taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839849/files/figure.png	http://doi.org/10.5281/zenodo.6839849	Fig. 1. Type specimens of M. lutko Evans, 1932, M. mimetica Higgins, 1940, and M. timandra Coutsis & van Oorschot, 2014. A, C, E. UPS. B, D, F. UNS. A–B. M. lutko, holotype, ♂ (BMNH). C– D. M. mimetica, paratype, ♀ (BMNH). E–F. M. timandra, holotype, ♂ (ZMA). A–D photos by A. Devyatkin; E–F, by Eulalia Gasso Miracle.	Fig. 1. Type specimens of M. lutko Evans, 1932, M. mimetica Higgins, 1940, and M. timandra Coutsis & van Oorschot, 2014. A, C, E. UPS. B, D, F. UNS. A–B. M. lutko, holotype, ♂ (BMNH). C– D. M. mimetica, paratype, ♀ (BMNH). E–F. M. timandra, holotype, ♂ (ZMA). A–D photos by A. Devyatkin; E–F, by Eulalia Gasso Miracle.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839851/files/figure.png	http://doi.org/10.5281/zenodo.6839851	Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839853/files/figure.png	http://doi.org/10.5281/zenodo.6839853	Fig. 3. Male genitalia (a = valva; b = aedeagus). A–G. Melitaea mimetica Higgins, 1940. H–I. M. lutko Evans, 1932. A. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. B. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C. Central Afghanistan, Ghor Prov., 17 km E Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. D. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt. 2700–2800 m. F. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. G. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. H. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. I. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m.	Fig. 3. Male genitalia (a = valva; b = aedeagus). A–G. Melitaea mimetica Higgins, 1940. H–I. M. lutko Evans, 1932. A. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. B. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C. Central Afghanistan, Ghor Prov., 17 km E Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. D. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt. 2700–2800 m. F. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. G. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. H. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. I. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839857/files/figure.png	http://doi.org/10.5281/zenodo.6839857	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839859/files/figure.png	http://doi.org/10.5281/zenodo.6839859	Fig. 5. Distribution of M. lutko Evans, 1932. For a description of the dots with letters see Fig. 30.	Fig. 5. Distribution of M. lutko Evans, 1932. For a description of the dots with letters see Fig. 30.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839861/files/figure.png	http://doi.org/10.5281/zenodo.6839861	Fig. 6. Adults of M. mimetica Higgins, 1940. A–H. UPS. I–P. UNS. A–B, I–J. ♂, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C, K. ♂, Afghanistan, Ghowr Prov., Bayan Range, 15 km S of Chagcharan, Kindival vall. D, L. ♂, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. E–F, M–N. ♀, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. G, O. ♀, Central Afghanistan, Bamian Prov., 8 km S of Bamian, Koh-e-Baba Mts, Dara-e-Khushkak, alt. 2930 m. H, P. ♀, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. B, J photos by Dr Eckweiler).	Fig. 6. Adults of M. mimetica Higgins, 1940. A–H. UPS. I–P. UNS. A–B, I–J. ♂, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C, K. ♂, Afghanistan, Ghowr Prov., Bayan Range, 15 km S of Chagcharan, Kindival vall. D, L. ♂, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. E–F, M–N. ♀, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. G, O. ♀, Central Afghanistan, Bamian Prov., 8 km S of Bamian, Koh-e-Baba Mts, Dara-e-Khushkak, alt. 2930 m. H, P. ♀, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. B, J photos by Dr Eckweiler).	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839863/files/figure.png	http://doi.org/10.5281/zenodo.6839863	Fig. 7. Eggs of Melitaea mimetica Higgins, 1940, Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	Fig. 7. Eggs of Melitaea mimetica Higgins, 1940, Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839865/files/figure.png	http://doi.org/10.5281/zenodo.6839865	Fig. 8. Distribution of Melitaea mimetica Higgins, 1940. For a description of the symbols with letters, see Fig. 30. = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954.	Fig. 8. Distribution of Melitaea mimetica Higgins, 1940. For a description of the symbols with letters, see Fig. 30. = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839867/files/figure.png	http://doi.org/10.5281/zenodo.6839867	Fig. 9. Adults of Melitaea timandra timandra Coutsis & van Oorschot, 2014. A–H. UPS. I–P. UNS. A–D. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. E–H. ♀, Turkmenistan, Sary-Yazy, alt. 300 m. I–L. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. M–P. ♀, Turkmenistan, Sary-Yazy, alt. 300 m.	Fig. 9. Adults of Melitaea timandra timandra Coutsis & van Oorschot, 2014. A–H. UPS. I–P. UNS. A–D. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. E–H. ♀, Turkmenistan, Sary-Yazy, alt. 300 m. I–L. