taxonID	type	description	language	source
653F6C3CF32AFFACE33C77CC355FF873.taxon	discussion	Comments. No additional records are available for this species, but the tentative identification of an Ecuadorian specimen as A. aymara by Stark (2001) has subsequently been refuted in Zúñiga et al. (2006) when the specimen was placed as a paratype of A. chachis. The revised distribution is shown in Fig. 20.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32AFFACE15F72A3332FFBA2.taxon	materials_examined	Material examined. Colombia: Valle del Cauca Department, 6 miles W Cali, 20 March 1955, E. I. Schlinger, E. S. Ross, 1 ♂ (CAS).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32AFFACE15F72A3332FFBA2.taxon	discussion	Comments. The known collections of this species include the holotype male, and four male and four female specimens from the type locality in a protected area (Parque Nacional Farallones de Cali) with extensive riparian vegetation and clear, unpolluted water in Valle del Cauca Department, Colombia (Zúñiga & Stark 2002). Three additional males and one female specimen, collected in Risaralda and Nariño Departments in the Central Andes, seem to confirm the distribution of this species in the highlands between 2000 and 2600 m a. s. l. (Castillo et al. 2013; Zúñiga et al. 2007). The current known distribution is shown in Fig. 20.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32AFFAAE1E676D63425FBDB.taxon	description	(Figs. 1 - 5, 21)	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32AFFAAE1E676D63425FBDB.taxon	materials_examined	Material examined. Holotype ♂, Venezuela, Barinas State, 15 km W Altamira de Caceras, 1000 m, 30 December 1985, P. Kevarik, R. Jones (USNM). Paratype: Venezuela: Barinas State, 10 km southeast of Miri, 8 February 1970, 150 m, S. L. Wood, 1 ♂ (BYUC).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32AFFAAE1E676D63425FBDB.taxon	description	Adult habitus. General color brown with yellow to yellow-brown markings. Lappets dark brown, frons yellow-brown with slightly darker, diffuse-brown area forward of ocelli (Fig. 1). Pronotum brown over much of disc, but pale laterally and along median suture. Wings pale amber with darker veins; C and Sc pale amber. Dorsum of legs brown, but pale along ventral margins. Male. Forewing length 9 - 10 mm. Hammer thimble shaped, height greater than apical diameter (Fig. 2). Aedeagal apex truncate, slightly emarginate, simple, but bearing a ventral pair of small membranous lobes (Fig. 3), and a distinctive pair of dorsal keel lines which meet at midlength and curve outward nearer apex, approximating a Y-shape (Fig. 5). Dorsolateral margins along keel lines slant steeply to ventrolateral base, and apices of keel lines extend to midlength of apex in lateral aspect (Fig. 4) and form a slight marginal incision in subapical profile. Hooks subchelate (Fig. 3); apex of right hook of holotype (in ventral aspect) broken. Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32AFFAAE1E676D63425FBDB.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the Venezuelan state in which the holotype was collected.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32AFFAAE1E676D63425FBDB.taxon	diagnosis	Diagnosis. The dorsal keel of the new species is almost identical to that of A. toni Zúñiga & Stark 2002, but the aedeagus of that species has a slight subapical constriction causing the truncate apex to be wider than the subapical area. In A. barinas, the lateral subapical aedeagal margins narrow to a slightly emarginate apex (Figs. 3 - 5).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32AFFAAE1E676D63425FBDB.taxon	discussion	Comments. One of us (BPS) first studied the specimen now designated as holotype from the Monte L. Bean Life Science Museum, Brigham Young University, in 1998, and labelled it “ Anacroneuria n. sp. ” Recently, it was returned for additional study through the courtesy of R. W. Baumann, and a second specimen, collected by S. L. Wood from a nearby site was also included. Both sites are located in the Cordillera de Mérida, the highest mountain range in the Venezuelan Andes. The species is a member of the A. aymara complex (Maldonado et al. 2002; Zúñiga & Stark 2002; Zúñiga et al. 2006; Stark et al. 2012), which currently includes the species listed in Table 1. Distributions (by Department, Province or State) for these 12 species are shown in Figs. 20 - 23. The key below will assist in recognition of male specimens assigned to this complex.