taxonID	type	description	language	source
1B0987B1FF89024DFE84FA60D0DCFEC7.taxon	description	(Figs. 1 - 15, 36 - 37, 42, 45) http: // lsid. speciesfile. org / urn: lsid: Plecoptera. speciesfile. org: TaxonName: 3555	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF89024DFE84FA60D0DCFEC7.taxon	materials_examined	Material examined. UNITED STATES: California: Modoc Co., Rush Creek, Hwy 299, Upper Rush Creek Campground, 24 May 2007, R. W. Baumann, B. C. Kondratieff, 1 ♂ (BYU). Colorado: Eagle Co., Deep Creek, 17 Rd, 27 June 1997, B. C. Kondratieff, 11 ♂, 1 ♀ (CSUIC). Gilpin Co., South Boulder Creek, 2 miles W Tolland, 7 July 1991, B. C. Kondratieff, R. Durfee, 16 ♂, 10 ♀ (CSUIC). Grand Co., Colorado River, Kawuneeche Valley, Rocky Mountain National Park, 18 June 1980, B. C. Kondratieff, 3 ♂, 6 ♀ (CSUIC). Gunnison Co., Quartz Creek, 76 Rd, N of Pitkin, 7 July 2008, K. W. Stewart, 25 ♂, 38 ♀ (BPSC). Idaho: Bannock Co., Mill Creek, Summit Campground, 5 June 2010, B. Stark, R. W. Baumann, 2 ♂ (BPSC). Nevada: Elko Co., Lamoille Creek, Thomas Canyon Campground, 40 ° 39 ’ N, - 115 ° 24.4 ’ W. 23 June 2006, S. M. Clark, 7 ♂, 10 ♀ (BYU). Humboldt Co., Indian Creek, Santa Rosa Range, 29 June 1999, A. L. Sheldon, 10 ♂, 19 ♀ (BYU). Mahogany Creek, Summit Lake Mountain, 1 July 1999, A. L. Sheldon, 16 ♂, 14 ♀ (BYU). Nye Co., South Fork Pine Creek, Toquima Range, 7 July 1999, A. L. Sheldon, 14 ♂, 5 ♀ (BYU). Washoe Co., Rock Creek, Granite Range, 5 June 1998, A. L. Sheldon, 1 ♂ (BYU). White Pine Co., Berry Creek, Schell Creek Range, Berry Creek Campground, 24 June 2006, R. W. Baumann, B. C. Kondratieff, 15 ♂, 9 ♀ (BYU). Timber Creek, Timber Creek Campground, Schell Creek Range, 39 ° 24 ’ N, - 114 ° 38 ’ W, R. W. Baumann, B. C. Kondratieff, 35 ♂, 14 ♀ (BYU). New Mexico: Rio Arriba Co., Rio de las Vallecitos, 4 miles N Cañón Plaza, 11 June 1985, R. L. Hassage, 1 ♂ (BYU). San Miguel Co., Terrero, Pecos River, 10 June 1974, B. Stark, T. Wolfe, 3 ♂, 6 ♀ (BPSC). Oregon: Harney Co., McCoy Creek, 1 mile above Fish Lake, Steens Mountains, 20 July 1969, S. G. Jewett, Jr. 3 ♂, 10 ♀ (BYU). Umatilla Co., Camas Creek, Hwy 395, Ukiah Dale State Park, 16 May 2001, B. Stark, K. W. Stewart, J. B. Sandberg, 3 ♂, 2 ♀ (BPSC). North Fork John Day River, Hwy 395, 12 miles S Ukiah, 16 May 2001, B. Stark, K. W. Stewart, J. B. Sandberg, 5 ♂, 1 ♀ (BPSC). Union Co., Grande Ronde River, Bird Track Springs Campground, Hwy 244, 45 ° 18.027 ’ N, - 118 ° 18.427 ’ W, 15 May 2014, B. Stark, A. Harrison, 1 ♂ (BPSC). South Dakota: Lawrence Co., Spearfish Creek, Hwy 85, Cheyenne Crossing, 7 June 1995, R. W. Baumann, B. Huntsman, 10 ♂, 14 ♀ (BYU). Utah: Box Elder Co., Clear Creek, Raft River Mountains, 18 June 1979, R. W. Baumann, G. M. Webb, 23 ♂, 20 ♀ (BYU). Juab Co., Granite Creek, Deep Creek Mountains, 14 April 1984, R. W. Baumann, 18 ♂, 5 ♀ (BYU). Salt Lake Co., Big Cottonwood Creek, 22 June 1974, B. Stark, 10 ♂, 5 ♀ (BPSC). Sanpete Co., Willow Creek, Ephraim Canyon, 39.29707 N, - 111.52377 W, 28 June 2005, Johnson, Anderson, 14 ♂, 16 ♀ (BYU). Tooele Co., Harker Creek, Harker Canyon, Sheep Rock Mountains, 39 ° 59 ’ N, - 112 ° 39 ’ W, 30 May 2012, R. W. Baumann, G. M. Webb, 8 ♂, 5 ♀ (BYU). South Willow Creek, Boy Scout Campground, Stansbury Mountains, 11 June 2007, R. W. Baumann, K. Anderson, 20 ♂, 21 ♀ (BYU). Wasatch Co., Cascade Springs, Uintah National Forest, 27 May 2000, B. Stark, 8 ♂, 4 ♀ (BPSC). Daniels Canyon, Whiskey Springs Picnic Area, 40 ° 24.677 ’ N, - 111 ° 19.278 ’ W, 13 June 2015, B. Stark. 