taxonID	type	description	language	source
03BB0738FFE7FFC7FEC0FEBFFED1F946.taxon	materials_examined	Type species. ‐ Perla limbata Pictet, 1841. Distribution. ‐ Eastern Palearctic and Oriental regions. Known from China, Japan, Thailand and Vietnam.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE7FFC7FEC0FEBFFED1F946.taxon	diagnosis	Diagnosis. Adult Togoperla are large, typically dark, and fully winged. Males of all known species have a well developed, truncate, emarginate or slightly notched lobe on the 5 th abdominal tergum and large mesal membranous areas on terga 6 ‐ 9 which often bear setal clusters but seldom have sensilla basiconica patches. Male hemiterga have a basal callus and a relatively short (in comparison to Agnetina) anterior process. The aedeagal tube of all species has a basodorsal lobe and most also have a pair of lateral lobes on the tube; the membranous apical sac is coarsely armed with small triangular spines. Females have a large parabolic subgenital plate, frequently with a median sclerite in the dorsal membrane of the plate. The subgenital plate in several species is emarginate or shallowly notched but no examples of narrow deep notches are known for the group. Egg collars are absent for most species, but two species have a sessile collar without a flanged rim and another has a small thickened polar disc. Larvae are associated for only one species, T. limbata (Pictet) (Sivec et al. 1988), which lacks anal gills, has few (usually 1 ‐ 2) slender intercalary setae on abdominal terga, and lacks a median dorsal fringe of long silky setae. Provisional keys for adult Togoperla are presented below the species accounts. Current species. 1) canilimbata (Enderlein, 1909); 2) clavata sp. n.; 3) fortunati Navas, 1926; 4) limbata (Pictet, 1841); 5) noncoloris Du & Chou, 1999; 6) perpicta Klapálek, 1921; 7) poilanina (Navas, 1934); 8) shan Stark & Sivec, 1991; 9) totanigra Du & Chou, 1999; 10) triangulata Du & Chou, 1999; 11) tricolor Klapálek, 1921.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE7FFC5FEE2F93EFC7DF8A0.taxon	description	(Figs. 1 ‐ 6, 57 ‐ 58)	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE7FFC5FEE2F93EFC7DF8A0.taxon	materials_examined	Material examined. China: Guangdong, Yim Na San, 11 ‐ 16 June 1936, L. Gressitt, 3 ♀ (MCZ, T. klapaleki holotype and paratypes). Same location, 11 ‐ 16 June 1936, L. Gressitt, 1 ♂, 1 ♀ (MCZ, T. sinensis holotype and paratype). Vietnam: Manson‐ Gebrige, 2 ‐ 3000 ’, April‐ May, H. Fruhstorfer, 2 ♀ (ZIP, T. canilimbata lectotype and paralectotype). Lao Cai, Sapa, large waterfall on road from Sapa to Lai Chau, UV light, 8 May 1995, ROM 956022, D. Currie, B. Hubley, J. Swann, 16 ♂, 1 ♀ (ROM, IEBR, BPS). Lao Cai, Sapa, Fan Si Pang Mountain, 22 ‐ 29 April 1995, 1600 m, 2 ♂ (ZMB). Adult habitus. General color dark brown, patterned with yellow. Head dark brown over ocelli, extending laterally over calluses and forward to M‐ line; a broad triangular dark patch is located forward of M‐ line (Fig. 1); head pale laterally and behind eyes on occiput. Pronotum brown with scattered paler rugosities. Wings dark brown except costal area pale to cord. Legs banded; both femora and tibiae with pale mesal band and dark basal and apical bands. Male. Forewing length 24 ‐ 26 mm. T 5 produced as an emarginate lobe; T 6 ‐ T 9 typical with anterior margin and lateral areas sclerotized and mid section membranous; membrane of T 6 ‐ T 9 and lobes of T 5 somewhat hairy (Fig. 2). Dorsal aspect of hemiterga relatively short, length from basal callus to hemitergal tip about equal to length of basal callus. Membranous aedeagal sac terminating in a pair of slender, bare lateral lobes and a slender median lobe armed with fine brown setal spines; subapical area of aedeagus armed with a coarse band of triangular spines and microtrichia (Figs. 5 ‐ 6); spine band narrowly interrupted mid‐ dorsally but forming a complete band to beyond mid‐ length on venter. Dorsobasal area of aedeagus bearing a pair of small, bare lateral lobes. Female. Forewing length 28 ‐ 30 mm. Subgenital plate parabolic, usually with a median notch or emargination; plate extends over much of S 9 (Fig. 3); dorsal sclerite of plate not reaching to lateral margins 2 5 6 of sternum 8. Vagina with small bulge at midlength (Fig. 4); vaginal accessory glands subequal to vagina in length. Egg. Length 0.42 ‐ 0.44 mm, width 0.30 ‐ 0.32 mm. Collar absent, general shape oval (Fig. 57). Micropylar row set near equator. Chorion smooth (Fig. 58).	