taxonID	type	description	language	source
0399AB12FFBBFFCFE91150695246FBC2.taxon	description	(Figs. 1 ‐ 5)	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBBFFCFE91150695246FBC2.taxon	materials_examined	Material examined. Holotype ♂, paratype ♂ and ♀ CHILE, Malleco, Nahuelbuta, Los Gringos, 37 ° 48 ʹ S; 73 ° 01 ʹ W, 15 ‐ 17 December 1993, CMF & OSF (USNM). Additional paratypes: 1 ♂, 1 ♀, CHILE, Malleco, Pino Hachado, 1480 m, 38 ° 39 ʹ S; 70 ° 58 ʹ W, 8 ‐ 9 January 1994 CMF & OSF; 2 ♂, Chile, Cautin, Conquillio, 1200 m, 4 ‐ 8 February 1988, L. E. Peña.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBBFFCFE91150695246FBC2.taxon	description	Dimensions. Male: body length 6.0 ‐ 7.0, forewing 7.5 ‐ 8.0; antenna 4.5. Female: body length 8.0; forewing 8.0 ‐ 9.0; antenna 5.0. Adult. Small slender light brown animals. Pronotum rectangular, with angles rounded. Head a little paler along sutures, muscle attachments on head and thorax a little darker, no prominent pattern. Ocelli present. Antenna brown, long, slender, individual segments distinctly expanded apically. Palps short, brown. Row of setae runs across apex of lacinia, in front of broad terminal teeth (see Fig. 68). Legs slender, essentially brown, base of femur a little paler, especially in front leg. Front and middle tibiae paler towards apex, hind tibiae with base to subgenual organ dark brown, remainder yellowish. Tibial spurs present. Wings (Fig. 1): Venation typical of the genus except for a very short fork of Rs without crossvein. The hind wing of specimens from Los Gringos (including the holotype) has the posterior branch of M coalesced with Cu 1 for a short distance. In other specimens the veins are separate but connected by a short crossvein. Veins pale brown and membrane with light brown tinge. Forewing with a darker tint around crossveins and in some specimens a purplish pigment from C to M but without speckles in cells. Abdomen sexually dimorphic. Cerci yellowish, short, downcurved with 8 ‐ 10 segments in males and 9 ‐ 11 in females. Pilosity inconspicuous. Male genitalia. (Figs. 2 ‐ 4): Abdomen brownish or purplish, only slightly sclerotised. Each tergite with an anterior sclerite band that is medially divided on anterior tergites but entire on tergites 9 ‐ 10, although it is deeply incised from behind on segment 9. Sternites incompletely sclerotised, segments 2 ‐ 7 with widely separate narrow transverse anterior sclerites plus a continuous sclerite band across posterior third of segment; sternite 8 completely sclerotised. Subgenital plate a broadly rounded transversely oval sclerite almost truncate posteriorly. Tergite 10 well sclerotised, except for a small notch in anterior margin and a median elongate pale band in front of a sharply pointed downcurved posterior sclerite (McLellan 1971). No epiproct. Paraprocts laterally dark brown with pale tips. The long slender sclerotised struts in side view are regularly curved and apically spatulate. Dorsoventrally the paraprocts have a narrow membranous triangular base that tapers until the sclerotised struts remain with their flat apices sometimes against each other. Penis structure not studied. Female genitalia. (Fig. 5): Abdomen pale, soft, dorsally unsclerotised except tergite 10, which is rounded apically. Sternites resemble those in males but anterior sclerites narrower and on sternite 7 posterior band shaped sclerite split into separate lateral sclerites that are slightly raised. Sternite 8 forming a sclerotised subgenital plate covering the posterior segments. Sides of subgenital plate parallel to slightly diverging posteriorly; distal margin with a broad, shallow excision. Subanal lobes triangular. The posterior sclerites on sternite 7, subgenital plate and ventrolateral face of the subanal lobes with a thick coat of stout curved colourless hairs which form a dense brush along the posterior margin of the subgenital plate. Larva. Unknown.