7 ♂, 12 ♀ (BPSC). Wyoming: Big Horn Co., West Tensleep Creek, N of West Tensleep Lake, 19 July 2005, D. E. Martin, 2 ♂ (BYU). Fremont Co., Jakeys Creek, Hwy 26, 5 miles E of Dubois, 20 July 1967, R. W. Baumann, 2 ♂, 2 ♀ (BYU). Washakie Co., 12.5 km ESE Tensleep, 25 June 1996, K. B. Miller, 9 ♂, 10 ♀ (CSUIC).	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF89024DFE84FA60D0DCFEC7.taxon	description	Adult habitus. Described by Needham & Claassen (1925). Male forewing length 7.0 - 7.5 mm, female 8.5 - 9.0 mm. Head and pronotum pale yellow with dark markings; ocelli connected by a V-shaped dark brown mark, M-line and anteromedian area of head forward of M-line dark brown. Pronotum pale yellow with almost complete dark brown margins and rugosities (Fig. 42). Abdomen brown with subtle, slightly darker median brown stripe. Cerci pale, antennae pale basally but brown in apical two thirds. Wing membrane pale, veins pale brown. Male epiproct (n = 12). Maximum width 320 - 370 μm, length 360 - 450 μm. Base relatively narrow, but gradually expanded beyond cowl, and reaching maximum width across lateral projections of epiproct body. Shoulders project conspicuously beyond hook base (Figs. 1 - 13, 45). Aedeagus (n = 3). Length 500 - 524 μm, width 590 - 760 μm. Entirely membranous; apex with a pair of small finger-shaped lobes in a parallel or V-pattern, apicolateral margins, each bearing a large, projecting, rounded lobe, and mesoventral area with a pair of eye-like lobes (Figs. 14 - 15); Fingershaped lobes bear scattered setae of variable length, apicolateral lobes bare; mesoventral lobes bare, but each with a transverse groove. Median ventral field below mesoventral eye-shaped lobes with a low nose-like tubercle. Female subgenital plate. Posterior margin of plate projecting over basal third to half of sternum 9; plate truncate, to slightly concave along posterior margin. Plate usually darkly pigmented over most of surface, contrasting with the pale sternum 9 and the pale basal and lateral areas of sternum 8 (Figs. 36 - 37).	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF89024DFE84FA60D0DCFEC7.taxon	diagnosis	Diagnosis and distribution. This species, in which the epiproct shoulders conspicuously project beyond the base of the epiproct hook (Fig. 45), is the common form in the central Rockies and Great Basin, and the probable “ true ” S. coloradensis. Published records exist for S. coloradensis from Alberta, British Columbia and Yukon Territory in Canada, and from Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, South Dakota, Utah, Washington and Wyoming in the United States (DeWalt et al. 2017; Surdick 1985). Many of the published records will apply to one of the two newly described species below. The Montana, Washington, Alberta, British Columbia and Yukon specimens examined by us have all been determined as S. lyrata sp. n., and the Arizona and western New Mexico specimens are determined as S. mogollonica sp. n. Specimens from the Central Rocky Mountains of Colorado and Utah are identified as S. coloradensis, Some states (e. g. Idaho and New Mexico) have populations of at least two members of the complex. We have recognized populations of S. coloradensis from the sites listed in the “ Material examined ” section above.	