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE7FFC5FEE2F93EFC7DF8A0.taxon	distribution	Distribution. Known from southeastern China and northern Vietnam.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE7FFC5FEE2F93EFC7DF8A0.taxon	discussion	Remarks. Sivec et al. (1988) considered T. canilimbata, T. fortunati and T. grahami as synonyms and this was repeated by Du & Chou (1999). Zwick (1973 b) found syntype females of T. canilimbata and illustrated the lectotype subgenital plate and vagina; although the subgenital plate is shown without emargination, this is present on the paralectotype and the two specimens agree in coloration and in vaginal shape. Evidently the subgenital plate margin shows some variation in the expression of a notch or emargination. One of us (IS) examined the lectotype and paralectotype and noted both femora and tibiae have wide yellow median bands which are larger on the hind legs. This color pattern is in agreement with the leg pattern of T. klapaleki and T. sinensis described by Banks (1939) as “ ... black band at base and tip of femora and tibiae as in perpicta. ” Types of T. fortunati and T. grahami, however, lack pale tibial bands and also lack a dark basal band on the femora, consequently these species are removed from synonymy with T. canilimbata. A consequence of these changes in synonymy is the male of this species is now based on T. sinensis and the recent material from Vietnam with short hemitergal lobes, rather than T. fortunati and T. grahami with long hemitergal lobes. Togoperla tonkinensis had been reported only from the holotype, supposedly a male, but with the abdomen missing. The damaged specimen from Vietnam, in the Museum National d’Histoire Naturelle has the head, thorax, wings and some legs intact. The legs show the characteristic banding pattern associated with T. canilimbata and allows the placement of T. tonkinensis as a synonym of that species. 8 11 12	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE2FFC2FEA0FEC5FB6FFAF1.taxon	materials_examined	Material examined. Holotype ♂ (alcohol) and 7 ♂, 1 ♀ (pinned) from Vietnam, Vinh Phu, Tam Dao Hill Station, stream 5 km along trail at west edge of town, UV light, 11 May 1996, B. Hubley, M. Hanson, ROM 961013 (ROM). Additional paratypes: Vietnam: Vinh Phu, Tam Dao, lower waterfall of stream flowing through town, UV light, 11 May 1996, B. Hubley, D. C. Darling, ROM 961030, 4 ♂ (ROM, IEBR). Vinh Phu, Tam Dao Hill Station, 2 km along trail at west edge of town, 4 ‐ 31 May 1996, ROM 961005, B. Hubley, 1 ♀ (pinned, ROM). Thua Thien‐ Hue, Bach Ma National Park, Campsite trail, edge of small stream, 1200 m, 2 June 2000, ROM 2000502, B. Hubley, 1 ♀ (pinned, ROM). Adult habitus. General color dark brown. Head mostly brown except for pale M‐ line and sublateral areas between lappets and dark area along anterior frons (Fig. 7); area between eyes almost entirely dark brown. Pronotum dark brown with darker rugosities and slightly paler area along median suture. Wings dark brown, veins darker except for pale band on costa from base to cord. Femora banded with basal yellow band and dark brown apical band; tibiae and tarsi dark brown. Male. Forewing length 21 ‐ 23 mm. Hairbrushes well developed on metathoracic basisternum and abdominal sterna 6 ‐ 7, weakly developed on sterna 5 and 8. Abdominal tergum 5 produced into a truncate or emarginate lobe partially covering T 6; T 6 ‐ 9 with mesal sclerite set in large membranous posteromesal field bearing long setae (Fig. 8). Hemitergal finger process longer than basal callus, area forward of callus ca. 1.8 X longer than callus; process deeply grooved between lateral margin and callus in dorsal aspect, and somewhat club shaped in lateral aspect (Figs. 13 ‐ 14). Basolateral lobes of aedeagal tube well developed and covered with fine spines; aedeagal sac armature consists of coarse triangular spines and smaller spicules covering most of sac in lateral and dorsal aspects (Figs. 11 ‐ 12); patch terminates at level of lateral lobes and is constricted on dorsal surface forward of lobes; patch is very narrowly divided along median field. Sac bears a pair of apical dorsolateral lobes and a smaller mesal one above a terminal ventroapical polyp shaped lobe; the latter lobe bears a sparse coat of fine, long setal‐ like spines. Female. Forewing length 26 ‐ 29 mm. Subgenital plate parabolic with small apical emargination (Fig. 9); dorsal sclerite present on plate; intersegmental membrane between sterna 9 and 10 covered with microtrichia. Vagina slightly swollen posterior to midlength (Fig. 10) and lined with pale brown spinules. Egg. Outline oval, collar absent, chorion smooth, similar to T. canilimbata. Larva. Unknown.