taxonID	type	description	language	source
250A8780EF0CDE04FF4EFD68FE9653D4.taxon	materials_examined	Holotype ♂ (MEUV), Alto Anchicayá, Valle Del Cauca, Colombia	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE04FF4EFD68FE9653D4.taxon	materials_examined	Material examined. Ecuador: Pastaza, Cushueme, Rio Cushueme, 320 m, 150 km SE Puyo, 15 - 30 May 1971, B. Malkin, 1 ♂ (CASC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE04FF4EFD68FE9653D4.taxon	discussion	Comments. This species was previously reported from Colombia and from Cotopaxi, Napo and Pichincha provinces in Ecuador (Stark et al., 1999; Stark 2001).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE04FF46FB62FAE552B5.taxon	description	(Figs. 1 - 5)	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE04FF46FB62FAE552B5.taxon	materials_examined	Material examined. Holotype ♂ and 1 ♂ paratype, Ecuador, Province Imbabura, Canton Cotacachi, Rio Intag, Apuela, 5020 ft, 00 ° 21.2 ’ N, 78 ° 31.0 ’ W, 6 November 2009, S. M. Clark, H. R. Hinkson (Holotype USNM, paratype BYUC). Adult habitus. General color brown patterned with darker brown. Ocellar region covered with moderately dark brown pigment extending to central frons, becoming darker forward of poorly defined Mline; lappets and antennae brown (Fig. 1). Pronotum pale along median suture and along lateral margins; disk predominantly brown with scattered pale rugosities. Wing membrane pale brown, veins darker except costa pale. Femora dark over apical half and along dorsal and ventral margins for most of length; tibiae dark along outer margin and in basal and apical fourth. Male. Forewing length 10 mm. Apical section of aedeagus gradually narrowed from shoulders, then slightly widened subapically and narrowed again to slightly notched tip (Fig. 3). Dorsal keel consists of two narrowly separated and broken lines which diverge sharply and reach lateral margins near subapical wide point (Fig. 5). Ventral aedeagal apex covered by a large pair of membranous lobes. Hooks large, subchelate and narrowed to acute tips. Lateral aspect of apical section shaped somewhat like a bird’s head (Fig. 4). Hammer thimble shaped (Fig. 2). Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE04FF46FB62FAE552B5.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the type locality.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE04FF46FB62FAE552B5.taxon	diagnosis	Diagnosis. This species is a member of the A. aymara Stark & Sivec, 1998 species group discussed by Zúñiga & Stark (2002). The group also includes A. chaima Stark, 1999, A. cotacachi Stark, 2001, A. farallonensis Rojas & Baena, 1993 (see Zúñiga & Stark 2002), A. perija Stark, 1999, A. portilla Stark & Rojas, 1999 (in Stark et al. 1999), A. ricki Zúñiga & Stark, 2002, A. segnini Stark & Maldonado, 2002 (in Maldonado et al. 2002), and A. toni Zúñiga & Stark, 2002, and appears to be widely distributed in the northern and central Andes. The new species is most similar to A. toni in structure of the dorsal aedeagal keel, sharing with that species the long, apically divergent pair of keel lines. However, in A. apuela these lines are broken, and appear as a series of small bumps separated by narrow gaps. In addition, the ventral membranous lobes are much larger than in A. toni (see Zúñiga & Stark 2002, Figs. 17 - 19).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE07FC17FA7AFC6B5078.taxon	materials_examined	Holotype ♂ (NHML). Amazon River, São Paulo de Olivença, Brazil	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE07FC17FA7AFC6B5078.taxon	materials_examined	Holotype ♂ (USNM). Iquitos, Loreto, Peru. Syn. Froehlich, 2008: 129.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE07FC17FA7AFC6B5078.taxon	materials_examined	Material examined. Ecuador: Pastaza, Rio Cushueme, Cushueme, 150 km SE Puyo, 320 m, 15 - 30 May 1971, B. Malkin, 1 ♂ (CASC). Same site, 23 July 1971, B. Malkin, 2 ♂ (CASC). Paraguay: Dept. Canindeyu, R. N. B. Mbaracayu, 183 m, 24 ° 08.1 ’ S, 55 ° 31.7 ’ W, 8 - 10 January 2012, D. Brzoska, 1 ♂, 4 ♀ (CUSC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0CDE07FC17FA7AFC6B5078.taxon	discussion	Comments. This species has been widely reported from the upper Amazonas region of Brazil, Colombia and Peru (Froehlich 2008; Stark & Sivec 1998; Stark et al. 2001), but not previously from Ecuador or Paraguay.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0EDE06FE9AF92AFB855078.taxon	materials_examined	Material examined. Ecuador: Pichincha, Santo Domingo de Los Colorados, 500 m, 3 - 8 April 1971, B. Malkin, 1 ♂ (CASC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0EDE06FE9AF92AFB855078.taxon	discussion	Comments. Stark (2001) previously reported this species from three sites in the general vicinity of this collection in Pichincha.