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. M–P. ♀, Turkmenistan, Sary-Yazy, alt. 300 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839869/files/figure.png	http://doi.org/10.5281/zenodo.6839869	Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis & van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis & van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839871/files/figure.png	http://doi.org/10.5281/zenodo.6839871	Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014, S Turkmenistan, SaryYazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-eBinalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014, S Turkmenistan, SaryYazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-eBinalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839877/files/figure.png	http://doi.org/10.5281/zenodo.6839877	Fig. 12. Eggs of Melitaea timandra timandra Coutsis & van Oorschot, 2014, S. Turkmenistan, SaryYazy. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	Fig. 12. Eggs of Melitaea timandra timandra Coutsis & van Oorschot, 2014, S. Turkmenistan, SaryYazy. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839879/files/figure.png	http://doi.org/10.5281/zenodo.6839879	Fig. 13. Biotopes and biology of Melitaea timandra timandraCoutsis & van Oorschot, 2014.A. Bumpycellular overgrown sands on the left bank of the Murgab River, alt. 300 m, April 1993, the habitat of M. timandra timandra. B. Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm. – host plant for M. timandra timandra. The female laying eggs is visible under the leaves of the plant. C. The female of M. timandra timandra is ovipositing on the soil under the host plant leaf. D. Egg clutch of M. timandra timandra on the basal leaf rosette of Ph. regeliana.	Fig. 13. Biotopes and biology of Melitaea timandra timandraCoutsis & van Oorschot, 2014.A. Bumpycellular overgrown sands on the left bank of the Murgab River, alt. 300 m, April 1993, the habitat of M. timandra timandra. B. Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm. – host plant for M. timandra timandra. The female laying eggs is visible under the leaves of the plant. C. The female of M. timandra timandra is ovipositing on the soil under the host plant leaf. D. Egg clutch of M. timandra timandra on the basal leaf rosette of Ph. regeliana.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839881/files/figure.png	http://doi.org/10.5281/zenodo.6839881	Fig. 14. Distribution of Melitaea timandra timandra Coutsis & van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status.	Fig. 14. Distribution of Melitaea timandra timandra Coutsis & van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839883/files/figure.png	http://doi.org/10.5281/zenodo.6839883	Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding.	Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839885/files/figure.png	http://doi.org/10.5281/zenodo.6839885	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839887/files/figure.png	http://doi.org/10.5281/zenodo.6839887	Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, SaryYazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, SaryYazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839891/files/figure.png	http://doi.org/10.5281/zenodo.6839891	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839893/files/figure.png	http://doi.org/10.5281/zenodo.6839893	Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.	Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839895/files/figure.png	http://doi.org/10.5281/zenodo.6839895	Fig. 20. Eggs of Melitaea timandra binaludica subsp. nov., Iran, Kuh-e-Binalud Mts. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	Fig. 20. Eggs of Melitaea timandra binaludica subsp. nov., Iran, Kuh-e-Binalud Mts. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839897/files/figure.png	http://doi.org/10.5281/zenodo.6839897	Fig. 21. Adults of Melitaea shahvarica sp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–C. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m. D. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., Tohar v. vicinity, alt. 2200 m. E–H. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200– 2400 m. I–K. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., 2200–2400 m. L. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, Tohar v. vicinity, alt. 2200 m. M–P. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m.	Fig. 21. Adults of Melitaea shahvarica sp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–C. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m. D. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., Tohar v. vicinity, alt. 2200 m. E–H. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200– 2400 m. I–K. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., 2200–2400 m. L. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, Tohar v. vicinity, alt. 2200 m. M–P. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839899/files/figure.png	http://doi.org/10.5281/zenodo.6839899	Fig. 22. Male genitalia of Melitaea shahvarica sp. nov. (a = valva; b = aedeagus). Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m.	Fig. 22. Male genitalia of Melitaea shahvarica sp. nov. (a = valva; b = aedeagus). Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839901/files/figure.png	http://doi.org/10.5281/zenodo.6839901	Fig. 23. Eggs of Melitaea shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	Fig. 23. Eggs of Melitaea shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839905/files/figure.png	http://doi.org/10.5281/zenodo.6839905	Fig. 24. First instar caterpillar of Melitaea shahvarica sp. nov. A. Head, bottom view. B. Head, front view. C–D. Head, lateral view. E–F. Caterpillar, lateral view.	