, B. Stark, K. W. Stewart, 5 ♂, 3 ♀ (BPS). Tillamook Co., Nehalem River at Cronin Creek jct., 4 June 1991, B. Stark, R. W. Baumann, C. Henderson, 1 ♀ (BPS). Wyoming: Park Co., Clarks Fork Yellowstone River, below Cooke City, 20 July 1989, B. Stark, 3 ♂, 1 ♀ (BPS). Egg. Outline oval with short, wide collar (Fig. 21); collar base encircled by a prominent basal ring, ca. 137 - 176 μm wide (Figs. 23, 25, 27, 29); sides of egg body along basal ring bearing a series of vertical ridges separated by wide depressions. Egg length ca. 428 - 484 μm, equatorial width ca. 312 - 324 μm, collar length ca. 34 - 55 μm, median collar width ca. 41 - 58 μm, rim width ca. 89 - 116 μm. Sides of collar with several prominent vertical ridges, rim flanged and irregularly incised. Chorionic surface appearing smooth (Figs. 21, 26, 28, 30) but with obscure hexagonal follicle cell impressions sometimes apparent at higher magnifications. Micropyles located near equator; orifices raised slightly (Fig. 24). Comments. Hesperoperla eggs are generally similar to those of Calineuria californica in shape, collar form and in having a basal ridge encircling the collar base (Baumann & Stark 1980; Isobe 1997). The most conspicuous feature distinguishing the eggs of the two genera is the row of vertical struts on the egg body which connect to the basal ridge in Hesperoperla (Fig. 25) which are absent in eggs of Calineuria (Fig. 2). No consistent morphological differences were found among eggs of different populations of H. pacifica, or between eggs of H. hoguei and H. pacifica. However, variations are evident in the images of H. pacifica eggs from different sites (Figs. 21 - 30), especially in shape of the anterior pole (e. g. Figs. 26, 28), shape of the collar (e. g. Figs. 25, 27) and in length of lines delimiting cells surrounding the basal ridge (= vertical struts). The specimen shown from the Gunnison River, Colorado, for example, has a few long lines of at least 57 μm (Fig. 25), whereas the longest vertical struts on the specimen shown from Trout Creek, Montana (Fig. 27) are only 30 μm.	en	Stark, Bill P., Green, Stephen (2011): Eggs Of Western Nearctic Acroneuriinae (Plecoptera: Perlidae). Illiesia 7 (17): 157-166, DOI: http://doi.org/10.5281/zenodo.4760209