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF8B0240FC4DFE50D2E1FEC7.taxon	description	(Figs. 16, 18 - 29, 38 - 39, 43, 46) http: // lsid. speciesfile. org / urn: lsid: Plecoptera. speciesfile. org: TaxonName: 502536	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF8B0240FC4DFE50D2E1FEC7.taxon	materials_examined	Material examined. Holotype ♂, 3 ♂, 1 ♀ paratypes, Washington, Skagit Co., Friday Creek, Prairie Rd, 5 miles N of Burlington, 16 April 1967, K. E. Vander Mey (holotype, United States National Museum, paratypes BYU). Additional paratypes: CANADA: Alberta: Cameron Creek, Cameron Lake, Waterton Lakes National Park, 19 July 1965, A. R. Gaufin, 26 ♂, 30 ♀ (BYU). British Columbia: Duhamel Creek, Six Mile Rd, 30 May 2010, B. Stark, R. W. Baumann, 2 ♂, 1 ♀ (BPSC). Yukon: Klondike River, Dempster Hwy, 23 June 1996, J. C. Abbott, K. W. Stewart, 8 ♂, 7 ♀ (BPSC). Mayo River, Hwy 11, 63 ° 36 ’ N, - 135 ° 55 ’ W, 24 June 1996, K. W. Stewart, J. C. Abbott, 5 ♂, 3 ♀ (BYU). McQuesten River, Hwy 2, 63 ° 33 ’ 15.88 ” N, - 137 ° 24 ’ 37.88 ” W, 24 June 1996, K. W. Stewart, J. C. Abbott, 3 ♂, 6 ♀ (BYU). Ogilvie River, Dempster Hwy, 65 ° 21 ’ 38.82 ” N, - 138 ° 18 ’ 20.73 ” W, 21 June 1996, K. W. Stewart, J. C. Abbott, 17 ♂, 5 ♀ (BYU). UNITED STATES: Idaho: Bonner Co., Pack River, Pack River Rd, 26 May 2010, B. Stark, R. W. Baumann, 5 ♂, 8 ♀ (BPSC). Montana: Gallatin Co., Duck Creek, 8 miles N West Yellowstone, 9 June 1987, B. C. Kondratieff, 1 ♂, 2 ♀ (CSUIC). Gallatin River, Squaw Creek Rd, 10 July 1999, B. Stark, L. Stark, 1 ♂, 1 ♀ (BPSC). Mineral Co., St. Regis River, Little Joe Rd, St. Regis, 25 May 2010, B. Stark, R. W. Baumann, 28 ♂, 8 ♀ (BPSC). St. Regis River, Riverside, 11 June 1969, A. R. Gaufin, 4 ♂, 7 ♀ (BYU). Trout Creek, Trout Creek Campground, 4 June 2010, B. Stark, R. W. Baumann, 23 ♂, 6 ♀ (BPSC). Washington: Okanogan Co., Buck Creek, 20 miles N of Winthrop, 9 June 1995, MacKenzie, 4 ♂, 1 ♀ (BYU). Pend Oreille Co. Sullivan Creek, below Sullivan Lake, 2 June 2010, B. Stark, R. W. Baumann, 35 ♂, 43 ♀ (BPSC). Whatcom Co., Padden Creek, 33 rd Street, Bellingham, 27 April 1994, Graham, MacKenzie, 6 ♂, 1 ♀ (BYU). Adult habitus. Male forewing length 6.2 - 6.7 mm; female 7.0 - 7.5 mm. Similar to S. coloradensis with hieroglyphic-like markings on head and pronotum (Fig. 43). However considerable variation occurs in the pigmentation patterns of individuals within and between sites. Our impression is that the three species overlap in pigment pattern.	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF8B0240FC4DFE50D2E1FEC7.taxon	description	Male epiproct (n = 14). Maximum width 310 - 410 μm, length 360 - 470 μm. Base conspicuously narrowed at stem and epiproct relatively wide across body at lateral projections giving a distinct stalked appearance (Figs. 18 - 29). Shoulders scarcely, or not at all projecting beyond base of hook (Fig. 46). Aedeagus (n = 1). Length from base to base of apical finger-like lobes 466 μm, width across eye-like lobes to apices of lateral lobes 525 μm. Lateral lobes broad basally near outer margins of eye-like lobes; margins of lateral lobes strongly convergent, forming an overall football-shape to the aedeagal body (Fig. 16). Female terminalia. Posterior margin of subgenital plate truncate, overlapping that of S. coloradensis in shape (Figs. 38 - 39). Larva. Unknown.	