Fig. 24. First instar caterpillar of Melitaea shahvarica sp. nov. A. Head, bottom view. B. Head, front view. C–D. Head, lateral view. E–F. Caterpillar, lateral view.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839907/files/figure.png	http://doi.org/10.5281/zenodo.6839907	Fig. 25. I–IV instar caterpillars of Melitaea shahvarica sp. nov. (a = view from above; b = lateral view). A. First instar caterpillar after hatching. B. First instar caterpillar before molting. C. Second instar caterpillar. D. Third instar caterpillar. E. Fourth instar caterpillar.	Fig. 25. I–IV instar caterpillars of Melitaea shahvarica sp. nov. (a = view from above; b = lateral view). A. First instar caterpillar after hatching. B. First instar caterpillar before molting. C. Second instar caterpillar. D. Third instar caterpillar. E. Fourth instar caterpillar.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839909/files/figure.png	http://doi.org/10.5281/zenodo.6839909	Fig. 26. V–VI instar caterpillars of Melitaea shahvarica sp. nov. (a = view from above; b = lateral view). A. Fifth instar caterpillar. B. Sixth instar caterpillar. C. Sixth instar caterpillar, head capsule, front view. D. Sixth instar caterpillar, head capsule, lateral view.	Fig. 26. V–VI instar caterpillars of Melitaea shahvarica sp. nov. (a = view from above; b = lateral view). A. Fifth instar caterpillar. B. Sixth instar caterpillar. C. Sixth instar caterpillar, head capsule, front view. D. Sixth instar caterpillar, head capsule, lateral view.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839911/files/figure.png	http://doi.org/10.5281/zenodo.6839911	Fig. 27.Biotope, host plant, and adult of Melitaea shahvaricasp. nov. in nature. A. Shahvar Mt., E Elburs Ridge. B. Biotope with Phlomoides molucelloides (Bunge) Salmaki. C. Host plant Ph. molucelloides. D. Female on Ph. molucelloides.	Fig. 27.Biotope, host plant, and adult of Melitaea shahvaricasp. nov. in nature. A. Shahvar Mt., E Elburs Ridge. B. Biotope with Phlomoides molucelloides (Bunge) Salmaki. C. Host plant Ph. molucelloides. D. Female on Ph. molucelloides.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839913/files/figure.png	http://doi.org/10.5281/zenodo.6839913	Fig. 28.Eggs and caterpillars of Melitaea shahvarica sp. nov. in nature and in the laboratory.A–B. Freshly laid eggs under a leaf of a host plant, May 2018, Iran, Shahvar Mt., alt. 2200 m. C–D. IV–V instar caterpillars on the leaves of the host plant Phlomoides molucelloides (Bunge) Salmaki, July 2019, Iran, Shahvar Mt., alt. 2500 m. E. I instar caterpillars in the laboratory, Moscow, May 2018. F. VI instar caterpillars during diapause, Moscow, October 2018.	Fig. 28.Eggs and caterpillars of Melitaea shahvarica sp. nov. in nature and in the laboratory.A–B. Freshly laid eggs under a leaf of a host plant, May 2018, Iran, Shahvar Mt., alt. 2200 m. C–D. IV–V instar caterpillars on the leaves of the host plant Phlomoides molucelloides (Bunge) Salmaki, July 2019, Iran, Shahvar Mt., alt. 2500 m. E. I instar caterpillars in the laboratory, Moscow, May 2018. F. VI instar caterpillars during diapause, Moscow, October 2018.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839915/files/figure.png	http://doi.org/10.5281/zenodo.6839915	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD3B3279147F896FBF1F8B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6841901/files/figure.png	http://doi.org/10.5281/zenodo.6841901	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839849/files/figure.png	http://doi.org/10.5281/zenodo.6839849	Fig. 1. Type specimens of M. lutko Evans, 1932, M. mimetica Higgins, 1940, and M. timandra Coutsis & van Oorschot, 2014. A, C, E. UPS. B, D, F. UNS. A–B. M. lutko, holotype, ♂ (BMNH). C– D. M. mimetica, paratype, ♀ (BMNH). E–F. M. timandra, holotype, ♂ (ZMA). A–D photos by A. Devyatkin; E–F, by Eulalia Gasso Miracle.	Fig. 1. Type specimens of M. lutko Evans, 1932, M. mimetica Higgins, 1940, and M. timandra Coutsis & van Oorschot, 2014. A, C, E. UPS. B, D, F. UNS. A–B. M. lutko, holotype, ♂ (BMNH). C– D. M. mimetica, paratype, ♀ (BMNH). E–F. M. timandra, holotype, ♂ (ZMA). A–D photos by A. Devyatkin; E–F, by Eulalia Gasso Miracle.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839851/files/figure.png	http://doi.org/10.5281/zenodo.6839851	Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839853/files/figure.png	http://doi.org/10.5281/zenodo.6839853	Fig. 3. Male genitalia (a = valva; b = aedeagus). A–G. Melitaea mimetica Higgins, 1940. H–I. M. lutko Evans, 1932. A. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. B. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C. Central Afghanistan, Ghor Prov., 17 km E Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. D. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt. 2700–2800 m. F. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. G. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. H. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. I. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m.	Fig. 3. Male genitalia (a = valva; b = aedeagus). A–G. Melitaea mimetica Higgins, 1940. H–I. M. lutko Evans, 1932. A. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. B. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C. Central Afghanistan, Ghor Prov., 17 km E Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. D. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt. 2700–2800 m. F. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. G. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. H. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. I. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839857/files/figure.png	http://doi.org/10.5281/zenodo.6839857	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839859/files/figure.png	http://doi.org/10.5281/zenodo.6839859	Fig. 5. Distribution of M. lutko Evans, 1932. For a description of the dots with letters see Fig. 30.	Fig. 5. Distribution of M. lutko Evans, 1932. For a description of the dots with letters see Fig. 30.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839885/files/figure.png	http://doi.org/10.5281/zenodo.6839885	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839891/files/figure.png	http://doi.org/10.5281/zenodo.6839891	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839915/files/figure.png	http://doi.org/10.5281/zenodo.6839915	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD2B32A916CFE8AFAA5FA63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6841901/files/figure.png	http://doi.org/10.5281/zenodo.6841901	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700–2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700–2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839849/files/figure.png	http://doi.org/10.5281/zenodo.6839849	Fig. 1. Type specimens of M. lutko Evans, 1932, M. mimetica Higgins, 1940, and M. timandra Coutsis & van Oorschot, 2014. A, C, E. UPS. B, D, F. UNS. A–B. M. lutko, holotype, ♂ (BMNH). C– D. M. mimetica, paratype, ♀ (BMNH). E–F. M. timandra, holotype, ♂ (ZMA). A–D photos by A. Devyatkin; E–F, by Eulalia Gasso Miracle.	Fig. 1. Type specimens of M. lutko Evans, 1932, M. mimetica Higgins, 1940, and M. timandra Coutsis & van Oorschot, 2014. A, C, E. UPS. B, D, F. UNS. A–B. M. lutko, holotype, ♂ (BMNH). C– D. M. mimetica, paratype, ♀ (BMNH). E–F. M. timandra, holotype, ♂ (ZMA). A–D photos by A. Devyatkin; E–F, by Eulalia Gasso Miracle.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839853/files/figure.png	http://doi.org/10.5281/zenodo.6839853	Fig. 3. Male genitalia (a = valva; b = aedeagus). A–G. Melitaea mimetica Higgins, 1940. H–I. M. lutko Evans, 1932. A. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. B. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C. Central Afghanistan, Ghor Prov., 17 km E Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. D. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt. 2700–2800 m. F. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. G. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. H. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. I. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m.	Fig. 3. Male genitalia (a = valva; b = aedeagus). A–G. Melitaea mimetica Higgins, 1940. H–I. M. lutko Evans, 1932. A. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. B. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C. Central Afghanistan, Ghor Prov., 17 km E Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. D. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt. 2700–2800 m. F. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. G. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. H. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. I. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839857/files/figure.png	http://doi.org/10.5281/zenodo.6839857	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839861/files/figure.png	http://doi.org/10.5281/zenodo.6839861	Fig. 6. Adults of M. mimetica Higgins, 1940. A–H. UPS. I–P. UNS. A–B, I–J. ♂, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C, K. ♂, Afghanistan, Ghowr Prov., Bayan Range, 15 km S of Chagcharan, Kindival vall. D, L. ♂, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. E–F, M–N. ♀, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. G, O. ♀, Central Afghanistan, Bamian Prov., 8 km S of Bamian, Koh-e-Baba Mts, Dara-e-Khushkak, alt. 2930 m. H, P. ♀, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. B, J photos by Dr Eckweiler).	Fig. 6. Adults of M. mimetica Higgins, 1940. A–H. UPS. I–P. UNS. A–B, I–J. ♂, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C, K. ♂, Afghanistan, Ghowr Prov., Bayan Range, 15 km S of Chagcharan, Kindival vall. D, L. ♂, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. E–F, M–N. ♀, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. G, O. ♀, Central Afghanistan, Bamian Prov., 8 km S of Bamian, Koh-e-Baba Mts, Dara-e-Khushkak, alt. 2930 m. H, P. ♀, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. B, J photos by Dr Eckweiler).	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839863/files/figure.png	http://doi.org/10.5281/zenodo.6839863	Fig. 7. Eggs of Melitaea mimetica Higgins, 1940, Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	Fig. 7. Eggs of Melitaea mimetica Higgins, 1940, Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839865/files/figure.png	http://doi.org/10.5281/zenodo.6839865	Fig. 8. Distribution of Melitaea mimetica Higgins, 1940. For a description of the symbols with letters, see Fig. 30. = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954.	