taxonID	type	description	language	source
03E487F6023C937B0A98FBD84F3F86E8.taxon	description	(Figs. 1 - 5)	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023C937B0A98FBD84F3F86E8.taxon	materials_examined	Material examined. Holotype ♂, 6 ♂, 6 ♀ paratypes (all pinned) from Guyana, District 8, Mount Wokomung, 1234 m, 05 ° 06 ’ 34.8 ” N, 59 ° 49 ’ 15.3 ” W, Mercury vapor light, 27 October- 1 November 2004, ROM 2004515, B. Hubley (ROM).	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023C937B0A98FBD84F3F86E8.taxon	description	Adult habitus. General appearance pale yellow brown with obscure darker markings. Head entirely pale except for pale brown antennae; pronotum pale over most of disk but with narrow, dark brown pigment bands near lateral margins. Legs pale except for narrow, dark bands at distal end of femora. Wings pale but with slightly darker yellow-brown longitudinal band over bases of anal veins, extending to apical margin but with an obscure pale circular area beyond cord; first cubital vein and cord crossveins slightly darker. Male. Forewing length 10 - 11 mm. Hammer long and slender, posteromesal margin of sternum 9 prolonged (Fig. 1). Aedeagal apex conspicuously narrower than shoulders, and bearing a poorly defined pair of ventral membranous lobes (Fig. 3); dorsal keel poorly defined, consisting of a short stem and a pair of divergent arms which circumscribe upturned tip (Fig. 5); keel lines interrupted along length, consisting of a linear series of small raised areas. Apex turned ventrad giving a smoothly curved lateral profile (Fig. 4). Aedeagal hooks slender. Female. Forewing length 12 - 13 mm. Subgenital plate 4 - lobed (Fig. 2); mesal lobes larger than lateral ones and separated by a relatively deep, V-shaped notch; lateral lobes separated by shallow V-shape notches from mesal lobes. Mesal sclerite of sternum 9 with a pair of prominent lateral patches of setae, separated by a median, triangular patch of fine short setae. Posterior transverse sclerite obscure. Larva. Unknown.	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023C937B0A98FBD84F3F86E8.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the Pakaraima Mountain range of Guyana.	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023C937B0A98FBD84F3F86E8.taxon	diagnosis	Diagnosis. This species is similar in pigment pattern to A. wokomung, described below, however, the pattern is not as distinct in the specimens available, and might be overlooked. Adults of these two Mount Wokomung species are separated on the basis of genital structures; externally A. pakaraima has the hammer long and slender (Fig. 1), whereas that of A. wokomung is wide with parallel margins (Fig. 8). The aedeagal apex of the two species are also quite distinctive with A. wokomung having a short, but conspicuous dorsal keel line (Fig. 11) and A. pakaraima having inconspicuous keel lines that diverge and circle around a portion of the curved aedeagal tip (Fig. 5). The female subgenital plates of the two species are also distinct with A. pakaraima having a much deeper mesal notch (Figs. 2, 13).	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023F937A0817FF4F49C38461.taxon	description	(Figs. 6 - 13)	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023F937A0817FF4F49C38461.taxon	materials_examined	Material examined. Holotype ♂ and 2 ♀ paratypes from Guyana, District 8, Mount Wokomung, 736 m, 05 ° 07 ’ 43.2 ” N, 59 ° 48 ’ 40.0 ” W, Malaise trap, 21 - 26 October 2004, ROM 2004531, B. Hubley (ROM). Additional paratypes (pinned). Guyana: District 8, Mount Wokomung, 1411 m, 05 ° 05 ’ 33.4 ” N, 59 ° 50 ’ 34.5 ” W, Mercury vapor light, 2 - 8 November 2004, ROM 2004521, B. Hubley 4 ♂, 1 ♀ (ROM).	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023F937A0817FF4F49C38461.taxon	description	Adult habitus. General appearance in alcohol pale white with limited brown pigment pattern. Head entirely pale except for pale brown antennae (Fig. 6); pronotum pale over most of disk, but with narrow, dark brown pigment bands near lateral margins. Legs pale except for narrow dark bands at distal ends of tibiae and femora. Wings pale along costal margins including subcosta and radius, but with pale amber pigment forming a longitudinal band over bases of anal veins, extending across cubito-median veins to apical margin (Fig. 7). A circular unpigmented area is located distal to cord crossveins; the first cubital vein and cord crossveins are dark brown. Male. Forewing length 10.5 - 11 mm. Hammer strongly sclerotized with parallel lateral margins; height about equal to width (Fig. 8). Aedeagal apex broadly rounded in lateral aspect (Fig. 10), bearing a short recurved, ventroapical tip (Fig. 9), and a short, thick dorsal keel line (Fig. 11). Ventral membranous lobes absent, but basolateral area of aedeagal apex bulging slightly. Aedeagal hooks slender. Female. Forewing length 13 - 14 mm. Subgenital plate 4 - lobed (Fig. 13). Inner lobes separated by a shallow, median notch, but notch separating outer lobes acute. Mesal sclerite of sternum 9 bearing a basomedian patch of fine short setae and patches of more prominent setae extending to posterior margin of segment. Posterior sclerite poorly developed. Intersegmental membrane armed with fine microtrichia. Egg. Outline spindle shaped, typical of genus (Fig. 12). Larva. Unknown.	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023F937A0817FF4F49C38461.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the type locality.	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023F937A0817FF4F49C38461.taxon	diagnosis	Diagnosis. Because the wing pigment pattern is similar, this species is identified as A. paria Stark, 1999 in the key proposed by Stark (1999) for Anacroneuria known for northeastern South America. However, the ventroapical region of the aedeagus in that species appears trilobed (see Fig. 34 in Stark 1999), the hammer is a short, thimble-shaped structure (see Fig. 33 in Stark 1999) and the female subgenital plate bears a moderately deep median notch (see Fig. 54 in Stark 1999). The male and female reproductive structures for A. wokomung are not similar to other regional species (Stark 1999; 2000). The following key to males of Guyana Anacroneuria is presented to aid in recognition of the two new species. Figure numbers referenced in the key refer to Stark (1995, 1999; 2000), or to those presented in this study.	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
03E487F6023F937D0A64FC924953848C.taxon	discussion	(Male A. wapishana Stark 1999 unknown)	en	Stark, Bill P. (2012): Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae). Illiesia 8 (10): 114-118, DOI: http://doi.org/10.5281/zenodo.4760603
