taxonID	type	description	language	source
03A621374903FF96FC04EFC6FE60357E.taxon	discussion	Zwick (1986 a) recognized 30 Bornean Neoperla species and also made several informal designations based on unassociated female and larval material. Subsequently Stark & Sivec (2007) described N. darlingi from West Kalimantan, and provided a few new records of N. edmundsi and N. theobromae from East Kalimantan.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF96FE9BE8B7FF1F375D.taxon	materials_examined	Material examined. Brunei Darussalam: Temburong District, Sungai Belalong, Kuala Belalong Field Studies Centre, 04.54822 ° N, 115.15823 ° E, ALS B 43 ‐ 08, 11 ‐ 16 July 2008, A. L. Sheldon, 1 ♂ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF96FE9BE8B7FF1F375D.taxon	discussion	Remarks. This species was previously known from a single male collected in “ British North Borneo ” in 1959.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF96FEBFEA54FB943083.taxon	description	(Figs. 1 ‐ 5)	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF96FEBFEA54FB943083.taxon	materials_examined	Material examined. Holotype ♂ and 2 ♂ paratypes from Malaysia, Sarawak, Lanjak Entiman, 1.39 ° N, 112.16 ° E, 23 June 2008, U. Grafe (holotype USNM, paratypes BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF96FEBFEA54FB943083.taxon	description	Adult habitus. General color pale yellow brown. Head pale except for obscure slightly darker region on frons and over ocelli (Fig. 1). Pronotum pale brown with scattered darker rugosities. Femora pale brown, tibiae similar but with dark brown band at knee. Wing membrane pale amber, veins brown. Male. Forewing length 9.5 mm. Process of tergum 7 subtriangular with projecting sensilla basiconica around apex and on lateral margins (Fig. 2). Median sclerite of tergum 8 slightly rounded from lateral aspect and armed with a median row, and a few scattered sensilla basiconica. Tergum 9 with a pair of low humps armed with a sparse patch of ca. 7 ‐ 8 sensilla basiconica; area between humps partially sclerotized and bearing a few smaller sensilla basiconica. Hemiterga typical, anterior finger lobes slender, straight and strongly narrowed at apex. Aedeagal tube slender and sclerotized except for a partial, near apical, membranous zone armed with strong apically directed spines (Figs. 3 ‐ 5); apex of tube gradually curved ventrad. Aedeagal sac about as long as tube and armed along most of dorsal margin with scattered strong spines; apex of sac with a patch of smaller, densely packed spines and ventral margin with a linear cluster of subapical spines. Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF96FEBFEA54FB943083.taxon	etymology	Etymology. The patronym honors Professor Ulmar Grafe, collector of the type series, for his gracious assistance in this study.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF96FEBFEA54FB943083.taxon	diagnosis	Diagnosis. Neoperla grafei is a member of the N. parva species group recognized by Zwick (1986 a). It differs from other Bornean members of the complex (N. edmundsi Stark, N. parva Banks, N. starki Zwick) in details of aedeagal armature and tube shape. The new species is the only known member of the group with a strongly curved apex to the aedeagus, and it is also the only one to have the membranous spiny area of the tube set near the apex and extending as a partial ring across the dorsum; other species in the group have the spiny area distinctly separated into a pair of lateral patches located near midlength of the tube. This complex might be endemic to Borneo but Zwick (1986 a) suggests N. triangulata Kawai, a Sri Lankan species, is a possible member of the group. The holotype and paratypes of this species were collected with two males each of N. parva and N. rougemonti, and several females which do not appear to be any of these species.