Fig. 8. Distribution of Melitaea mimetica Higgins, 1940. For a description of the symbols with letters, see Fig. 30. = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839885/files/figure.png	http://doi.org/10.5281/zenodo.6839885	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839891/files/figure.png	http://doi.org/10.5281/zenodo.6839891	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839915/files/figure.png	http://doi.org/10.5281/zenodo.6839915	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFDDB32D9149FE8AFD5CF99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6841901/files/figure.png	http://doi.org/10.5281/zenodo.6841901	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839849/files/figure.png	http://doi.org/10.5281/zenodo.6839849	Fig. 1. Type specimens of M. lutko Evans, 1932, M. mimetica Higgins, 1940, and M. timandra Coutsis & van Oorschot, 2014. A, C, E. UPS. B, D, F. UNS. A–B. M. lutko, holotype, ♂ (BMNH). C– D. M. mimetica, paratype, ♀ (BMNH). E–F. M. timandra, holotype, ♂ (ZMA). A–D photos by A. Devyatkin; E–F, by Eulalia Gasso Miracle.	Fig. 1. Type specimens of M. lutko Evans, 1932, M. mimetica Higgins, 1940, and M. timandra Coutsis & van Oorschot, 2014. A, C, E. UPS. B, D, F. UNS. A–B. M. lutko, holotype, ♂ (BMNH). C– D. M. mimetica, paratype, ♀ (BMNH). E–F. M. timandra, holotype, ♂ (ZMA). A–D photos by A. Devyatkin; E–F, by Eulalia Gasso Miracle.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839867/files/figure.png	http://doi.org/10.5281/zenodo.6839867	Fig. 9. Adults of Melitaea timandra timandra Coutsis & van Oorschot, 2014. A–H. UPS. I–P. UNS. A–D. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. E–H. ♀, Turkmenistan, Sary-Yazy, alt. 300 m. I–L. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. M–P. ♀, Turkmenistan, Sary-Yazy, alt. 300 m.	Fig. 9. Adults of Melitaea timandra timandra Coutsis & van Oorschot, 2014. A–H. UPS. I–P. UNS. A–D. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. E–H. ♀, Turkmenistan, Sary-Yazy, alt. 300 m. I–L. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. M–P. ♀, Turkmenistan, Sary-Yazy, alt. 300 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839869/files/figure.png	http://doi.org/10.5281/zenodo.6839869	Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis & van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis & van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839871/files/figure.png	http://doi.org/10.5281/zenodo.6839871	Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014, S Turkmenistan, Sary- Yazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-e- Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014, S Turkmenistan, Sary- Yazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-e- Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839877/files/figure.png	http://doi.org/10.5281/zenodo.6839877	Fig. 12. Eggs of Melitaea timandra timandra Coutsis & van Oorschot, 2014, S. Turkmenistan, Sary- Yazy. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	Fig. 12. Eggs of Melitaea timandra timandra Coutsis & van Oorschot, 2014, S. Turkmenistan, Sary- Yazy. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839879/files/figure.png	http://doi.org/10.5281/zenodo.6839879	Fig. 13. Biotopes and biology of Melitaea timandra timandraCoutsis & van Oorschot, 2014.A. Bumpycellular overgrown sands on the left bank of the Murgab River, alt. 300 m, April 1993, the habitat of M. timandra timandra. B. Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm. – host plant for M. timandra timandra. The female laying eggs is visible under the leaves of the plant. C. The female of M. timandra timandra is ovipositing on the soil under the host plant leaf. D. Egg clutch of M. timandra timandra on the basal leaf rosette of Ph. regeliana.	Fig. 13. Biotopes and biology of Melitaea timandra timandraCoutsis & van Oorschot, 2014.A. Bumpycellular overgrown sands on the left bank of the Murgab River, alt. 300 m, April 1993, the habitat of M. timandra timandra. B. Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm. – host plant for M. timandra timandra. The female laying eggs is visible under the leaves of the plant. C. The female of M. timandra timandra is ovipositing on the soil under the host plant leaf. D. Egg clutch of M. timandra timandra on the basal leaf rosette of Ph. regeliana.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839887/files/figure.png	http://doi.org/10.5281/zenodo.6839887	Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary- Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary- Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839893/files/figure.png	http://doi.org/10.5281/zenodo.6839893	Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.	Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839915/files/figure.png	http://doi.org/10.5281/zenodo.6839915	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6841901/files/figure.png	http://doi.org/10.5281/zenodo.6841901	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFD8B33492A3FE8AFC4EFA38.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839881/files/figure.png	http://doi.org/10.5281/zenodo.6839881	Fig. 14. Distribution of Melitaea timandra timandra Coutsis & van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status.	