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32DFFABE32D70F136BEF997.taxon	materials_examined	Material examined. Colombia: [Putumayo Department], Santa Rosa de Sucumbios, Kofan Indian Village, Rio San Miguel, 400 m, 26 August- 9 September 1971, B. Malkin, 1 ♂ (CAS).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32DFFABE32D70F136BEF997.taxon	discussion	Comments. The holotype specimen, and the holotype for the synonym are both from Colombia, but this is the first new record of the species from Colombia since the original descriptions were published, and the first where the collection sites are clearly defined. Both holotypes were taken from the Rio Magdalena, however the precise locality along the approximate 1500 km of stream length is unknown. The “ Kofan Indian Village ” site for our specimen is located in the Amazon Natural Region. The San Miguel River is a tributary of the Putumayo River and together these streams form the southeastern border of Colombia with Ecuador and Peru.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32DFFABE37474C53232FCFF.taxon	materials_examined	Material examined. Venezuela: Lara State, Quebrada Los Frailes, 8.3 km from San Pedro, 9 ° 48.358 ’ N, 70 ° 05.297 ’ W, 1892 m, 23 June 2001, Holzenthal, Blahnik, Paprocki, Cressa, 3 ♂ (pinned, BPSC).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32DFFABE37474C53232FCFF.taxon	discussion	Comments. This species is reported from Colombia, Ecuador and five states in Venezuela (Maldonado et al. 2002; Stark 1995; 1999, 2001; Zúñiga et al. 2007). These are the first records from Lara State in western Venezuela. This region has an abundance of xerophitic vegetation, but the mountainous regions have remnants of cloud forests. Streams in this area flow into the Caribbean or into the Orinoco River drainage basin. The Colombian specimens are from the Region Natural Andina in the Cordillera Occidental and the Cauca	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32DFFABE19C7164327DF88D.taxon	materials_examined	Material examined. Venezuela: Falcón State, Quebrada El Charo, Cataratas, 10 ° 46.771 ’ N, 69 ° 12.173 ’ W, 425 m, 12 June 2001, Holzenthal, Blahnik, Paprocki, Cressa, 1 ♂, 1 ♀ (pinned, BPSC). Falcón State, Parque Nacional Cueva de la Quebrada del Toro, Quebrada del Toro, 10 ° 49.581 ’ N, 69 ° 07.990 ’ W, 530 m, 11 June 2001, Holzenthal, Blahnik, Paprocki, Cressa, 1 ♀ (pinned, BPSC). Lara State, Quebrada Los Frailes, 8.3 km from San Pedro, 09 ° 48.358 ’ N, 70 ° 05.297 ’ W, 1892 m, 23 June 2001, Holzenthal, Blahnik, Paprocki, Cressa, 1 ♀ (pinned, BPSC).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32DFFABE19C7164327DF88D.taxon	discussion	Comments. Anacroneuria bifasciata was previously reported from three states (Aragua, Carabobo, Lara) and the Distrito Federal of Venezuela, in addition to two sites in Colombia (Froehlich 2010). Only one of the Colombian sites is specifically known, and it lies within Antioquia Department in the Región Natural Caribe and the Cauca River drainage basin. The specimens listed above include the first reported from Falcón State, which is located in northwestern Venezuela in the Caribbean region. Parque Nacional Cueva de la Quebrada del Toro is the site of a noted underground stream.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32EFFA9E37877EB3699FEA5.taxon	description	(Figs. 6 - 8)	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32EFFA9E37877EB3699FEA5.taxon	materials_examined	Material examined. Ecuador: Esmeraldas Province, Cantón San Lorenzo, Río Lita, 1830 ’, 00 ° 51.64 ’ N, 78 ° 27.16 ’ W, 2 December 2008, S. M. Clark, R. C. Mower, 2 ♂, 1 ♀ (BYUC).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32EFFA9E37877EB3699FEA5.taxon	discussion	Comments. This species was recently recorded by Stark et al. (2012) from two male specimens collected together with the additional males (listed above) and first female specimen recorded here and described below. The adult head and pronotum are also illustrated for the first time (Fig. 6). The Rio Lita collection site is located in the Northwestern Ecuadorian Pacific Zone, which forms part of the Choco Biogeographic Region, an area known for its hot, humid climate.