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBBFFCFE91150695246FBC2.taxon	discussion	Remarks. Sternite 9 of the female paratype is completely telescoped under sternite 8. The generic name describes the hooked (Latin: uncus, hook; cauda, tail) caudal end of the male and the pirate ‐ like hook of tergite 10 posterior sclerite of the male suggested the specific name. Affinities. The new taxon is a member of Gripopteryginae in the sense of McLellan (1977). The insect has no striking autapomorphies but at the same time fits into none of the existing genera. Analysis of its affinities is hampered by lack of its larva. Therefore, generic rank as well as placement are somewhat provisional. The known apomorphic characters of Uncicauda suggest relations with some of the smaller genera which Illies (1963) classified as Paragripopteryginae. However, all of the apomorphies in question represent reductions that may have occurred repeatedly and independently, especially shortening of the fork of Rs and of the cerci. Even loss of the male epiproct, although very unusual and known only among South American Gripopterygidae, has occurred independently. Within Gripopteryx which, by its bizarre larvae, is undoubtedly monophyletic several species have very small epiprocts, and G. serrensis Froehlich lacks it completely (Froehlich 1993). The remaining genera without male epiproct (Paragripopteryx, Tupiperla, Guaranyperla (all NE South American) and Potamoperla (Andean )) have a double point of male tergite 10 which is unique in the subfamily and suggests these genera represent a monophylum. We believe the new genus Uncicauda with a sharp single point of tergite 10 may be an early member or the sister taxon of this small group of genera.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBDFFCCE93357CD571AFD03.taxon	description	(Figs. 6 ‐ 17, 71)	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBDFFCCE93357CD571AFD03.taxon	materials_examined	Material examined. Holotype ♂, Chile, Arauco, San Alfonso above Caramavida, 16 ‐ 17 October 1969, OSF & GB (USNM). Paratypes: ♂ and ♀ and 3 paratype larvae, Chile, Malleco, Pino Hachado, 1480 m, 38 ° 39 ʹ S; 70 ° 58 ʹ W, 8 ‐ 9 January 1994, CMF & OSF (USNM); 1 m (with everted conical penis), 1 ♀, Argentina, Neuquen, Lago Lolog, 40 ° 02 ʹ S; 71 ° 24 ʹ W, Quebrada, 19 km N of San. Martin del Los Andes, 1 ‐ 3 January 1994, CMF, OSF; 1 ♂, Chile, Biobio / Malleco, Volcan Callaquen, 5 ‐ 7 February 1959, LEP. 1 ♂, Argentina, Neuquen, Rio Litran, 5 km. N. Lago Alumine 1 March 1978 C. MF & OSF; 2?, Chile, Malleco, Nahuelbuta, bog outlet, 37 ° 47 ʹ S; 73 ° 01 ʹ W, 16 ‐ 19 December 1993, Malaise trap, CMF & OSF; 1 ♀, Chile, Palena, Rio Ventisquero, 16 km S Pto. Cardenas, 100 m, 23 ‐ 24 January 1987, CMF & OSF; 1 ♀, Chile, Nuble, El Purgatorio, 3 March 1968, OSF, MLP, 1 ♀, Argentina, Neuquen, brooklet above Pucara, 1 February 1974, OSF; 1 ♂, 1 ♀, Chile, Nuble, Las Cabras, Cordillera Chillan, 8 ‐ 15 February 1959, LEP. All paratypes (USNM). Additional material: 8 LA, Chile, Nuble, Las Trancas, 1300 m, 21 km E Recinto, high waterfall ex moss, 17 January 1979, MD, DD, BA; 9 LA (1 small), Chile, Nuble, Las Trancas, 1300 m, 21 km E Recinto, high waterfall, ex moss, 17 January 1979, MD & DD & BA; 3 LA, Argentina, Neuquen, cascades, 6 km N Lago Alumine 1100 m, 3 February 1987, CMF & OSF, 1 LA, Chile, Osorno, stream near the Carabino Post in Antillanca, 12 December 1972, MLP; 1 LA, Argentina, Neuquen, Ao Punameco, NW Lago Alumine, 1 March 1978, CMF & OSF; 1 LA, Argentina, Neuquen, Ao Punameco, NW Lago Alumine, 1 March 1978, CMF & OSF.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBDFFCCE93357CD571AFD03.taxon	description	Dimensions. Male: body length 7.0; forewing 8.0; antenna 5.0. Female: body length 9.0; forewing 9.5; antenna 5.0. Larva (last instar female): body length 8.5; antenna 3.0; cercus 1.7. Adult. A medium sized stonefly. Head strongly deflexed, narrowed towards down ‐ turned mouthparts. Epicranial suture deeply grooved. Area between 3 distinct ocelli appears raised. Apical segment of maxillary palpus 3 X length of penultimate segment. Pronotum rectangular, twice as wide as long, with raised anterior and posterior margins and a paramedian callus on disk. Brown to dark brown with pale stripe along midline and around