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE2FFC2FEA0FEC5FB6FFAF1.taxon	etymology	Etymology. The species name refers to the club shaped lateral aspect of the male hemitergal lobes.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE2FFC2FEA0FEC5FB6FFAF1.taxon	diagnosis	Diagnosis. The long hemitergal lobes and banded femora are similar to those of T. poilanina (see below) and T. fortunati. It differs from males of the latter in shape of the hemitergal lobes in lateral aspect and in aedeagal armature and shape. The aedeagus for T. fortunati is more cylindrical, lacks the basolateral lobes and also differs in armature patterns. Externally the presence of brown sclerites on terga 6 ‐ 9 is typical of T. clavata but not T. fortunati. Two of the Chinese species described by Du & Chou (1999) also share these tergal sclerites, but neither have the same pattern of aedeagal armature and one of them (T. noncoloris) also differs in pigment pattern and in hemitergal lobe shape. The subgenital plate form will not distinguish females of this species from other Togoperla, but the barrel shape of the vagina and the color pattern will help in recognition.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE2FFC0FC5FFAE8FCE9F8E2.taxon	description	(Figs. 15 ‐ 20)	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE2FFC0FC5FFAE8FCE9F8E2.taxon	materials_examined	Material examined. China: Kwieyang, Kouy‐ Tcheou, Fujian Province, 1906, P. P. Cavalerie and Fortunat, 1 ♂ (MNHM, T. fortunati holotype). Sichuan Province, Lin Ngai Si, near Kuanshien, 3500 ’, 20 September 1930, D. C. Graham, 1 ♂ (USNM, T. grahami holotype). Sichuan Province, Emei Shan, 710 m, 20 June 1979, 1 ♂, 1 ♀ (ZIAS). Sichuan Province, Won Chuan, August 1938, D. C. Graham, 1 ♀ (USNM). Sichuan Province, Kuanshien, 3 ‐ 13 July 1937, 1 ♂, 3 ♀ (MCZ). Adult habitus. General color brown to dark brown. Ocellar area covered by dark brown quadrangular spot and area forward of M‐ line with smaller triangular dark area (Fig. 15); occiput pale brown. Pronotum dark brown with scattered rugosities. Wings brown with pale costal area. Femora banded, pale basally and dark apically; tibiae dark brown. Male. Forewing length 20 ‐ 22 mm. T 5 produced into an emarginate lobe; T 6 ‐ 9 with large mesal, membranous field covered with fine setae (Fig. 16). Projecting portion of hemitergum about twice as long as basal callus in dorsal aspect, gradually tapered to rounded tip in lateral aspect (Fig. 18). Aedeagus cylindrical, relatively straight and armed with microtrichia and small triangular spines over most of apical half (Fig. 17); apex terminating in a slender cylindrical lobe armed with fine brown setal spines; basolateral lobes absent. Female. Forewing length 24 ‐ 26 mm. Subgenital plate covers most of sternum 9 and is slightly emarginate or entire on posterior margin (Fig. 19). Vagina constricted near midlength; accessory glands less than half as long as vagina (Fig. 20). Egg. Outline oval. Collar absent. Chorion smooth, similar to T. canilimbata. Larva. Unknown.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE2FFC0FC5FFAE8FCE9F8E2.taxon	discussion	Remarks. This species is known from southwest China, primarily from the T. fortunati and T. grahami 21 22 type series. Sivec et al. (1988) listed it as valid, but erroneously placed T. canilimbata as a synonym; had T. canilimbata and T. fortunati remained synonyms, the former species would have priority. As discussed above under T. canilimbata, these species are now regarded as distinct, primarily due to differences in leg banding patterns and distribution. 