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32EFFA9E37877EB3699FEA5.taxon	description	Female. Forewing length 17 mm. Subgenital plate pale, hairy and bilobed, mesal notch relatively narrow and shallow (Fig. 8). Posterior transverse sclerite on sternum 9 pale, mesal field of sternum 9 uniformly covered with moderately long pale setae. Egg. Length ca. 0.32 mm, greatest width ca. 0.19 mm. Spindle shaped with button-like collar (Fig. 7).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32FFFA9E32C73D234A8FC00.taxon	materials_examined	Material examined. Colombia: Caldas Department: 20 miles W Manizales, 16 March 1955, E. I. Schlinger, E. S. Ross, 1 ♂ (CAS). Tolima Department: 18 miles W Honda, 16 March 1955, E. I. Schlinger, E. S. Ross, 1 ♂ (CAS).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32FFFA9E32C73D234A8FC00.taxon	discussion	Comments. This species was previously known from two males collected in Valle del Cauca Department. This site on Rio Azul is located in the Pacific Region and the western Andes (Stark et al. 1999). The new records extend the species distribution to the Central Andes and the middle Magdalena River valley.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32FFFA9E37C76753204FB93.taxon	description	(Figs. 9 - 11)	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32FFFA9E37C76753204FB93.taxon	materials_examined	Material examined. Colombia: [Putumayo Department]: Santa Rosa, Rio San Miguel headwaters, Kofan Indian Village, 2 - 24 October 1970, B. Malkin, P. Burchard, 7 ♂, 2 ♀ (CAS). Santa Rosa de Sucumbios, Kofan Indian Village, Rio San Miguel, 400 m, 26 August- 9 September, 1971, B. Malkin, 1 ♂ (CAS).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32FFFA9E37C76753204FB93.taxon	discussion	Comments. The original description of this species included male and larval specimens, but the adults were in poor condition and the habitus could not be adequately described. This larger sample of male and female specimens allows the descriptions of adult habitus, female genitalia and the egg. This species was previously reported from Guyana and Venezuela. These first records from Colombia are from the Amazon Natural Region. Rio San Miguel joins Rio Putumayo and forms one of the major tributaries of the Amazon River basin.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32FFFA9E37C76753204FB93.taxon	description	Adult habitus. General color yellow-brown, patterned with dark brown. Head with obscure, yellow-brown spot forward of ocelli and dark brown lappets and antennae (Fig. 9). Pronotum brown over most of disc but with moderately pale anterolateral area and pale median band. Dark brown bands on apical third of hind femora; fore and mid femora with wider dark bands; tibiae brown but with darker proximal and distal brown bands. Wings dark amber, veins pale brown except Sc, R and Cord crossvein dark brown. Female. Forewing length 13.5 mm. Subgenital plate with four, approximately equal sized lobes; median notch deeper than lateral notches (Fig. 11). Posterior margin of sternum 9 with narrow sclerotized band bearing several thick setae; median field of sternum 9 with patch of shorter setae lying in median notch of sternum 8, additional longer setae occur posterolateral to median patch. Egg. Length ca. 0.3 mm, greatest width ca. 0.16 mm. Outline spindle shaped with low, button-like collar (Fig. 10). Anchor bearing a small membranous knob which lies over collar.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32FFFA9E18D76C435CEF876.taxon	materials_examined	Material examined. Colombia: Cauca Department: San Andreas [= San Andrés], Tierradentro, ~ 1500 m, 21 - 22 November 1971, B. Malkin, 1 ♂ (CAS).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF32FFFA9E18D76C435CEF876.taxon	discussion	Comments. The species was originally described from five male specimens collected in Cauca Department. The “ San Andreas ” location listed on the label for Malkin’s specimen is the same site as “ San Andrés de Pisimbalá ” from the original description. The site is in the Central Andes of southwestern Colombia and the species is known only from this region in a narrow elevational range of 1500 - 1750 m a. s. l. The site is the seat of the indigenous Paez community (Zúñiga & Stark 2002).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF320FFA6E2D0741D3294F880.taxon	materials_examined	Material examined. Colombia: Cauca Department, San Andreas [= San Andrés], Tierradentro, ~ 1500 m, 21 - 22 November 1971, B. Malkin, 1 ♂ (CAS).