Fig. 14. Distribution of Melitaea timandra timandra Coutsis & van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839851/files/figure.png	http://doi.org/10.5281/zenodo.6839851	Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839869/files/figure.png	http://doi.org/10.5281/zenodo.6839869	Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis & van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis & van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839871/files/figure.png	http://doi.org/10.5281/zenodo.6839871	Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014, S Turkmenistan, Sary- Yazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-e- Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014, S Turkmenistan, Sary- Yazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-e- Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839881/files/figure.png	http://doi.org/10.5281/zenodo.6839881	Fig. 14. Distribution of Melitaea timandra timandra Coutsis & van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status.	Fig. 14. Distribution of Melitaea timandra timandra Coutsis & van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839883/files/figure.png	http://doi.org/10.5281/zenodo.6839883	Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding.	Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839887/files/figure.png	http://doi.org/10.5281/zenodo.6839887	Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary- Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary- Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839893/files/figure.png	http://doi.org/10.5281/zenodo.6839893	Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.	Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839895/files/figure.png	http://doi.org/10.5281/zenodo.6839895	Fig. 20. Eggs of Melitaea timandra binaludica subsp. nov., Iran, Kuh-e-Binalud Mts. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	Fig. 20. Eggs of Melitaea timandra binaludica subsp. nov., Iran, Kuh-e-Binalud Mts. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839915/files/figure.png	http://doi.org/10.5281/zenodo.6839915	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC0B33C913AFAF8FADBFD92.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6841901/files/figure.png	http://doi.org/10.5281/zenodo.6841901	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839857/files/figure.png	http://doi.org/10.5281/zenodo.6839857	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839881/files/figure.png	http://doi.org/10.5281/zenodo.6839881	Fig. 14. Distribution of Melitaea timandra timandra Coutsis & van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status.	Fig. 14. Distribution of Melitaea timandra timandra Coutsis & van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839885/files/figure.png	http://doi.org/10.5281/zenodo.6839885	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839891/files/figure.png	http://doi.org/10.5281/zenodo.6839891	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839897/files/figure.png	http://doi.org/10.5281/zenodo.6839897	Fig. 21. Adults of Melitaea shahvarica sp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–C. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m. D. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., Tohar v. vicinity, alt. 2200 m. E–H. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200– 2400 m. I–K. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., 2200–2400 m. L. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, Tohar v. vicinity, alt. 2200 m. M–P. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m.	Fig. 21. Adults of Melitaea shahvarica sp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–C. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m. D. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., Tohar v. vicinity, alt. 2200 m. E–H. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200– 2400 m. I–K. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., 2200–2400 m. L. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, Tohar v. vicinity, alt. 2200 m. M–P. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839899/files/figure.png	http://doi.org/10.5281/zenodo.6839899	Fig. 22. Male genitalia of Melitaea shahvarica sp. nov. (a = valva; b = aedeagus). Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m.	Fig. 22. Male genitalia of Melitaea shahvarica sp. nov. (a = valva; b = aedeagus). Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839901/files/figure.png	http://doi.org/10.5281/zenodo.6839901	Fig. 23. Eggs of Melitaea shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	Fig. 23. Eggs of Melitaea shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. A–C. Lateral view. D–F. View from above. G–I. Micropile area.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839905/files/figure.png	http://doi.org/10.5281/zenodo.6839905	Fig. 24. First instar caterpillar of Melitaea shahvarica sp. nov. A. Head, bottom view. B. Head, front view. C–D. Head, lateral view. E–F. Caterpillar, lateral view.	Fig. 24. First instar caterpillar of Melitaea shahvarica sp. nov. A. Head, bottom view. B. Head, front view. C–D. Head, lateral view. E–F. Caterpillar, lateral view.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839907/files/figure.png	http://doi.org/10.5281/zenodo.6839907	Fig. 25. I–IV instar caterpillars of Melitaea shahvarica sp. nov. (a = view from above; b = lateral view). A. First instar caterpillar after hatching. B. First instar caterpillar before molting. C. Second instar caterpillar. D. Third instar caterpillar. E. Fourth instar caterpillar.	