perimeter. Legs without tibial spurs. Both wings with Rs and Cu 1 forks and 6 anal veins in hind wing (Fig. 6). Cerci downcurved, with about 10 ‐ 12 segments, basal one long. Male genitalia (Figs. 7 ‐ 9). Sternite 9 produced into wide, shallowly bilobed subgenital plate. Tergite 10 central sclerite curved downwards at half length to fit neatly between arms of epiproct base. Medially on rounded hind margin of tergite 10 is a minute cone shaped posterior sclerite. Arms of epiproct curve down and posteriorly, then join and curve upwards and posteriorly, terminating in a downcurved sharp tip. Paraprocts large, about as long as cerci, broadly falcate, inner face from base to about half length forms a triangular membranous sheet. Membranous penis inside sternite 9, visible only in cleared specimens. The everted penis appears relatively dark, apparently through tissue pushed into it by body fluid pressure. It is a large pointed cone slightly curved up between the paraproct bases, lying below the epiproct hook and is partly divided longitudinally by a lateral fold (Fig. 9). A tiny gonopore apparently at tip of slightly longer anterior part. A small triangular wing embracing base of paraproct observed only on the left side of body of the single specimen with everted penis. Female genitalia. (Fig. 10). Sternite 8 with well developed subgenital plate, with 2 lateral dark bands, tapers to a rounded or double pointed tip extending on to sternite 9. Subanal lobes slender, pointed. Cerci short, of 12 segments. Tip of tergite 10 produced, rounded. Larva. (Figs. 11 ‐ 17). Short, stout, hairless and dusty brown. Surface matt because of numerous shallow pupillate punctures. Head curved down, with shallow central depression between ecdysial suture and indistinct ocelli. Head capsule much narrowed from wide base to narrow front i. e. wedge shaped, genae in front view visible as flat surfaces below and in front of eyes and antennal foramen. Clypeus and labrum narrow. Mandible base (Fig. 13, stippled) prominently exposed. Mandibles (Fig. 17) exceptionally flat, their long apices blade ‐ like, the sharp serrate edges of the pair interdigitate. Base of mandible with distinct setal rosette, mola far from mandible base. Left mola serrate anteriorly, right mola concave. Maxillae (Fig. 16) long, with strongly elbowed base often quite visible in dorsal view; apparently, the maxilla can reach far forward. Maxillary palpus 4 ‐ segmented, apex of last segment soft, sometimes folded inward. Galea and lacinia unusually slender. Galea with apical rosette of sensilla, lacinia with few narrow well sclerotised apical teeth and a series of translucent fairly wide subterminal setae. Glossae shorter than simple paraglossae, with apical hair rosette (Fig. 15). 2 nd segment of labial palpus medially swollen and strongly sclerotised. Hypopharynx inconspicuous and smooth. 17 Pronotum subrectangular, wider than long; hind margins of mesonotum and metanotum convex. Thoracic tergites with relief of grooves and ridges whose outlines are difficult to define precisely because of rough matt surface and best seen in side view (Fig. 12). A raised cone near base of wingpad on meso ‐ and metanotum. Femora short and stout, dorsally crested, a raised spinulate hump mid ‐ dorsally. Tibia lacks spurs. Tarsal segments 1 and 2 both extremely short, segment 2 dorsally divided by narrow base of long 3 rd segment, ventral face of 3 rd tarsal segment spinose (Fig. 14). Tergites and sternites of abdominal segments 1 and 2 divided, those of 3 incompletely divided, with anterior notch. Abdominal tergite 10 produced and pointed, more so in males. Subanal lobes narrow, long, triangular. Cerci short, about 18 segments. Retractable anal gill rosette of many filaments with subanal lobes able to close over it.