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF320FFA6E2D0741D3294F880.taxon	discussion	Comments. This species was previously known from the holotype specimen collected in Ecuador. The new record is the first for Colombia and it represents a significant range extension from the Pacific slope of the Ecuadorian Andes to the Cordillera Central of southwestern Colombia (also see comments on the Colombian locality under A. inza).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA7E37972A33703F94E.taxon	materials_examined	Material examined. Colombia: [Cesar Department]: Socorpa Mission, Sierra de Perija, 1350 m, 20 - 21 August 1968, B. Malkin, 8 ♂ (CAS). Same location, 10 - 22 August 1968, B. Malkin, 16 ♂ (CAS). [Magdalena Department.], San Sebastian de Rabago, Sierra Nevada de Santa Marta, 2000 m, 1 - 10 April 1968, B. Malkin, 7 ♂, 1 ♀ (CAS). Venezuela: Lara State, Parque Nacional Dinira, Quebrada Buenos Aires, 9 ° 36.407 ’ N, 70 ° 04.178 ’ W, 1850 m, 18 - 19 June 2001, Holzenthal, Blahnik, Paprocki, Cressa, 3 ♂ (pinned, BPSC).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA7E37972A33703F94E.taxon	discussion	Comments. This species is known from specimens collected in Barinas and Mérida states, Venezuela (Stark 1995; Maldonado et al. 2002) and is also recorded from Santander Department, Colombia (Stark et al. 1999) and Trujillo State, Venezuela (Stark 1999), however the Colombian specimens from Santander figured in Stark et al. (1999) appear to have a shorter apical projection of the aedeagus and the keel is shown as an obscure V-shaped structure (Figs. 136 - 138 in Stark et al. 1999). The Colombian specimens reported in this study have a narrow, relatively long keel and the apical aedeagal projection is more slender and slightly longer than those shown for other populations. The records given above include the first Colombian specimens from Cesar and Magdalena Departments, the first from Serrania de Perijá, and the sixth Anacroneuria species to be reported from the Sierra Nevada de Santa Marta (Zúñiga et al. 2013). The new records from Lara State are from a national park in the foothills of the northern mountain range of the Venezuelan Andes. Streams in this region drain into several basins, including the Orinoco River and Lake Maracaibo.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA7E37F74BB328FFD65.taxon	materials_examined	Holotype ♂ (United States National Museum), 1.5 km SE Las Piedras de Cocollar, Rio Cocollar, Sucre State, Venezuela	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA7E37F74BB328FFD65.taxon	materials_examined	Material examined. Venezuela: Lara State, Parque Nacional Terepaima, Quebrada San Antonio, 9 ° 51.754 ’ N, 69 ° 13.098 ’ W, 631 m, 17 June 2001, Holzenthal, Blahnik, Paprocki, Cressa, 6 ♀ (pinned, BPSC).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA7E37F74BB328FFD65.taxon	discussion	Comments. This species was previously known from the type locality in Sucre State, Venezuela. These are the first specimens reported from Lara State in the southern foothills of the Sierra de Portuguesa in the Venezuelan Andes. The site is in a protected area and is part of an important biodiversity center for the Andean Cloud Forests ecoregion.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA4E1F77091359CFD64.taxon	description	(Figs. 12 - 16)	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA4E1F77091359CFD64.taxon	materials_examined	Material examined. Holotype ♂, Colombia, [Cesar Department], Socorpa Mission, Sierra de Perijá, ~ 1350 m, 10 - 22 August 1968, B. Malkin (CAS).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA4E1F77091359CFD64.taxon	description	Adult habitus. General color pale brown. Head pale yellow-brown with slightly darker lappets and frons, forward of ocelli (Fig. 12). Antennae dark brown. Eyes with numerous fine, white setae giving a hairy appearance. Pronotum pale yellow-brown with darker sublateral bands; broad median field pale. Wing membrane pale amber with darker veins; C vein pale, Sc and R much darker. Legs pale except for narrow brown distal band superimposed on black band; tibiae with narrow proximal and distal brown bands. Male. Forewing length 12 mm. Hammer thimbleshaped, height subequal to apical diameter (Fig. 13). Aedeagal apex abruptly narrowed from shoulders, forming a slender, median projecting lobe, offset from shoulders by moderately deep grooves (Fig. 14); shoulders slightly scalloped along anterior margin. Dorsal keel short in median field, but with small basolateral struts which connect to longer, dark, arcuate supports (Fig. 16). Apical area in lateral aspect with a basodosal ear-shaped lobe and a slightly more anterior slender crescent lobe; most of apical region membranous (Fig. 15). Hooks moderately wide, saber-like. Female. Unknown. Larva. Unknown.