25	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFC1FEB4FEC5FA96FBA3.taxon	description	(Figs. 21 ‐ 27)	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFC1FEB4FEC5FA96FBA3.taxon	materials_examined	Material examined. Japan: Unknown locality, 4 July 1981, S. Uchida, 1 ♂ (PMSL). Hozuyama, Kuwada Gori, Tamba, June 1897, 3 ♀ (??). Honshu, Obama, May 1926, 1 ♂ (CSU). Adult habitus. General color brown. Head with dark brown pigment over ocelli and forward of M‐ line; occiput dusky brown. Pronotum brown but usually with pale anterolateral areas on disc (Fig. 21). Wings brown except costal area pale. Legs entirely brown, without distinctive banding. Male. Forewing length 19 ‐ 22 mm. T 5 produced into a notched or emarginate lobe; T 6 ‐ T 9 typical; membrane of T 6 ‐ T 9 bearing patches of mixed setae and scattered sensilla basiconica. Dorsal aspect of hemiterga moderately long and attenuated from basal callus to tip; length from basal callus to hemitergal tip about 1.3 ‐ 1.5 times as long as basal callus (Fig. 22); lateral aspect of hemiterga slender across basal curve and gradually narrowed to form a slender triangular structure (Fig. 23); sensilla basiconica clustered on basal callus and sparse on hemitergal tip. Membranous aedeagal sac armed with a coarse band of variably sized spines from subapical lobes to about midlength of tube (Fig. 25); largest spines occur in a band somewhat proximal to margin of spinous zone; apex of sac extends as a nose like lobe below a pair of membranous, unarmed dorsal lobes; apical lobe armed at tip with a zone of slender, long spines; tube with a pair of midlateral spiny lobes. Female. Forewing length 24 ‐ 29 mm. Subgenital plate parabolic without apical emargination or notch. Dorsal membrane of plate with a short sclerotized mesal band reaching less than halfway to base of plate (Fig. 26). Vagina with essentially parallel sides, or sides slightly constricted at back of chamber (Fig. 27); accessory glands about half as long as vagina. Egg. Length 0.38 ‐ 0.42 mm, width 0.31 ‐ 0.34 mm. Collar expressed as a small dark brown disc. Micropyles oriented almost perpendicular to long axis of egg; micropylar row set almost halfway between equator and pole.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFC1FEB4FEC5FA96FBA3.taxon	distribution	Distribution. Known only from Japan.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFC1FEB4FEC5FA96FBA3.taxon	discussion	Remarks. Sivec et al. (1988) proposed the synonymy of T. kawamurae and T. matsumurae with T. limbata but omitted supporting data. Uchida (1990) supported these synonymies in his thesis, but this has remained unpublished. We accept and validate the lectotype for T. matsumurae, previously selected by Uchida (1990) in his thesis, which bears label data of, Haki, 8 / 10, Perla matsumurae Okam., Sesuji‐ kawagera, & Lectotype, Perla matsumurae Okamoto, S. Uchida det. 1984, HU, and is in the Hokkaido University collection, Sapporo. Uchida (1990) also described a second Japanese Togoperla species, generally similar to T. limbata, but differing in details of aedeagal armature, however this description has not been published and we have seen no material. Male hemiterga of this species are generally similar to those of T. fortunati, T. poilanina and others, however it is easily distinguished by virtue of geography and by the pattern of armature on the aedeagus.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFC1FC7EFBA6FBD6FA17.taxon	discussion	Remarks. We have seen no material of this species, but it appears distinctive by virtue of its pale wing pigmentation. The hemitergal lobes are short, the basolateral lobes of the aedeagal tube are small and bare, and the dorsoapical spine patch is fused mesally (Du & Chou 1999).	