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBDFFCCE93357CD571AFD03.taxon	discussion	Remarks. The genus is named for the first locality from which the species was recorded. This species is dedicated to Carol and Oliver Flint who not only collected the types but also were responsible for collecting a sizeable portion of all the material we examined. The male epiproct, paraprocts and the bilobed penis are characteristic of the Australian Dinotoperlinae and the wing venation with forks in Rs and Cu 1 is characteristic of Trinotoperla and Illiesoperla in that subfamily. Lack of tibial spurs is uncommon but despite this the other characters point more to the inclusion of the genus in Dinotoperlinae rather than Gripopteryginae or Antarctoperlinae. A young wingless larva of this species was illustrated and briefly described by Illies (1963), as “ Larventyp X (Teutoperla n. gen.?) ”. The general shape, the spinose, dorsally raised femora (Illies’ text refers to ‘ tibiae’ by mistake but his figure is correct), the pointed tergite 10, the long triangular paraprocts, the narrow maxilla and its 4 ‐ segmented palpi clearly indicate the conspecificity of his specimens with ours. Our interpretation is also confirmed by specimens in the Illies collection. The larva of this species was verified by dissection of a mature male larva. Illies (1963) found the larvae in moss at the bottom of streams or on rocks. The present material additionally suggests that the larvae dwell in or at least near waterfalls. The exceptional shape of head capsule and mouthparts suggested some special food, but the fore ‐ gut of a single penultimate instar larva (from Neuquen Cascades) was filled with fine particulate matter, including many small diatom valves, fungal hyphae and spores, fragments of macrophyte tissue with tracheae, fine mineral particles, but no recognisable animal remains. The larva appears to be a gatherer, possibly obtaining its food from special microsites.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBEFFCCEBA7518157A0F919.taxon	discussion	This subfamily is well supported, also by larval characters. These include the very rounded massive head with a frontal depression, ring ‐ shaped abdominal segments lacking a soft pleural area, the long parallel ‐ sided posteriorly extended segment X which covers the retracted gills and the paraprocts from above, and the modified cerci. The cerci are short and thin, the apical section is thread ‐ like. The terrestrial Megandiperla with its very short but thick conical cerci is the only exception. Larvae and adults lack tibial spurs. However, larvae of some other South American taxa also lack them (see the key). Additional characters characteristic of larvae of Antarctoperlinae but not shared by all of them are the thickened and often very pilose base of antennal flagellum, prominent or extended pronotal corners, paired dorsal processes on head, pronotum and / or abdomen, and camouflage of detritus containing fine sand firmly attached to body hairs. Unassociated Larvae In the Flint collection there are Antarctoperlinae larvae without associated adults. Several are last instar specimens but none is sufficiently mature to dissect the pharate adult. These larvae are described below, under informal designations.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBEFFC2EBB455EB57CCFB5A.taxon	description	(Figs. 18 ‐ 23)	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBEFFC2EBB455EB57CCFB5A.taxon	materials_examined	Material examined. 1 LA Chile, Malleco, Vegas Blancas, 27 km W Angol, 700 m, 17 January 1987, CMF & OSF; 1 LA Chile, Llanquihue Prov., a stream 15 km SW of Las Quemas 17 December 1972, MLP; 1 LA Chile, Osorno, Rio El Gringo Park, Puyehue, 1075 m, 13 December 1972, MLP; 2 female LA, Chile, Malleco, Vegas Blancas, 27 km W Angol, 700 m, 17 January 1987, CMF & OSF.