Fig. 25. I–IV instar caterpillars of Melitaea shahvarica sp. nov. (a = view from above; b = lateral view). A. First instar caterpillar after hatching. B. First instar caterpillar before molting. C. Second instar caterpillar. D. Third instar caterpillar. E. Fourth instar caterpillar.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839909/files/figure.png	http://doi.org/10.5281/zenodo.6839909	Fig. 26. V–VI instar caterpillars of Melitaea shahvarica sp. nov. (a = view from above; b = lateral view). A. Fifth instar caterpillar. B. Sixth instar caterpillar. C. Sixth instar caterpillar, head capsule, front view. D. Sixth instar caterpillar, head capsule, lateral view.	Fig. 26. V–VI instar caterpillars of Melitaea shahvarica sp. nov. (a = view from above; b = lateral view). A. Fifth instar caterpillar. B. Sixth instar caterpillar. C. Sixth instar caterpillar, head capsule, front view. D. Sixth instar caterpillar, head capsule, lateral view.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839911/files/figure.png	http://doi.org/10.5281/zenodo.6839911	Fig. 27.Biotope, host plant, and adult of Melitaea shahvaricasp. nov. in nature. A. Shahvar Mt., E Elburs Ridge. B. Biotope with Phlomoides molucelloides (Bunge) Salmaki. C. Host plant Ph. molucelloides. D. Female on Ph. molucelloides.	Fig. 27.Biotope, host plant, and adult of Melitaea shahvaricasp. nov. in nature. A. Shahvar Mt., E Elburs Ridge. B. Biotope with Phlomoides molucelloides (Bunge) Salmaki. C. Host plant Ph. molucelloides. D. Female on Ph. molucelloides.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839913/files/figure.png	http://doi.org/10.5281/zenodo.6839913	Fig. 28.Eggs and caterpillars of Melitaea shahvarica sp. nov. in nature and in the laboratory.A–B. Freshly laid eggs under a leaf of a host plant, May 2018, Iran, Shahvar Mt., alt. 2200 m. C–D. IV–V instar caterpillars on the leaves of the host plant Phlomoides molucelloides (Bunge) Salmaki, July 2019, Iran, Shahvar Mt., alt. 2500 m. E. I instar caterpillars in the laboratory, Moscow, May 2018. F. VI instar caterpillars during diapause, Moscow, October 2018.	Fig. 28.Eggs and caterpillars of Melitaea shahvarica sp. nov. in nature and in the laboratory.A–B. Freshly laid eggs under a leaf of a host plant, May 2018, Iran, Shahvar Mt., alt. 2200 m. C–D. IV–V instar caterpillars on the leaves of the host plant Phlomoides molucelloides (Bunge) Salmaki, July 2019, Iran, Shahvar Mt., alt. 2500 m. E. I instar caterpillars in the laboratory, Moscow, May 2018. F. VI instar caterpillars during diapause, Moscow, October 2018.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839915/files/figure.png	http://doi.org/10.5281/zenodo.6839915	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFC8B30E9174FCA2FAA5F8D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6841901/files/figure.png	http://doi.org/10.5281/zenodo.6841901	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis & Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. & Hemsl.) Adylov, Kamelin & Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin & Makhm.; = Phlomoides laciniata (L.) Kamelin & Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin & Makhm.; = Phlomoides loasifolia (Benth.) Kamelin & Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin & Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839891/files/figure.png	http://doi.org/10.5281/zenodo.6839891	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	Fig. 18. Male genitalia and harpe. A–C. Melitaea shahvarica sp. nov. D–E. M. lutko Evans, 1932. F–I. M. mimetica Higgins, 1940. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D–E. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m. F–G. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m. H–I. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839885/files/figure.png	http://doi.org/10.5281/zenodo.6839885	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	Fig. 16. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea shahvarica sp. nov. D–H. M. mimetica Higgins, 1940. I. M. lutko Evans, 1932. A–C. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. D. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt, 2700– 2800 m. F. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. G. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. H. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali v. vicinity, alt. 2400 m. I. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839915/files/figure.png	http://doi.org/10.5281/zenodo.6839915	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839861/files/figure.png	http://doi.org/10.5281/zenodo.6839861	Fig. 6. Adults of M. mimetica Higgins, 1940. A–H. UPS. I–P. UNS. A–B, I–J. ♂, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C, K. ♂, Afghanistan, Ghowr Prov., Bayan Range, 15 km S of Chagcharan, Kindival vall. D, L. ♂, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. E–F, M–N. ♀, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. G, O. ♀, Central Afghanistan, Bamian Prov., 8 km S of Bamian, Koh-e-Baba Mts, Dara-e-Khushkak, alt. 2930 m. H, P. ♀, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. B, J photos by Dr Eckweiler).	Fig. 6. Adults of M. mimetica Higgins, 1940. A–H. UPS. I–P. UNS. A–B, I–J. ♂, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C, K. ♂, Afghanistan, Ghowr Prov., Bayan Range, 15 km S of Chagcharan, Kindival vall. D, L. ♂, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. E–F, M–N. ♀, Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. G, O. ♀, Central Afghanistan, Bamian Prov., 8 km S of Bamian, Koh-e-Baba Mts, Dara-e-Khushkak, alt. 2930 m. H, P. ♀, Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. B, J photos by Dr Eckweiler).	