identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
10F43D86F6B6539C83F6889F86AF3713.text	10F43D86F6B6539C83F6889F86AF3713.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coframalaxius bletteryi Le Cesne & Bourgoin, sp. nov.	<div><p>Coframalaxius bletteryi Le Cesne &amp; Bourgoin, sp. nov.</p><p>Genebank registration.</p><p>ON087562/ ON113342/ ON087640/ ON231299/ ON101633.</p><p>Etymology.</p><p>Dedicated to Jonathan Blettery who discovered the first specimen during a fieldtrip with the two first authors exploring caves around Nice in the south of France.</p><p>Diagnosis.</p><p>Small species externally similar to  Trigonocranus emmeae Fieber, 1876, from which it can easily be separated by the triangular areolet of the vertex (versus pentagonal (Emeljanov 2015: 69 and fig.21.6) in  T. emmeae), but also by the rounded posterior margin of the pronotum (versus angular), the conformation of the male genitalia with thinner spiniform processes also different in number and conformation and particularly by a unique internal distinct process inside the periandrium (Fig. 4).</p><p>Color.</p><p>Compound eyes, post clypeus, areolet, anterior part of prothorax behind vertex and mesonotum black, frons paler and carinae yellowish-brown. Tegmina translucent without color patches, pterostigma and vein pale brown, with setiferous granules darker; veins after nodal line darker. Legs pale brown. Metatibia and metatarsomere teeth black, median spines of metarsormere II with setae paler.</p><p>Male terminalia with anal tube regularly ovoid, in dorsal view more or less regularly convex lateroventrally in lateral view with a pair of lateroventral teeth directed posteroventral. Male genitalia asymmetrical with suspensorium X-like, connected to the perandrium basally shortly wide then distally tube-like; bearing 5 spiniform processes: a very basal and dorsal short straight spiniform process (1), on left side a long proximal spiniform process directed dorsoposteriorly then bent posteroventrally (2), a short acute ventral process (3), an elongate dorsal spiniform process forked at mid length in a short teeth-like (4) and a short internal hook-like process, located inside the periandrium (5). Gonopore opening large on apical right side, endosoma not visible, probably very weakly developed.</p><p>Female terminalia (Fig. 5) with paired sternite VI; each gonocoxa VIII developed in a wide plate slightly bilobed lateroapically; gonapophyses IX fused, slightly larger than gonapophyses VIII; gonapophyses VIII larger proximally in a wide triangular endogonocoxal lobe; gonoplacs longer than gonapophyses and enveloping them. Opening of ectodermic internal parts with developed lateral swellings of vestibulum, opening into posterior vagina developed in a strong wrinkled membranous pouch on left side, opening dorsally into the ductus bursae leading to a membranous translucent bursa copulatrix, and laterally into the anterior vagina bent at mid length and ending in the spermatheca; spermatheca with pars intermedialis, half-length of anterior vagina, opening into a diverticulum ductus bent, followed by a long pars intermedialis before vanishing into the the mesodermic glandula apicalis.</p><p>Material type.</p><p>Holotype male, pinned, original description: [Grotte de la  chèvre d’or; Roquefort-less-Pins; France (06)], [12-VII-2021; J. Blettery rec.], [ Coframalaxius bletteryi Le Cesne &amp; Bourgoin sp. nov.; M. Le Cesne det. 2022], [MNHN(EH) 24997].