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBEFFC2EBB455EB57CCFB5A.taxon	description	Dimensions. Larva (last instar): body length, 9.0 ‐ 10.0; antenna, 5.0 ‐ 7.0; cerci, 1.0. Larva. Pale brown, with a sand like matt surface; indistinct muscle attachments on head and thorax a darker brown; a brown ring across widest part of antennal flagellum, subgenual rings and apices of tibiae dark grey. Hairless, except as mentioned below. Head: Rounded, massive. Eyes large, no ocelli. Unpaired cephalic suture long, well visible, its fork distinct; depression in front of fork deep. Frons bulging over downwardly directed clypeus and labrum. Mandible (Fig. 19) with large sharp pointed teeth ventrally and a dorsal elongate oval spiny mola and a faint prosthecal hair brush but no other hairs; a transverse row of sensilla inward from ventral condyle. Maxilla (Fig. 20) with 5 ‐ segmented simple palp, basal segment very short, in ventral view not easily distinguished from palpiger. Galea simple, a few outwardly turned elongate sensilla at apex, medial face closely appressed to lacinia. Lacinia very broad at base, especially from lateral view, strongly tapered to narrow tip which has three very dark sharp slender teeth. Basally from teeth along mediodorsal edge are a series of pale setae with about five of the apical ones modified into large spike ‐ like structures. Antennae (Fig. 18) almost as long as body, scape large, and pedicel dark and basally strongly constricted. Flagellum distinctly enlarged basally with a dark band, segments on wide part very short and transverse, scarcely discernable, only distinguished by rings of curved short setae; distal segments narrow, almost twice longer than wide, hairless. Thorax. Dorsum flat and much wider than parts below. Laterally flanged with a sharp edge, along which there is a row of small sharp pale setae. Pronotum with all four angles produced into prominent angular lobes. Mesothoracic wing pads with parallel sides and free distal parts only about as long as wide. Metathoracic wing pads divergent and shorter. Legs simple, long, with inconspicuous pilosity. Outer face of tibiae with a series of 4 ‐ 7 long pale hairs along a narrow bare longitudinal space between two regular rows of very short setae. No tibial spurs, but a few stronger setae ventrally at apex. Tarsi (Fig. 21) ventrally with fine hair brushes, dorsally with some thin hairs. First segment longer than wide with narrow base, Segment 2 incomplete, a ventral lobe dorsally divided completely by insertion of segment 3; segment 3 longer than first two together. Abdomen formed of 10 complete segmental rings, each of the first nine medially strongly constricted, with fringe of setae arising from dark spots along bulging distal edge (Fig. 22). Most setae are short, truncate, of variable length but some long, extending into a soft pale filament which is probably broken in the truncate setae. Ventrally, the distal bulges are less prominent, setae are shorter, soft, pale, appearing woolly. Segment 10 (Fig. 23) forms a distally obliquely truncate cylinder and bears a regular fringe of short setae posteriorly along its upper edge. The ventrodistal opening of the segment is almost completely covered by the flat, hard subanal lobes from which rise a short hard apical spine. Cerci minute, pale and sparsely pilose; the longer one with seven segments but both are probably broken. The pale tips of numerous retracted gill filaments project between the subanal lobes.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFBEFFC2EBB455EB57CCFB5A.taxon	discussion	Remarks. The width and seeming shortness of the wing pads results from their flanged nature; the adult is probably fully winged. The mandibular mola resembles the “ Stachelmola ” as illustrated for Brachyptera, Wichard et al., 1995.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB0FFC0EBB457AA51AFF919.taxon	description	(Figs. 24 ‐ 29)	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB0FFC0EBB457AA51AFF919.taxon	materials_examined	Material examined. 2 LA, Argentina, Neuquen, Rio Litran, 5 km. N. Lago Alumine 1 March. 1978 CMF & OSF; 8 LA, Argentina, Neuquen, Ao. Purumeco, NW Lago Alumine 1 March 1978 CMF & OSF; 1 LA, Chile, Malleco, small stream 20 km E. Liucura, 1750 m, 4 February 1987, CMF & OSF, (February 97 in LFS); 1 LA, Argentina, Neuquen, small stream, 10 km S. Pino Hachado, 4. February 1987, CMF & OSF, (February 97 in LFS); 4 LA, Chile, Malleco Prov., Cordillera de las Raices, 37 km E Curacautin, 5 ‐ 6 February 1979, 1100 m, DD, MD & DA (97 in LFS); 39 LA, Chile, Malleco, small stream, 9 km W Lonquimay, 5 February 1987, CMF & OSF (3 remain as duplicates in LFS). 