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FEF4FF7EFE3A5533.taxon	materials_examined	Holotype ♂ (CMNH), Chical, Carchi, Ecuador	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FEF4FF7EFE3A5533.taxon	materials_examined	Material examined. Ecuador: Esmeraldas, Cantόn San Lorenzo, Rio Lita, 1830 ft, 00 ° 51.64 ’ N, 78 ° 27.16 ’ W, 2 December 2008, R. C. Mower, 2 ♂ (BYUC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FEF4FF7EFE3A5533.taxon	discussion	Comments. This species was known previously from the holotype (Stark 2001).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FEA2FDF8FDC95219.taxon	materials_examined	Holotype ♂ (CASC). 67 mi E Tingo Maria, Yurac, Peru	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FEA2FDF8FDC95219.taxon	materials_examined	Material examined. Ecuador: El Oro, 9 mi S Santa Rosa, 23 January 1955, E. I. Schlinger, E. S. Ross, 11 ♂ (CASC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FEA2FDF8FDC95219.taxon	discussion	Comments. Forewing lengths for these males range from 8 - 9 mm and the holotype forewing length is 10.5 mm. This difference and subtle differences in the aedeagal apex suggest these specimens may represent a sibling species in this complex. Specifically, the narrow dark markings along the posterior shoulder margins are less distinct for the Ecuadorian specimens and the lateral margins of the aedeagal apex are not constricted at the shoulder as they are in the holotype. A larger sample from the type locality in Peru is needed to more fully evaluate the status of these specimens.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FF51FACEFC2653BE.taxon	description	(Figs. 6 - 10)	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FF51FACEFC2653BE.taxon	materials_examined	Material examined. Holotype ♂ and 1 ♂ paratype, Ecuador, Pastaza, Rio Cushueme, Cushueme, 150 km SE Puyo, 320 m, 15 - 30 May 1971, B. Malkin (CASC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FF51FACEFC2653BE.taxon	description	Adult habitus. General color pale yellow patterned with pale brown. Head pattern indistinct, almost entirely yellow with only slightly darkened lappets (Fig. 6). Pronotum with sublateral pale brown bands; area adjacent to median suture and anterolateral margins pale. Femora pale brown, tibiae slightly darker at apex and base. Wings transparent, veins pale amber. Male. Forewing length 9 mm. Apical section of aedeagus with median, projecting lobe narrowed slightly from base and truncate at tip (Fig. 8). Shoulders strongly sclerotized, bulging laterad; body of aedeagus wider proximal to shoulder bases. Dorsal keel consists of two long, divergent ridges and two pairs of shorter ones near base of longer median keel lines (Fig. 10). Hooks relatively wide and acute. Lateral lobes on apical section project away from aedeagal body, one pair lunate and the other almost quadrate (Fig. 9). Hammer thimble shaped (Fig. 7). Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FF51FACEFC2653BE.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the type locality.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE01FF51FACEFC2653BE.taxon	diagnosis	Diagnosis. Based on aedeagal morphology, this species is related to the A. yameo Stark & Sivec 1998 group of species, which also includes A. pakitza Stark & Sivec 1998, A. pinza Stark 1995, and A. zunigae Stark 2001. The new species bears a greater similarity in aedeagal structure to A. yameo and A. pakitza but in these species the shoulder region bulges toward the aedeagal apex creating an apically trilobed structure; this is quite distinct in the dorsal aspect and differs from the comparable structures in A. cushueme which bulge more laterad. In A. cushueme the mesal projection of the aedeagal apex is truncate at the tip and broad at the base which also differs from both related species.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE02FCC2FB77FE805680.taxon	description	(Figs. 11 - 13)	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE02FCC2FB77FE805680.taxon	materials_examined	Material examined. Ecuador, Napo Province, 6.38 km W Baeza, Hwy E- 20, 1,987 m, 25 January 2012, B. C. Kondratieff, B. Gill 1 ♂, 1 ♀, 1 larva, 1 exuvium (CSUC). Napo Province, 8.68 km NW Baeza, Hwy E- 20, 2,134 m, 24 January 2012, B. C. Kondratieff, B. Gill, 1 ♂, 1 exuvium (CSUC). Larva. Body length pre-emergent female 16 - 17 mm. General color brown, patterned with dark brown. Head brown with pale areas at frontoclypeus, M-line, lateral to ocelli and over most of occiput (Fig. 11); Mline incomplete adjacent to pale mesal spot. Pronotum brown but with pale leg shaped