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF95FC3FEDE8FBCA327F.taxon	materials_examined	Material examined. Brunei Darussalam: Temburong District, Sungai Belalong, Kuala Belalong Field Studies Centre, 04.54822 ° N, 115.15823 ° E, ALS B 36 ‐ 08, 27 June 2008, A. L. Sheldon, 14 ♂, 44 ♀ (BPS, PMSL). Same site, ALS B 20 ‐ 08, 18 June 2008, A. L. Sheldon, 2 ♀ (BPS). Same site, ALS B 21 ‐ 08, 18 June 2008, A. L. Sheldon, 6 ♂, 4 ♀ (BPS). Same site, ALS B 29 ‐ 08, 23 June 2008, A. L. Sheldon, 3 ♂, 17 ♀ (BPS). Same site, ALS B 39 ‐ 08, 3 July 2008, A. L. Sheldon, 5 ♂ (BPS). Same site, ALS B 44 ‐ 08, 17 July 2008, A. L. Sheldon, 12 ♂, 5 ♀ (PMSL). Same site, ALS B 45 ‐ 08, 21 ‐ 23 July 2008, A. L. Sheldon, 18 ♂ (BPS). Temburong District, Sungai Esu, ALS B 30 ‐ 08, 28 June 2008, A. L. Sheldon, 1 ♀ (BPS). Temburong District, Sungai Belalong, Sungai Esu mouth, 04.453509 ° N, 115.16212 ° E, ALS B 37 ‐ 08, 4 July 2008, A. L. Sheldon, 2 ♀ (BPS). Malaysia: Sarawak: Lanjak Entiman, 1.39 ° N, 112.16 ° E, 23 June 2008, U. Grafe, 2 ♂ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374902FF95FC3FEDE8FBCA327F.taxon	discussion	Remarks. This species was previously known from two Bornean sites and five total specimens (Zwick 1986 a). These records include the first reports from Brunei Darussalam.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374901FF94FC1DEFBCFE8F3590.taxon	materials_examined	Material examined. Malaysia: Sarawak: Lanjak Entiman, 1.39 ° N, 112.16 ° E, 23 June 2008, U. Grafe, 2 ♂ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374901FF94FC1DEFBCFE8F3590.taxon	discussion	Remarks. This species was previously known from 11 specimens collected at two sites in Sarawak in 1932 and 1978.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374900FF93FEB0E89CFDCA370B.taxon	description	(Figs. 6 ‐ 10)	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374900FF93FEB0E89CFDCA370B.taxon	materials_examined	Material examined. Brunei Darussalam: Temburong District, Sungai Belalong, Kuala Belalong Field Studies Centre, 04.54822 ° N, 115.15823 ° E, ALS B 36 ‐ 08, A. L. Sheldon, 2 ♂ (BPS). Same site, ALS B 29 ‐ 08, 23 June 2008, A. L. Sheldon, 1 ♂, 1 ♀ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374900FF93FEB0E89CFDCA370B.taxon	description	Adult habitus. General color yellow brown. Head mostly pale but with brown arcuate mark behind ocelli and area between ocelli dark; M‐ line area and anteromedian area of frons slightly darker than background pigment (Fig. 6). Basal antennal segments pale brown, flagellum darker. Femora pale, tibiae darker; wing membrane pale amber, veins brown. Female. Forewing length 10.5 mm. Posterior margin of sternum 8 unmodified (Fig. 7). Receptacular stalk Cshaped and almost completely armed internally with closely packed brown scales (Fig. 8); vagina slightly constricted near midlength. Egg. Length from tip of lid to rim of anchor pole, ca. 225.0 μm; diameter, ca. 147.5 μm. Most of egg body covered by narrow, spiral striae (ca. 18 in lateral view) separated by narrow, punctuate sulci (Fig. 9). Base of anchor arises from center of convex, impunctate plate covering anchor pole; margins of plate scalloped with small projecting teeth over base of each striation (Fig. 10). Lid punctate with tiny aeropyles over entire surface.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374900FF93FEB0E89CFDCA370B.taxon	discussion	Remarks. Zwick (1986 a) described this species from five males taken at two localities in Sabah. These are the first specimens taken in Brunei Darussalam. The female association is based on a common pattern of brown pigment behind the ocelli. Eggs are similar to those of N. rougemonti (Zwick 1986 a) but have spiral striae rather than straight ones.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374907FF93FE99EA19FE0031B2.taxon	materials_examined	Material examined. Brunei Darussalam: Temburong District, Sungai Belalong, Kuala Belalong Field Studies Centre, 04.54822 ° N, 115.15823 ° E, ALS B 11 ‐ 08, 11 June 2008, A. L. Sheldon, 1 ♂ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374907FF93FE99EA19FE0031B2.taxon	discussion	Remarks. Zwick (1986 a) found this to be a variable species based on a few males collected at several Bornean sites. The aedeagus of the male listed above is very similar to Fig. 56 in Zwick (1986 a) but the median process of tergum 7 more closely resembles that of N. multilobata Zwick. This is the first record from Brunei Darussalam.