8 LA “ 9 / 12 o ” among unsorted material in coll. Illies, locality and date unknown.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB0FFC0EBB457AA51AFF919.taxon	description	Dimensions. Larva (last instar): body length 12.5 ‐ 13.0; antenna 7.0; cerci 2.5. Larva. Dark brown, muscle attachments on head and thorax darker. Scape and pedicel plus some flagellar segments (about segments 7 ‐ 12) darker brown than rest of antenna. Similarly, tip of first cercus segment and the next 6 ‐ 8 segments distinctly darker brown than rest of cercus. Clothed with short dark procumbent setae (Fig. 25) except side edges of thorax and caudal edges of abdominal segments which have setal fringes as described below. Head: Massive, dorsal face flat, occipital area strongly downcurved. Eyes large, no ocelli. Central depression shallow, indistinct. Unpaired cephalic suture long, branches initially diverging at right angle, then bent abruptly sidewards, meeting inner edge of eyes and curving forward alongside them, behind antennae. Frons descending stepwise to clypeus, labrum with large articulation membrane. Genae as high as the large antennal scape and cut off and rounded level with the scape. Mouthparts similar to Antarctoperlinae gen. sp. I. Antennae about half as long as body, scape large, pedicel dark, basally strongly constricted. Base of flagellum distinctly expanded, segments on wide part of flagellum very short and distinguished by rings of strong short setae whose tips curve back against flagellum; narrow distal segments almost twice longer than wide, hairs hardly noticeable. Thorax. Nota flanged, the edges with a fringe of prominent setae with truncate tips. Pronotum as wide as head, wider than long, almost rectangular with a short blunt extension of the anterior angles. Wingpads short and triangular. Legs. Short, hind legs shorter than the abdomen and clothed dorsally with a few dark setae. Tibiae triangular with a sharp outer edge. Tarsi with the usual short segment 1 and 2 and large claws. Abdomen short and wider medially, the segments are complete rings, their hind margins with a fringe of prominent dark setae. Hind margin of tergite 10 rounded. Subanal lobes each with a long pointed projection. Cerci short and threadlike, their extremities clothed with a ring of short setae.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB0FFC0EBB457AA51AFF919.taxon	discussion	Remarks. The short triangular wing pads probably indicate that some adults may be short winged. Both Antarctoperlinae gen. sp. I and II have a broad flanged thorax and long, almost straight paraproctal spines. The prominent pronotal corners and long setae of the abdominal hair fringes distinguish sp. I from sp. II. The latter has blunt corners of pronotum and short peg ‐ like setae in abdominal fringes.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB2FFC5E90055EB51F7FE71.taxon	description	(Figs. 30 ‐ 35)	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB2FFC5E90055EB51F7FE71.taxon	materials_examined	Material examined. 1 LA, Chile, Magal., Penin. Cordova Ba. Borja 7. October. 69 OSF; another larva from the same locality is additionally labelled “ pond ”; 2 LA (partly dissected, parts on slide), Chile, Magal., I. Desolacion, Pto. Churruca, 5 October 1969, OSF.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB2FFC5E90055EB51F7FE71.taxon	description	Dimensions. Larva (final instar): body length 12.0 ‐ 13.0; antenna 6.0; cerci 1.5. Larva. A moderately slender species, uniformly light brown and shining; the larva appears to be hairless under the dissecting microscope, except for colourless short hair fringes along the front edge of the clypeus and the rear edge of tergite 10. However, when pieces of integument are studied with a compound microscope and under transmitted light, the body is found to be covered by very fine, colourless hairs. Hairs on tergites are curved and strongly tapered towards the tip, being almost whip like on basal segments. Apices of hairs along distal segment margins are strongly curved inwards, towards the body. Floriform chloride cells on intersegmental