areas adjacent to median suture, and with a few additional pale rugosities. Lateral pronotal setal fringe incomplete anterolaterally. Femora pale brown but darker along ventral and apical margins; fine setal fringe well developed; basal patch of short thick setae extensive along dorsal margin; transverse setal row consists of ca. 7 long, thick setae; scattered thick setae sparse beyond transverse row and with ca. 16 on lower half of femoral surface. Ventral margin with a few thick setae and interspersed fine short setae (Fig. 12). Abdomen dark brown but apical segments with a small median pale spot (Fig. 11), terga without apparent intercalary setae. Cerci without welldeveloped fine setal fringe (Fig. 13), only a few fine setae occur along dorsal margin of apical segments; cercal segments bear apical whorls of short, thick setae, except on some basal segments.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF09DE02FCC2FB77FE805680.taxon	discussion	Comments. The male specimen from the first site listed above varies slightly from typical A. guambiana specimens in having the apical aedeagal section with convergent lateral margins and in having the aedeagal shoulders approaching the apical section at an obtuse angle. Larger samples are needed to resolve whether this specimen represents a sibling member of the A. guambiana complex. The larva is generally similar to that of A. paleta Stark 1995 (Maldonado et al. 2002), but that species has a complete M-line and more extensive dark pigment on the occiput. These specimens are the first representatives of this species to be reported for Ecuador. The first site listed in the materials section above (Figs. 32 - 33) is located in the lower montane cloud forest at 1,987 m, and is composed mainly of alternating step-pools. The substrate ranges from depositional fines to large boulders, however, size extremes of sand and boulders dominate. The stream channel has numerous debris dams and downed trees from adjacent riparian zones. Steep walls surrounding the stream are covered with plants that confine the channel and create approximately 71 % canopy cover. The channel is 1.56 m wide and depth averaged 0.13 m. Discharge measured during the collection at this site was 37.1 L s- 1. Interestingly, the substrate was partially covered at the time of the collection by filamentous algae not typically seen in the region. The second stream (Fig. 34) is characterized by stretches of cobble filled riffle between cascades formed by large boulders. The channel is braided and narrow at various points, and is relatively free of debris dams or downed trees. Channel width is 2.32 m and depth averages 0.19 m. Discharge was measured at 105.3 L s- 1 during collecting. Canopy cover varies from open to closed along the channel. Adjacent riparian areas are composed of vegetation typical of lower montane cloud forest.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE02FEF2FE25FD835471.taxon	materials_examined	Material examined. Ecuador: Imbabura Province, Cantόn Cotacachi, Rio Intag, Apuela, 5020 ft, 00 ° 21.2 ’ N, 78 ° 31.0 ’ W, 6 November 2009, S. M. Clark, H. R. Hinkson 8 ♂ (BYUC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE02FEF2FE25FD835471.taxon	discussion	Comments. These specimens are the first to be reported from Imbabura Province.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE02FF74FC37FB785451.taxon	description	(Figs. 14 - 18)	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE02FF74FC37FB785451.taxon	materials_examined	Material examined. Holotype ♂, Ecuador, Napo, 6.38 km W Baeza, Hwy E- 20, 1,987 m, 0 ° 27.123 ’ S, 77 ° 56.468 ’ W, 25 January 2012, B. C. Kondratieff, B. Gill (USNM). Adult habitus. General color pale yellow, white in alcohol. Head with slightly darker, but obscure markings over ocelli, lappets and posterolateral areas of occiput (Fig. 14). Pronotum pale along lateral margins and median suture, but with irregular, dark brown, sublateral bands. Ocelli narrowly separated; wings transparent, veins pale. Legs, antennae and palpi pale. Male. Forewing length 17 mm. Hammer almost cylindrical, but slightly wider at base than apex (Fig. 15). Aedeagal apex truncate, lateral margins slightly expanded and appearing swollen at base (Figs. 16, 18); hooks strongly sclerotized at base with a distinct dark line outlining inner basal margin, and bearing a strong ventral keel from near midlength to near tips. Dorsal keel consists of two close-set, low ridges, extending for about half of apical area length (Fig. 18). In lateral aspect much of aedeagal apex membranous (Fig. 17). Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE02FF74FC37FB785451.taxon	etymology	Etymology. We honor Dr. Kayce Anderson, Colorado State University, for her dedication to ecological studies in Ecuador, and her appreciation of Neotropical aquatic biology.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE02FF74FC37FB785451.taxon	diagnosis	Diagnosis. The aedeagus of this species is similar to that of the Peruvian-Bolivian species, A. pachacuti Stark & Sivec 1998, particularly in lateral aspect. However, in that species the apical section of the aedeagus is offset by prominent basolateral notches and the aedeagal apex is nipple-shaped rather than truncate (see Stark & Sivec 1998, Figs. 124 - 126). In addition, A. pachacuti is a smaller species (FWL ~ 11 mm) with a slightly darker habitus. The aedeagus is also similar to that of A. rawlinsi Stark, 2001, but in that species the apical aedeagal section lacks swollen basolateral lobes.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE02FF74FC37FB785451.taxon	discussion	Comments. The type locality (Figs. 32 - 33) is described under A. guambiana.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE0DFCF6FC1AFA805078.taxon	description	(Figs. 19 - 23)	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE0DFCF6FC1AFA805078.taxon	materials_examined	Material examined. Holotype ♂, Ecuador, Pastaza, Rio Cushueme, Cushueme, 150 km SE Puyo, 320 m, 15 - 30 May 1971, B. Malkin (CASC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE0DFCF6FC1AFA805078.taxon	description	Adult habitus. General color yellow-brown patterned with darker pigment. Head mostly yellow but with a large dark brown patch covering ocelli, and with dark brown lappets. Antennae dark brown, pronotum brown with scattered pale rugosities and with a pale median band and with areas of lighter pigment along lateral margins (Fig. 19). Femora brown in apical half with narrow black band at knee; basal half of femora pale, tibiae brown. Wings transparent with pale amber tint, veins dark amber with R vein dark brown. Male. Forewing length 8 mm. Aedeagal apex narrowed from shoulders, rounded at tip, and bearing a pair of membranous ventral lobes (Fig. 23). Dorsal keel consists of a pair of divergent, narrow ridges and a small transverse ridge set at the base of the divergent ridges (Fig. 21); lateral aspect of aedeagal apex broad basally, narrowed to an acute tip and directed toward apex (Fig. 22). Aedeagal hooks gradually curved to acute tips. Abdominal sternum 9 bears a thimble shaped hammer (Fig. 20). Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE0DFCF6FC1AFA805078.taxon	etymology	Etymology. The patronym honors the late Borys Malkin who collected the holotype of this and many other species. Mr. Malkin was a famed naturalist and friend of the well-known Plecoptera authority, Stanley G. Jewett, Jr. The specimens were part of the Jewett collection directed to us by R. W. Baumann.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF0ADE0DFCF6FC1AFA805078.taxon	diagnosis	Diagnosis. The aedeagus of this species is generally similar to that of A. tzapino Stark 2001, a species also known from Pastaza. The aedeagal apex of that species is slightly notched, the dorsal keel arms are parallel for most of their length, and the apical aedeagal region slants slightly ventrad in lateral aspect (see Stark 2001, Figs. 118 - 120). In addition, A. tzapino has a more extensive area of brown pigment on the head and the wings are darker, but also have an obscure pale spot at the cord (Stark 2001).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF04DE0FFF07F9D5FA7455C0.taxon	description	(Figs. 24 - 28)	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF04DE0FFF07F9D5FA7455C0.taxon	materials_examined	Material examined. Holotype ♂, Ecuador, Pichincha, Santo Domingo de Los Colorados, 500 m, 3 - 8 April 1971, B. Malkin (CASC).	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF04DE0FFF07F9D5FA7455C0.taxon	description	Adult habitus. General color yellow-brown patterned with dark brown. Head with a quadrangular area covering ocellar area forward to about mid-frons; pigmented area with a slight anteromedian notch (Fig. 24); lappets pale brown, antennae brown. Pronotum dark over much of disk but pale along median suture, anterolateral margins and bearing a few small, pale rugosities. Femora pale brown except for narrow, dark knee band; tibiae pale except for small dark bands at base and near apex. Wing membrane dark amber with darker veins; R-vein particularly dark in basal half. Male. Forewing length 10.5 mm. Apical section of aedeagus indistinctly trilobed with narrow median lobe projecting well beyond lateral, somewhat bulging shoulders (Figs. 26, 28); entire apical area covered by large ventral membranous lobes; shoulders with short dorsal ridges extending inwards toward bases of median, finger-like lobe. Body of aedeagus constricted near apical third, but with low lateral bulges projecting over bases of hooks; tips of hooks narrowly truncate. Lateral lobes of aedeagal apex project away from aedeagal body in ear-like fashion (Fig. 27). Abdominal sternum 9 bears a thimble shaped hammer (Fig. 25). Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF04DE0FFF07F9D5FA7455C0.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the type locality.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF04DE0FFF07F9D5FA7455C0.taxon	diagnosis	Diagnosis. Aedeagal morphology places this species near A. curiosa Stark 1998, a Mesoamerican species, and A. cosnipata Stark & Sivec 1998, a species known from several Peruvian sites. The latter species bears a broad, dorsomedian, mesa-like structure on the aedeagus, and the median lobe on the aedeagal apex is wider than in the new species (see Stark & Sivec 1998, Figs. 53 - 55). Neither of these related species has the aedeagus strongly constricted proximal to the apical section, neither have the hooks blunt at the tips and neither shares the distinctive color pattern of A. pichincha.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF07DE09FEC3F9D4FB275376.taxon	description	(Figs. 29 - 31)	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF07DE09FEC3F9D4FB275376.taxon	materials_examined	Material examined. Ecuador: Napo Province, 2.78 km W Papallacta, Hwy E- 20, 3296 m, 25 January 2012, B. C. Kondratieff, B. Gill, 1 ♀ (CSUC). Adult habitus. General color dark brown to black. Head yellow, with dark brown patches extending forward of each ocellus toward central frons; lappets dark brown, posterolateral occiput and anteromedian frons pale brown (Fig. 29). Antennae and legs dark brown, femora paler in ventral half. Pronotum dark brown over lateral third, and bearing a thin, dark, diagonal line near anterolateral margin. Wing membrane dark brown to black, veins brown except pale R vein. Male. Unknown. Female. Forewing length 31 mm. Subgenital plate bilobed with truncate posterior margins; lobes separated by a shallow V-shaped notch. Sternum 9 bearing an obscure transverse sclerite, median sclerite covered by dark setae (Fig. 31). Egg. Somewhat thimble shaped with broadly rounded anterior pole almost as wide as collar end (Fig. 30). Collar small and button shaped. Chorionic surface without follicle cell impressions.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
250A8780EF07DE09FEC3F9D4FB275376.taxon	discussion	Comments. Females representing more than 30 Anacroneuria species have been described under temporary designations consisting of a two letter national code followed by a number. Ten have been proposed for Colombia (Stark et al. 1999; Zuniga & Stark, 2002; Zuniga et al. 2007), five for Guyana (Stark 1999; 2000), three for Paraguay (Stark & Baumann 2011), one for Peru (Stark & Sivec 1998), and twelve for Venezuela (Stark 1995; Maldonado et al. 2002). Several additional species have been proposed as Anacroneuria sp., or as Anacroneuria sp. A, etc., without a prefix; six of these (sp. A, B, C, D, E, F) were proposed by Harper (1992) for the Panamanian fauna. Only one of these unassociated females, A. CO- 5 (FWL 24 - 26 mm), has a forewing length approaching that of this species; both species also have the subgenital plate bilobed, however the former species is quite pale in comparison to A. EC- 1. Among large Anacroneuria species where sufficient specimens are known to generate a significant sample size [e. g. A. magnirufa Jewett 1958 or A. plutonis (Banks 1914)], male forewing length is approximately 75 % of female forewing length (Stark 1998). If this is applied to the current female, the male could be expected to display a forewing length of about 23 mm. The site (Fig. 35) where this female was collected is a high gradient stream characterized by a series of large waterfalls and short stretches of riffles. The channel width is 1.63 m and average depth at the time of collection was 0.12 m. Discharge was 31.3 L s- 1 at the time of the collection, and canopy cover was about 47 %. Vegetation is typical of high altitude Andean cloud forest; however, it has been disturbed by adjacent anthropogenic activities. This stream is one of the few in the Papallacta area where larvae of the gripopterygid Claudioperla cf. tigrina (Klapálek) were found in oxygenated areas with smaller substrate in large numbers.	en	Stark, Bill P., Kondratieff, Boris C., Gill, Brian (2012): New Species And Records Of Anacroneuria (Plecoptera: Perlidae) From Ecuador And Paraguay. Illiesia 8 (6): 78-93, DOI: http://doi.org/10.5281/zenodo.4760778