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF8B0240FC4DFE50D2E1FEC7.taxon	etymology	Etymology. The species name refers to the lyre- shaped dorsal aspect of the epiproct of this species. Diagnosis and distribution. Sweltsa lyrata is recognized by the prominent lateral projections, small apical projections and abruptly narrowed epiproct stem that give the epiproct a lyre-like shape. Males of this species are distinguished from those of S. mogollonica by the enlarged lateral lobes of the epiproct (Figs. 17 - 28), and from S. coloradensis by the prominent apical lobes of the epiproct in that species (Figs. 2 - 13). The aedeagus of this species (based on one specimen) is shorter than both additional species of the complex, and has the lateral lobes wider and more pointed at their apex. Populations exhibiting the S. lyrata epiproct type are known from the Northern Rocky Mountains including sites in Alberta, British Columbia and Yukon Territory and adjacent areas in Washington, Montana and Idaho.	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF860247FEF0FE5CD1E4FAD4.taxon	description	(Figs. 17, 30 - 35, 40 - 41, 44, 47) http: // lsid. speciesfile. org / urn: lsid: Plecoptera. speciesfile. org: TaxonName: 502537	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF860247FEF0FE5CD1E4FAD4.taxon	materials_examined	Material examined. Holotype ♂, 1 ♂, 10 ♀ paratypes, Arizona, Apache Co., North Fork White River, Hwy 473, 12 June 1974, B. Stark, (holotype United States National Museum, paratypes BPSC). Additional paratypes, Arizona: Graham Co., Grant Creek, Hospital Flat Campground, Graham Mountains, 5 June 1982, R. W. Baumann, S. M. Clark, 12 ♂, 16 ♀ (BYU). Greenlee Co., Hannagan Creek, Hannagan Meadow, Sitgreaves National Forest, 29 May 1993, C. R. Nelson, J. K. Gelhaus, 29 ♂, 14 ♀ (BYU). New Mexico: Catron Co., Willow Creek, Willow Creek Campground, 11 June 1974, B. Stark, 8 ♂, 7 ♀ (BPSC). Adult habitus. Pigment pattern not distinct from that of S. coloradensis or S. lyrata (Fig. 44). Male forewing length 6.8 - 7.5 mm, female forewing length 7.8 - 9.0 mm. Male epiproct. Maximum width 320 - 340 μm, length 390 - 430 μm. Base and stem relatively wide and usually, more or less continuous with epiproct body (Figs. 30 - 35). Apical lobes not projecting from shoulders (Fig. 47). Aedeagus (n = 1). Length 357 μm, width 614 μm. Similar in shape and setation to S. coloradensis but conspicuously shorter in the only everted specimen available (Fig. 17). Female terminalia. Subgenital plate not always distinct from that of S. coloradensis or S. lyrata. The specimen illustrated in Fig. 39 - 40 has a more rounded subgenital plate than is typical for the other species. Larva. Unknown.	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF860247FEF0FE5CD1E4FAD4.taxon	etymology	Etymology. This species name is based on the Mogollon Rim of Arizona and New Mexico.	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
1B0987B1FF860247FEF0FE5CD1E4FAD4.taxon	diagnosis	Diagnosis and distribution. Epiprocts of specimens of this species have, at most, slightly projecting shoulders near the hook base and are relatively wide across the lateral lobes (Figs. 29, 32). The new species is similar to S. lyrata in the former character but the two are distinguished on the basis of the relatively greater width of the epiproct for males of S. lyrata (compare Figs. 19 and 29). The aedeagus may also be distinct (see Fig. 17), but verification is needed from additional material. Specimens are known from three Arizona sites, and an adjacent site in New Mexico.	en	Stark, Bill P., Baumann, Richard W. (2018): Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae). Illiesia 14 (2): 30-43, DOI: http://doi.org/10.5281/zenodo.4761184