membranes at basal abdominal segments numerous, each resembling a goblet with long basal stalk. Head massive, simple, with distinct shallow central depression, eyes small, and ocelli indistinct. Antennae seemingly bare, with large squarish scape and narrow bell ‐ shaped pedicle; basal third of flagellum indistinctly inflated. Genae normal, simple; frons gently bulging down to the short clypeus, labrum trapezoidal, with narrow base. Sclerotised mouthparts worn, details of molar structure no longer visible, lacinial apex apparently with only a few blunt massive scoop ‐ like apical teeth. Galea longer than lacinia, with apical group of small rod shaped sensillae. Palpus simple, segment 4 a little shorter than both segments 3 and 5. Labium normal, except terminal segment of palpi wide, distinctly ovoid, fairly densely pilose. Thorax. Pronotum a little wider than long, the anterior angles a little produced forward but barely outward. Rear angles pronounced but blunt, not produced. Meso ‐ and metanotum simple, with short wing pads. Legs short, stout, hind tarsi not reaching abdominal tip. Tibiae simple, no crests or keels. Tarsi with segment 2 short, dorsally completely divided by the basal part of distal segment. Abdominal segments ring ‐ shaped, a little constricted just forward of the middle of each so that the segment contour appears bell ‐ shaped. Two minute dark spots on tergites 2 ‐ 9 appear to be part of the adult pattern rather than larval marks; the narrowly triangular apophyses of the antecosta of tergite 10 faintly visible by transparency. Segment 10 large, with protruding rounded posterior edge; bases of cerci and paraprocts concealed inside last segment. Cerci of about 20 segments but very short, downcurved, the apical part extremely fine, narrow and translucent. Paraprocts with sclerotised, medially partly overlapped bases from which rises a sharply pointed upcurved process; in dorsal view, the two processes appear like the forceps of an earwig (Dermaptera) while in side view the basal two thirds of their ventral edge are slightly concave to nearly straight. Gill rosette large, many fine filaments but contracted in the specimens examined. 30 Adult. Not known; however, a few traits of the developing adult were visible by transparency or were dissected out from the larvae from Isola Desolación. The wings are about 3 mm long when artificially extended in hot lactic acid. Wings were insufficiently developed to recognize details of the much reduced venation; however, Rs and M are distinctly forked, crossveins are not visible. Both mature larvae are females; a pale median strip and darker paramedian lobes that probably extend backward in the emerged adult are visible by transparency inside sternite 8. Sternite 9 contains a deeply forwardly arched darker structure.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB7FFC5E900515C50E3F9C4.taxon	description	(Figs. 36 ‐ 42)	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB7FFC5E900515C50E3F9C4.taxon	materials_examined	Material examined. 3 LA, Prov. Linares, Catillo, 29 March 1975, CC (in coll. Zwick).	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB7FFC5E900515C50E3F9C4.taxon	description	Dimensions. Larva (last instar): body length 9.0; antenna 6.0; cercus 1.5. Larva. Slender, fully winged, long ‐ legged, larva. Pale matt brown, a narrow sharply delimited median longitudinal line on the abdomen. Antennae very long and slender with a darker ring in basal quarter and another just before mid ‐ length. Subgenual rings on tibiae dark brown. Pilosity not obvious, but very complex and varied at high magnification. Head not distinctive, dorsally a little concave but depression in front of occipital fork not deep; frons bulging over downwardly directed clypeus and labrum. Eyes large, distinct ocelli. Genae distinct, not as deep as width of antennal scape. Mandibles, maxillae and labium resembling Antarctoperlinae II, but hypopharynx simple, no distinct lobes. Antennae about 2 / 3 body length, structurally simple, no swollen or conspicuously setose parts, distal segments very thin and several times