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFCFFC5AF9D5FE38F912.taxon	description	(Figs. 28 ‐ 36)	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFCFFC5AF9D5FE38F912.taxon	materials_examined	Material examined. China: Hong Kong, February‐ May, 1 ♂ (BMNH, T. perpicta lectotype). Fujian, Fuzhou (Foochow), 1935 ‐ 1936, M. S. Yang, 1 ♂ (BMNH). Same site, 9 May 1936, M. S. Yang, 1 ♂ (BMNH). Fujian, Fuzhou, San Chiang, 1927, C. H. Pope, 1 ♂. Fujian Province, Tachulan, 4500 ’, foot of Kuatun Mountain, 21 ‐ 26 April 1948, J. Fu, 2 ♂, 1 ♀ (USNM). Same site, 21 ‐ 25 May 1948, J. Fu, 1 ♂, 1 ♀ (USNM). Same site, 16 ‐ 19 June 1948, J. Fu, 1 ♂ (USNM). Zhejiang (Chekiang), Hangzhou (Hangtcheou), 1925, A. Pichon 1 ♀ (MNHN, T. pichoni holotype). Adult habitus. General color brown. Head with dark pigment over ocellar region, occiput and forward of M‐ line; pale around eyes and on lateral margins (Fig. 34). Pronotum brown, wings brown except costal field. Legs banded, proximal and distal parts of femora and tibiae brown, mesal area yellow. Male. Forewing length 18 ‐ 21 mm. T 5 produced into a truncate to emarginate lobe; T 6 ‐ T 9 typical with membrane bearing patches of long setae; T 5 lobe with apical cluster of sensilla basiconica (Fig. 35). Dorsal aspect of hemiterga moderately long and only slightly narrowed from basal curve to tip; length from basal callus to hemitergal tip about 2.3 ‐ 2.8 times as long as basal callus; lateral aspect of hemiterga gradually narrowed from basal curve to midlength but scarcely narrowed beyond midlength to rounded tip (Figs. 31 ‐ 33). Membranous aedeagal sac coarsely armed with a broad basal spine band, interrupted on median dorsal surface; sac beyond spine band bearing a pair of dorsal unarmed or sparsely armed lobes and terminating with a long cylindrical lobe armed with fine long setal spines in apical half (Fig. 36); tube membranous and bearing a pair of spiny dorsolateral lobes near sac base. Female. Forewing length 24 ‐ 27 mm. Subgenital plate parabolic with a truncate apex (Fig. 28). Dorsal membrane of plate with a sclerotized mesal band extending about halfway from margin to base. Vagina expanded slightly at midlength (Fig. 29); accessory glands about two thirds vaginal length. Egg. Length about 0.53 mm, width about 0.39 mm. Collar absent, outline oval with broadly rounded poles. Micropylar canals strongly slanted; micropylar row set near equator. Chorion smooth.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFCFFC5AF9D5FE38F912.taxon	distribution	Distribution. Known from mainland China, Hong Kong and Vietnam.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFE1FFCFFC5AF9D5FE38F912.taxon	discussion	Remarks. Sivec et al. (1988) illustrated this species and established the synonymy given above. Because types for the synonyms are either lost or badly damaged we have relied on illustrations by Klapálek (1923) and Wu & Claassen (1934) for T. bifoveolata and T. elongata, respectively. The T. pichoni type female in Paris is in very poor condition but the subgenital plate, vagina and leg pattern are consistent with those of T. perpicta.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEFFFCCFF0FF8CBFBCDF912.taxon	description	(Figs. 37 ‐ 43, 63 ‐ 68)	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEFFFCCFF0FF8CBFBCDF912.taxon	materials_examined	Material examined. Vietnam: Quang Tri [Annam], 1925, Poilane, 1 ♀ (MNHN, holotype). Vinh Phu Province, Tam Dao, 800 ‐ 1100 m, 21 ° 28 ’ N, 105 ° 38 ’ E, 19 May‐ 13 June, 1995, H. Malicky, 1 ♂, 1 ♀ (PMSL). Adult habitus. General color dark brown. Head with dark brown area covering ocelli and forward to M‐ line. Pronotum brown with scattered rugosities and dark lateral margins (Fig. 37). Wings dark brown except for pale costal margin to cord. Femora pale on basal third to half and dark apically; tibiae entirely dark brown. Male. Forewing length about 19 mm. Hemitergal lobes about twice as long as basal callus (Fig. 41); lobes gradually tapered from base to apex in lateral aspect (Fig. 38). T 6 ‐ 9 with large mesal membranous area surrounded laterally and along anterior margin by sclerotized area; entire membranous area covered by setae. Tergum 5 produced as an emarginate lobe. Aedeagus sinuate in lateral aspect and swollen apically; apex with a dorsolateral pair of ear‐ like lobes and a terminal cylindrical lobe; armature ending subterminally and barely extending onto bases of lateral lobes; armature absent from basolateral lobes and from a large lateral area but covering most of venter; dorsal armature patch strongly narrowed to a small strip that approaches, but does not reach, basolateral lobes (Figs. 42 ‐ 43). Female. Forewing length 27 ‐ 29 mm. Subgenital plate parabolic and slightly emarginate or shallowly notched. Dorsal sclerite on plate slender, extending to level of lateral margins of sternum 8 (Fig. 39). Lateral margins of vagina parallel for most of length (Fig. 40). Egg. Outline barrel shaped; length ca. 0.37 mm, width ca. 0.30 mm. Collar sessile but wide and surrounded by a smooth rim with irregular lobing (Figs. 63 ‐ 66). Chorion coarsely punctate except on rim (Figs. 67). Micropylar orifices surrounded by five or six chorionic punctations (Fig. 68). Larva. Unknown.