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374907FF93FE95ECF1FAD13592.taxon	materials_examined	Material examined. Brunei Darussalam: Temburong District, Sungai Belalong, Kuala Belalong Field Studies Centre, 04.54822 ° N, 115.15823 ° E, ALS B 36 ‐ 08, 27 June 2008, A. L. Sheldon, 1 ♂ (BPS). Same site, ALS B 45 ‐ 08, 21 ‐ 23 July 2008, A. L. Sheldon, 2 ♂ (BPS). Same site, ALS B 21 ‐ 08, 18 June 2008, A. L. Sheldon, 1 ♂ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374907FF93FE95ECF1FAD13592.taxon	discussion	Remarks. This species was previously known from three sites in Sabah and one each in East Kalimantan and Sarawak (Stark & Sivec 2007; Zwick 1986 a). This is the first report from Brunei Darussalam.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374907FF92FC03EDA6FA4234E0.taxon	discussion	Zwick (1982, 1986 b) gave formal recognition to seven Bornean Phanoperla species and left one potentially distinct form (P. species cf maculata) without resolution. Stark & Sivec (2007) recognized two additional species (P. flabellare, P. tuberosa) and also gave new records for P. flaveola (Klapálek) and P. pumilio (Klapálek) from East Kalimantan. The following provisional key for males will assist in recognizing the 11 nominal Bornean species.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374906FF92FC49E80CFC23369F.taxon	materials_examined	Material examined. Malaysia: Sarawak, Lanjak Entiman, 1.39 ° N, 112.16 ° E, 23 June 2008, U. Grafe, 1 ♂ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374906FF92FC49E80CFC23369F.taxon	discussion	Remarks. The aedeagus of this specimen is very similar to the illustration (Fig. 15) in Zwick (1986 b) which shows a “ Phanoperla bakeri x anomala hybrid ”. Larger samples are needed in order to evaluate this hypothesis.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374906FF90FC18EB96FD2D34E0.taxon	description	(Figs. 11 ‐ 12)	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374906FF90FC18EB96FD2D34E0.taxon	materials_examined	Material examined. Holotype ♂ from Brunei Darussalam, TemburongDistrict, SungaiBelalong, KualaBelalongField Studies Centre, 04.54822 ° N, 115.15823 ° E, ALS B 43 ‐ 08, 11 ‐ 16 July 2008, A. L. Sheldon (USNM).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374906FF90FC18EB96FD2D34E0.taxon	description	Adult habitus. Body color pale white in alcohol. Head without distinctive pigment pattern; ocelli almost touching. Antennal bases pale but flagellum beyond basal segments, dark brown. Pronotum with pale brown rugosities and darker marginal and median sutures. Tibiae about as wide as femora and slightly darker. Wings pale with pale yellow veins. Male. Forewing length 7.5 mm. Tergum 8 unmodified, tergum 9 with a pair of lateral sensilla basiconica patches and a median patch of fine, short setae (Fig. 11). Hemiterga short and relatively wide with rounded tips in dorsal aspect; apices hooked downward in lateral aspect and bearing a few enlarged sensilla basiconica; notch separating basal callus narrow and deep. Aedeagal tube shortandplump with a fewfine setalspineson bulb; apex swollen and heavily armed with spines (Fig. 12); Largest and most prominent armature consists of a subapical ring of ca. 22 black spines and a pair of star‐ like, lateral clusters. Additional small, triangular spines occur over a broad zone, proximal to star clusters and on apical zone distal to spine ring. Frontal area below spine ring, bare and produced into a membranous nose‐ like process. Dorsal sclerite slender and poorly sclerotized. Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374906FF90FC18EB96FD2D34E0.taxon	etymology	Etymology. The species name refers to the star pattern of large spines on the aedeagus sac.