longer than wide, no distinct setae. Thorax. Pronotum with all four angles conspicuously horned laterally, the anterior horns curved posteriorly and the posterior ones curved anteriorly. Anterior and posterior margins convex, hind margin with rough short modified hairs. Raised anteromedially part of meso ‐ and metanotum with similar pilosity. Both wing pads divergent, well developed, adult obviously fully winged. Legs simple, long (apex of hind femur reaching to tergite VI), almost hairless, no keels, spines or tibial spurs. Tarsi similar to other Antarctoperlinae, but slender, first segment twice as long as wide. Abdominal segments ring ‐ shaped, middle and posterior ones strongly constricted medially, with dark spots along bulging distal edge; the setae in these insertion points are very short, club ‐ shaped to almost spatulate with the wide apices pale and transparent. Interspersed with the setae are a few long thin twisted or curled pale hairs which are best seen in slanting light and then give the larva an untidy appearance. On the tergal surfaces, in transmitted light, between the few pale club ‐ shaped setae are numerous minute thinly stalked floriform chloride cells which have only a few delicate cuticular rods surrounding the central vesicle (normally, there is a dense basket ‐ like arrangement of cuticular rods around it). Segment 10 with its distal dorsal edge regularly rounded and fringed with short setae, its ventro ‐ distal face covered by elongate triangular, prominently pointed, sclerotised subanal lobes. In side view, the ventral edge of the subanal lobe forms a straight line, from base to tip. Cerci directed postero ‐ laterally. They are basally light brown, regularly segmented and sparsely pilose but distally completely pale, indistinctly segmented and hairless. Gill rosette large, purplish.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB7FFC5E900515C50E3F9C4.taxon	discussion	Remarks. The male larva is insufficiently developed to show distinct secondary sexual characters, except for an indistinctly bilobed slight mediodistal extension of sternite 9. The dark dorsal line is most distinct and longest in the smallest larvae, but least developed in the largest specimen. However, this may be related to the condition of the specimens rather than being typical of given instars. By the shape of their wing pads, the three available larvae belong to the last three instars; their body proportions suggest wing development is in three steps, as in most Plecoptera (Zwick 2003). Slender shape and pronotal angles suggest this may be the larva of some Plegoperla species. However adult Plegoperla have forwardly directed anterior angles of pronotum and the short extension of the posterior angles is directed backward. Also, Plegoperla adults have very long antennal scapes and tibial spurs which are lacking in the Antarctoperlinae IV larva.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
0399AB12FFB7FFDEEB0755C95771FE02.taxon	discussion	Family diagnosis. Gripopterygidae are distinguished by a supra ‐ anal retractable rosette of unsegmented, often purplish gill filaments. Only Notoperla lacks gills but can be separated from the gill ‐ less South American family, Notonemouridae, by its large furcasternum widely separating the coxae and by furcal pits located far from the spina. In Notonemouridae, the furcasternum is very small, coxae are close together, and furcal pits are adjacent to the spina. 44 46 47 The larva of Andiperlodes is unknown. Taxa that may or may not be dirt covered are identified by their morphology, not mainly by the dirt cover There is little information on morphological change during larval development of South American Gripo pterygidae, which may be important (Figs 43 ‐ 47). Change may also concern changes in late instars that are not related to wing development, for example Antarctoperlinae sp. II and IV. The key is to last instar larvae.	en	McLellan, Ian D., Zwick, Peter (2007): New Species Of And Keys To South American Gripopterygidae (Plecoptera). Illiesia 3 (4): 20-42, DOI: http://doi.org/10.5281/zenodo.4758655