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEFFFCCFF0FF8CBFBCDF912.taxon	discussion	Remarks. The association of the K. poilanina holotype with fresh material from Tam Dao was confirmed through comparison of SEM micrographs of eggs from the holotype and from the new locality. The two samples were found to be very similar and to be atypical for Togoperla, However the associated male from Tam Dao and the presence of a large subgenital plate with dorsal sclerite indicate this species is a true member of Togoperla. This species occurs in the same area as T. clavata (described above), although not necessarily from the same stream, and the two species have similar male hemiterga, general coloration and leg patterns. Males of the two may be distinguished by the presence of brown sclerites in the membrane of terga 6 ‐ 9 for T. clavata and by the club shaped hemiterga in lateral aspect for that species. In addition the aedeagal armature for T. clavata is narrowly divided along the mid‐ dorsum and the basolateral lobes are covered with scale‐ like triangular spines. The color patterns also differ slightly on the head with T. clavata having a much more extensive area of black pigment between the compound eyes.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFECFFCAFC55F8D5FD7BFC1D.taxon	description	(Figs. 44 ‐ 50, 59 ‐ 60)	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFECFFCAFC55F8D5FD7BFC1D.taxon	description	45 47 48	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFECFFCAFC55F8D5FD7BFC1D.taxon	materials_examined	Material examined. Thailand: Chiang Mai, amp. Fang, tambon Papah, 18 ‐ 20 April 1975, A. E. Gordon, 14 ♂, 4 ♀ (USNM, BPS, PMSL). Adult habitus. General color brown. Head with dark pigment over ocelli and forward of M‐ line. Pronotum brown with indistinct rugosities (Fig. 44). Legs brown without distinct banding. Male. Forewing length 19 ‐ 21 mm. T 5 produced into a truncate or slightly emarginate lobe; T 6 ‐ T 9 typical; T 5 lobe with marginal tufts of long setae (Fig. 45). Dorsal aspect of hemiterga moderately long with prominent sulcus and with tips curved slightly laterad; length from basal callus to hemitergal tip about 1.6 ‐ 2.0 times as long as basal callus; lateral aspect triangular in shape but tip blunt (Fig. 46); sensilla basiconica clustered on basal callus and scattered on hemitergal tip. Membranous aedeagal sac coarsely armed with a broad band of short thick spines; band ends near sac apex on dorsal surface but continues onto tube on venter (Figs. 47 ‐ 48); sac apex trilobed; tube with a pair of unarmed lateral lobes. Female. Forewing length 29 ‐ 31 mm. Subgenital plate parabolic with emarginate apex (Fig. 49). Vagina constricted beyond midlength (Fig. 50); accessory glands short, much less than half vaginal length. Egg. Length 0.45 ‐ 0.47 mm, width 0.35 ‐ 0.37 mm. Outline pear shaped with wide posterior pole and narrow collar; collar width 0.12 ‐ 0.14 mm, length about 0.01 ‐ 0.02 mm. Collar distinct but sessile, rim smooth but with a row of pits circling base (Fig. 59). Chorion relatively smooth but with faint follicle cell impressions (Fig. 60). Micropyles subequatorial with canals slanted.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFECFFCAFC55F8D5FD7BFC1D.taxon	distribution	Distribution. Known only from Thailand.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFECFFCAFC55F8D5FD7BFC1D.taxon	discussion	Remarks. Other than T. poilanina, this is the only Togoperla species known to have a distinct egg collar but in male and female genitalic features it is typical of the genus. The eggs of T. poilanina are coarsely punctate and those of T. shan lack punctations.