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374906FF90FC18EB96FD2D34E0.taxon	diagnosis	Diagnosis. The aedeagus and external genitalia of this species is generally similar to that of P. flaveola (Zwick 1982) and it appears to be a member of that group. These structures are also generally similar to, or perhaps the same as, Phanoperla cf. maculata of Zwick (1986 b). That designation was based on a pair of Bornean specimens from the 1932 Oxford University Expedition studied by Zwick (1986 b), but whose status was not formally defined. The present specimen has the same general pattern of spines on the aedeagus, however there are fewer large lateral spines and they are arranged in a more distinct star pattern than in Zwick’s specimens. In addition, fine triangular spines are shown by Zwick (1986 b) along the frontal zone, and no nose shaped process is shown for his specimens. The new species is also distinguished from P. flaveola by the star cluster of lateral aedeagal spines. The lateral grouping of that species forms a linear group of 5 ‐ 6 large spines, and in addition, the dorsal aedeagal sclerite is much larger for P. flaveola than for the new species.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374904FF90FE9BE80CFB05357E.taxon	description	(Figs. 13 ‐ 14)	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374904FF90FE9BE80CFB05357E.taxon	materials_examined	Material examined. Holotype ♂ from Brunei Darussalam, Temburong District, Sungai Belalong, Kuala Belalong Field Studies Centre, 04.54822 ° N, 115.15823 ° E, ALS B 39 ‐ 08, 3 July 2008, A. L. Sheldon (USNM). Paratypes. Same site but ALS B 24 ‐ 08, 18 June 2008, A. L. Sheldon, 1 ♂ (BPS). Same site but ALS B 36 ‐ 08, 27 June 2008, A. L. Sheldon, 1 ♂ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374904FF90FE9BE80CFB05357E.taxon	description	Adult habitus. Body color pale yellow brown. Head yellow brown without pattern; ocelli almost touch. Basal 4 ‐ 5 antennal segments pale, rest of flagellum brown. Pronotum pale with darker rugosities and dark brown median and marginal sutures. Femora pale, tibiae brown. Wings pale, veins amber. Male. Forewing length 7.5 mm. Tergum 8 unmodified, tergum 9 with lateral and mesal sensilla basiconica patches united, or almost so (Fig. 13). Hemiterga with attenuated tips; basal callus offset from apical section of hemiterga by broad, shallow emargination. Aedeagal tube short and plump with a few fine setal spines on bulb; apex of tube bearing a pair of ventromedian lobes and a pair of dorsolateral lobes (Fig. 14); apical half of tube, including lobes armed with fine triangular spines. Aedeagal sac bearing lateral groups of 6 ‐ 7 large black spines in fan shaped clusters on either side of a cylindrical lobe densely armed with black, scale‐ like spines; dorsad to large fan shaped clusters, a small membranous lobe, armed with brown, triangular spines occurs on each side of sac. Dorsal sclerite spear shaped but apex curved dorsad over base of sac. Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374904FF90FE9BE80CFB05357E.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the type locality of Sungai Belalong and the Kuala Belalong Field Studies Centre.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374904FF90FE9BE80CFB05357E.taxon	diagnosis	Diagnosis. The aedeagal features of this species are similar to both P. flabellare Stark & Sivec and to P. anomala (Banks) (Stark & Sivec 2007; Zwick 1982). It differs from the former in bearing two pairs of prominent, membranous, spine‐ covered lobes ventroapically and dorsolaterally on the tube. In addition, a small spiny lobe occurs dorsad to the large fan shaped clusters of spines on the sac. The latter species shares the ventroapical spiny lobes on the aedeagal tube but the sac is armed with an almost complete ring of large black spines.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374904FF90FC6DE8BDFC31304F.taxon	materials_examined	Material examined. Brunei Darussalam, Temburong District, Sungai Belalong, Kuala Belalong Field Studies Centre, 04.54822 ° N, 115.15823 ° E, 21 July 2008, ALS B 45 ‐ 08, A. L. Sheldon, 1 ♂ (BPS). Same site, 27 June 2008, ALS B 36 ‐ 08, A. L. Sheldon, 1 ♂ (BPS).	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
03A621374904FF90FC6DE8BDFC31304F.taxon	discussion	Remarks. The surviving type series of this species includes a male and two females from Sebroang, Borneo (Zwick 1982) and a second male and five female specimens were reported by Stark & Sivec (2007) from East Kalimantan. These males represent the first record of the species from Brunei Darussalam.	en	Stark, Bill P., Sheldon, Andrew L. (2009): Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species. Illiesia 5 (2): 11-19, DOI: http://doi.org/10.5281/zenodo.4759022