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA4E1F77091359CFD64.taxon	etymology	Etymology. The species name honors the Yuko people who inhabit the region near the type locality. The name is used as a noun in apposition.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA4E1F77091359CFD64.taxon	diagnosis	Diagnosis. The aedeagal structure of A. yuko is similar to that of A. arcabuca Zúñiga & Stark 2006 (in Zúñiga et al. 2006,) and to A. tejon Baena & Stark 1999 (in Stark et al. 1999), but it differs from both in the lateral profile of the apical aedeagal region. In the new species this structure projects more strongly away from the aedeagal body (Fig. 15), whereas, in the two known species, this process is more dorsally oriented. The new species is more similar to the former species but is smaller (17.5 mm forewing in A. tejon), has a more defined keel and less pronounced scalloping along the anterior margins.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF321FFA4E1F77091359CFD64.taxon	discussion	Comments. This new species is the second Anacroneuria to be reported from the Serranía de Perijá range which forms part of the border between northwestern Colombia and adjacent areas of Venezuela (see comments under A. paleta above). The collection site for the holotype lies on the northwestern slope of the mountain range and is in the drainage basin of Rio Cesar which enters the Magdalena River and the Colombian Caribbean. This mountain range is considered part of the biogeographic unit known as the Cinturón árido Pericaribeño.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF322FFA2E35D74F037EDFBB0.taxon	description	(Figs. 17 - 19)	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF322FFA2E35D74F037EDFBB0.taxon	materials_examined	Material examined. Venezuela: Lara State, Parque Nacional Terepaima, Rio Aura near Sabana Alta, 9 ° 44.740 ’ N, 69 ° 16.614 ’ W, 480 m, 16 June 2001, Holzenthal, Blahnik, Paprocki, Cressa, 1 ♀ (pinned, BPSC).	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF322FFA2E35D74F037EDFBB0.taxon	description	Adult habitus. General color dark on head and pronotum and yellow-brown over meso and metathorax and abdomen. Head dark over central frons and yellow brown on anterior half of frons; antennae black (Fig. 17). Pronotal disc dark except for pale anterolateral area. Wings with dark pigment over most of membrane, but with a pale circular window spot beyond cord which extends from above Rs vein to at least the first median vein (Fig. 19); coastal area pale from base to end of Sc vein. Legs mostly black, but hind femora banded, dark apically and pale in basal half. Male. Unknown. Female. Forewing length 10 mm. Subgenital plate with four lobes, similar in length but outer lobes wider than inner pair (Fig. 18). Median sclerite of sternum 9 rather uniformly covered with setae; outer zones of patch with slightly longer and thicker setae. Intersegmental membrane between sterna 9 and 10 densely armed with microtrichia. Egg. Unknown. Larva. Unknown.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
653F6C3CF322FFA2E35D74F037EDFBB0.taxon	discussion	Comments. This species is another small, dark, distinctively pigmented and unassociated female Anacroneuria. Although several similar females have been described in previous studies (e. g. A. wapishana Stark, 1999), this specimen appears distinct from these. The informal designation “ VZ- 13 ” indicates this is the thirteenth Anacroneuria species from Venezuela to be provisionally recognized from unassociated females or larvae (Stark 1995; Maldonado et al. 2003). The collection site is within a national park in the mountainous region of Lara and Portuguesa states. Vegetation in the highlands is cloud forest and streams in this area enter the Orinoco River basin.	en	Stark, Bill P., Zúñiga, Maria del Carmen (2014): New Species And Records Of Colombian, Ecuadorian And Venezuelan Anacroneuria (Plecoptera: Perlidae), With A Review Of The Anacroneuria Aymara Stark & Sivec Complex. Illiesia 10 (8): 66-79, DOI: http://doi.org/10.5281/zenodo.4752749