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEAFFCAFED0FBC3FDFEF9FF.taxon	discussion	Remarks. We have seen no material of this species, but it seems distinctive by virtue of a black mesal sclerite on T 9, a character not otherwise reported among Togoperla species. The hemitergal lobes are short, the basolateral lobes of the aedeagal tube are small and bare, and the dorsoapical spine patch of the aedeagal sac is completely divided by a narrow, median line. The legs are presumed to be dark since Du & Chou (1999) indicate the species is “ … similar to Togoperla chekianensis ” [= T. tricolor].	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEAFFCAFEE7F9E6FC67FEC9.taxon	discussion	Remarks. We have seen no material of this species which is said to be “ … similar to Togoperla perpicta … ” and presumably has banded femora and tibiae. The hemitergal lobes are long and “ triangular ” and the basolateral lobes of the aedeagal tube lack spines (Du & Chou 1999).	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEAFFCBFC5FFEBFFAA6F967.taxon	description	(Figs. 51 ‐ 56, 61 ‐ 62)	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEAFFCBFC5FFEBFFAA6F967.taxon	materials_examined	Material examined. China: Kiu Kiang, 1 ♀ (BMNH, T. tricolor holotype). Fujian Province, Tachulan, 4500 ’, foot of Kuatun Mountain, 1 ‐ 5 May 1948, J. Fu, 1 ♂ (USNM). Adult habitus. General color dark brown. Head with dark brown area covering ocelli. Pronotum, femora and wings dark brown. Male. Forewing length about 22 mm. T 5 produced into a truncate or slightly emarginate lobe with lateral angles projecting (Fig. 51); T 6 ‐ T 9 typical, membranous areas bearing long setae. Dorsal aspect of hemiterga short, with a prominent sulcus and broadly rounded tip; length from basal callus to hemitergal tip slightly less than length of callus; sensilla basiconica clustered on basal callus and hemitergal tips (Fig. 56). Membranous aedeagal sac coarsely armed with short thick spines in a wide band encircling mesal section of sac (Figs. 52 ‐ 53); distal margin of dorsal patch strongly notched and proximal margin of dorsal patch terminating in a pair of short pointed areas. dorsolateral surface with a pair of small unarmed lobes; apical lobe cylindrical and armed with an apical patch of long thin setae; dorsobasal lobes unarmed. Female. Forewing length 25 ‐ 32 mm. Subgenital plate parabolic, usually with a shallow apical notch or emargination. Dorsal membrane of plate with a sclerotized mesal band extending less than half the distance from margin to base (Fig. 54). Vagina swollen near midlength and strongly constricted near orifice (Fig. 55); accessory glands about half as long as vagina. Egg. Length 0.44 ‐ 0.47 mm, width 0.31 ‐ 0.32 mm. Collar absent, outline oval with broadly rounded poles (Fig. 61). Micropylar canals strongly slanted and subequatorial in position (Fig. 62).	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEAFFCBFC5FFEBFFAA6F967.taxon	distribution	Distribution. Known from east China.	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFEAFFCBFC5FFEBFFAA6F967.taxon	discussion	Remarks. Sivec et al. (1988) considered the three species listed above as synonymous (but incorrectly placed them in the synonymyof T. sinensis), whereasDu & Chou (1999) apparently overlooked T. tricolor and treated T. chekiangensis as the valid name with T. sinensis, T. valvulata, and T. klapaleki assynonyms. Typesfor T. chekiangensis and T. valvulata are lost butfiguresin Wu (1938) showmalesof these species to have the short wide hemiterga typical of T. tricolor and both are reported to have uniformly dark legs similar to those of T. tricolor, consequently we continue to 51 support the synonyms proposed by Sivec et al. (1988), except T. tricolor is recognized as the valid name. The dorsal armature pattern on the aedeagus of this species is similar to the pattern of T. noncoloris in having a distal V‐ shaped incursion of membrane, but that species has dark mesal sclerites on terga 6 ‐ 9 which are not present in this species. In addition, Du & Chou (1999) do not show the proximal edge of the dorsal aedeagal armature to project in the same way as in this species, and the wing pigmentation patterns also differ. 56	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
03BB0738FFF6FFD6FC87FE98FACBFAF1.taxon	discussion	(T. noncoloris, T. totanigra, T. triangulata not included)	en	Stark, Bill P., Sivec, Ignac (2008): The Genus Togoperla Klapálek (Plecoptera: Perlidae). Illiesia 4 (20): 208-225, DOI: http://doi.org/10.5281/zenodo.4754843