</p><p>Paratypes: 1 female, pinned, with genitalia in a separate microvial, original label: [Grotte de la  chèvre d’or; Roquefort-les-Pins; France (06)], [ Coframalaxius bletteryi Le Cesne &amp; Bourgoin sp. nov.; M. Le Cesne det. 2022], [MUSEUM PARIS; 12-VII-2021; T. Bourgoin rec.], [Museum Paris; MNHN(EH) 24998]; 4 females, pinned, original label: [France, 06; Roquefort-les-Pins; grotte de la  chèvre d’or],[MUSEUM PARIS; 12-VII-2021; T. Bourgoin rec.], [ Coframalaxius bletteryi Le Cesne &amp; Bourgoin sp. nov.; M. Le Cesne det. 2022], [Museum Paris; MNHN(EH) 25177, 25178, 25179, 25180]; 2 males, pinned, with genitalia in a separate microvial, original label: [France, 06; Roquefort-les-Pins; grotte de la  chèvre d’or],[MUSEUM PARIS; 17-VII-2021; M. Le Cesne rec.], [ Coframalaxius bletteryi Le Cesne &amp; Bourgoin sp. nov.; M. Le Cesne det. 2022], [Museum Paris; MNHN(EH) 25181, 25182]; 3 males, pinned, original label: [France, 06; Roquefort-les-Pins; grotte de la  chèvre d’or],[MUSEUM PARIS; 12-VII-2021; T. Bourgoin rec.], [ Coframalaxius bletteryi Le Cesne &amp; Bourgoin sp. nov.; M. Le Cesne det. 2022], [Museum Paris; MNHN(EH) 25183, 25184, 25185]; 3 males, pinned, original label: [France, 06; Roquefort-les-Pins; grotte de la  chèvre d’or],[MUSEUM PARIS; 12-VII-2021; J. Blettery rec.], [ Coframalaxius bletteryi Le Cesne &amp; Bourgoin sp. nov.; M. Le Cesne det. 2022], [Museum Paris; MNHN(EH) 25186, 25187, 25188].</p><p>Other material.</p><p>Several other nymphs at various instars, pinned.</p><p>Type locality.</p><p>Roquefort-les-Pins, Alpes-Maritimes, France</p><p>Phylogeny.</p><p>Coframalaxius bletteryi was sequenced for: COI, Cytb, H2A, 18S and 28S (D3-D5, D6-D7), but only successfully sequenced for 28S (D6-D7) for  Trigonocranus emmeae . Comparison between the two taxa shows significant differences in both total characters and base frequencies, that differs by 37 characters (4.4%) (Table 4). The resulting topology of the molecular analysis (Fig. 8) confirms the placement of  Coframalaxius into  Oecleini as sister to  Trigonocranus .</p></div>	https://treatment.plazi.org/id/10F43D86F6B6539C83F6889F86AF3713	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Le Cesne, Maxime;Bourgoin, Thierry;Hoch, Hannelore;Luo, Yang;Zhang, Yalin	Le Cesne, Maxime, Bourgoin, Thierry, Hoch, Hannelore, Luo, Yang, Zhang, Yalin (2022): Coframalaxius bletteryi gen. et sp. nov. from subterranean habitat in Southern France (Hemiptera, Fulgoromorpha, Cixiidae, Oecleini). Subterranean Biology 43: 145-168, DOI: http://dx.doi.org/10.3897/subtbiol.43.85804, URL: http://dx.doi.org/10.3897/subtbiol.43.85804
F7BB43F68C925E80A97420CF1D6D8E4A.text	F7BB43F68C925E80A97420CF1D6D8E4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coframalaxius Bourgoin & Le Cesne 2022	<div><p>Coframalaxius Bourgoin &amp; Le Cesne gen. nov.</p><p>Type species.</p><p>Coframalaxius bletteryi Le Cesne &amp; Bourgoin (by present designation and monotypy).</p><p>Etymology.</p><p>Arbitrary combination of the first syllabus of author (T. Bourgoin) four grandsons with suffix -xius from  Cixius, type genus of the family  Cixiidae .</p><p>Diagnosis.</p><p>Small cixiids, in habitus resembling  Trigonocranus Fieber, 1875, but can be distinguished from the latter by the combination of the following characters: 1) pygofer longer in lateral view, expanded in a triangular lobe on its latero-posterior margin, 2) anal tube with proximal pair of lateroventral teeth, 3) posterior part of gonostyli wider and 4) aedeagus with one internal spine-like process. Female pygofer elongated, without wax plates.</p><p>Description.</p><p>Head capsule. Vertex with posterior compartment sub-rectangular, anterior compartment triangular; subapical carina straight weakly marked, apical carina well distinct, median carina weak vanishing at subapical carina level; in lateral view, slightly surpassing lateral carina. Frons wider at ventral level of antennae in frontal view; frontoclypeal suture slightly arched dorsally, median carina weak, distinct only in dorsal part and not reaching median ocellus; in lateral view, regularly convex, slightly surpassing laterofrontal carina. Postclypeus with lateral margins slightly concave in basal 1/3; in frontal view, median carina weak in ventral 2/3. Anteclypeus lacking median carina. Compound eye thinly elongated in dorsal view. Antennal socket wide, emarginated, almost touching ventral margin of compound eye; scape short, pedicel globular with distinct transversal margin in frontal view, flagellum with basal swelling well developed, almost five times as long as pedicel, surpassing in length the level of lateral side of abdomen (Fig. 2). Lateral ocelli present, separated from compound eye by 1X their length. Labium almost reaching metacoxae; apical segment 1/4 length of subapical one, slightly thinner medially, proximal segment half as long as subapical one.</p><p>Thorax. Prothorax anterior margin widely roundly concave (Fig. 2); posterior margin more sharply roundly concave (Fig. 2); median carina present, not reaching posterior margin; postocular carinae well distinct, not meeting posterior margin and running subapically to its ventral margin in frontal view (Fig. 2B, C). Mesonotum tricarinate with median and lateral carinae. Forewing elongated; stem ScP+R+MP slightly longer than basal cell length, forking at 1/4 of forewing length, before level of PCu+A1 fusion; anterior and posterior margins subparallel. C1 cell distinctly curved; anterior branch of MP (MP1+2) forking twice: in MP1 and MP2, then MP1a, and MP1b. Posterior branch MP3+4 single, unforked. C5 cell short, diamond-shaped, ending with CuA1+CuA2 fused; icu reaching apex of clavus (Fig. 3A). Hindwing with MP and CuA connecting in I-type (Fig. 3C).</p><p>Hindleg with metatibia laterally unarmed, with 6 apical teeth separated in two groups of 3 by a wide diastema, and outermost tooth largest; first metatarsomere elongate, not dilated apically, with 8-9 apical teeth; second metatarsomere with (7-8) apical teeth, without setae under the first one or two teeth on each side, with one long straight setae under the three to five medium teeth; metatibitarsal formula: 0-(3d3)/(8-9)/(7-8)</p><p>Male genitalia. Anal tube symmetrical, with an anterior pair of lateroventral conspicuous hook-like spines. Pygofer symmetrical, dorsocaudally produced into a triangular lobe; suspensorium developed, X-shaped, attached to ventral margin of anal tube (Fig. 4). Male copulatory organ with periandrium tube-like, elongated, bearing spines and processes. Aedeagus s.s. very short, endosoma not or very shortly developed. Gonostyli bilaterally symmetrical, with proximal portion slender, apically developed into a spoon-shaped extension with a nearly pentagonal outline, directed dorsally.</p><p>Female genitalia of orthopteroid type, sword-shaped ovipositor, following paired hemisternite VII medially divided by a membranous portion (Fig. 5). Pygofer of elongated type without ventral wax plate. Gonoplac fused, apically separated. Ductus receptaculi regular, short, not developed in a helix-twirled structure (Fig. 5C).</p><p>Nymphs. Two short laterometatibial spines in 5th instar (Fig. 6). With abdominal paired tergal wax plates on tergites VI to VIII divided into 6 subplates separated by one sensory pits, those ones ranged in raw in subanterior position (Fig. 6). In instars 3 to 5: procoxa with one sharp anterior process bearing small (sensorial?) dark triangular microcuticular sculptures; profemur with a conspicuous latero-extern process bearing a row of short sensorial setae-like structures, protibia short distally truncate, bearing two tarsomeres.</p></div>	https://treatment.plazi.org/id/F7BB43F68C925E80A97420CF1D6D8E4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Le Cesne, Maxime;Bourgoin, Thierry;Hoch, Hannelore;Luo, Yang;Zhang, Yalin	Le Cesne, Maxime, Bourgoin, Thierry, Hoch, Hannelore, Luo, Yang, Zhang, Yalin (2022): Coframalaxius bletteryi gen. et sp. nov. from subterranean habitat in Southern France (Hemiptera, Fulgoromorpha, Cixiidae, Oecleini). Subterranean Biology 43: 145-168, DOI: http://dx.doi.org/10.3897/subtbiol.43.85804, URL: http://dx.doi.org/10.3897/subtbiol.43.85804