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839893/files/figure.png	http://doi.org/10.5281/zenodo.6839893	Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.	Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839887/files/figure.png	http://doi.org/10.5281/zenodo.6839887	Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary- Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary- Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839853/files/figure.png	http://doi.org/10.5281/zenodo.6839853	Fig. 3. Male genitalia (a = valva; b = aedeagus). A–G. Melitaea mimetica Higgins, 1940. H–I. M. lutko Evans, 1932. A. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. B. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C. Central Afghanistan, Ghor Prov., 17 km E Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. D. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt. 2700–2800 m. F. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. G. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. H. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. I. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m.	Fig. 3. Male genitalia (a = valva; b = aedeagus). A–G. Melitaea mimetica Higgins, 1940. H–I. M. lutko Evans, 1932. A. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. B. Pakistan, Balochistan, Quetta, Urak, alt. 2400–2700 m. C. Central Afghanistan, Ghor Prov., 17 km E Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. D. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. E. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak v. vicinity, alt. 2700–2800 m. F. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. G. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. H. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. I. Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839851/files/figure.png	http://doi.org/10.5281/zenodo.6839851	Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839869/files/figure.png	http://doi.org/10.5281/zenodo.6839869	Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis & van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis & van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839899/files/figure.png	http://doi.org/10.5281/zenodo.6839899	Fig. 22. Male genitalia of Melitaea shahvarica sp. nov. (a = valva; b = aedeagus). Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m.	Fig. 22. Male genitalia of Melitaea shahvarica sp. nov. (a = valva; b = aedeagus). Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839867/files/figure.png	http://doi.org/10.5281/zenodo.6839867	Fig. 9. Adults of Melitaea timandra timandra Coutsis & van Oorschot, 2014. A–H. UPS. I–P. UNS. A–D. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. E–H. ♀, Turkmenistan, Sary-Yazy, alt. 300 m. I–L. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. M–P. ♀, Turkmenistan, Sary-Yazy, alt. 300 m.	Fig. 9. Adults of Melitaea timandra timandra Coutsis & van Oorschot, 2014. A–H. UPS. I–P. UNS. A–D. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. E–H. ♀, Turkmenistan, Sary-Yazy, alt. 300 m. I–L. ♂, Turkmenistan, Sary-Yazy, alt. 300 m. M–P. ♀, Turkmenistan, Sary-Yazy, alt. 300 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839883/files/figure.png	http://doi.org/10.5281/zenodo.6839883	Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding.	Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839897/files/figure.png	http://doi.org/10.5281/zenodo.6839897	Fig. 21. Adults of Melitaea shahvarica sp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–C. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m. D. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., Tohar v. vicinity, alt. 2200 m. E–H. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200– 2400 m. I–K. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., 2200–2400 m. L. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, Tohar v. vicinity, alt. 2200 m. M–P. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m.	Fig. 21. Adults of Melitaea shahvarica sp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–C. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m. D. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., Tohar v. vicinity, alt. 2200 m. E–H. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200– 2400 m. I–K. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., 2200–2400 m. L. ♂. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, Tohar v. vicinity, alt. 2200 m. M–P. ♀. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mt., alt. 2200–2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839857/files/figure.png	http://doi.org/10.5281/zenodo.6839857	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	Fig. 4. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–D. M. shahvarica sp. nov., Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. E. M. lutko Evans, 1932, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F–I. M. mimetica Higgins, 1940. F. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. G–H. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. I. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
EB42F330FFF9B31393A7FE8BFAA5FC8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6839871/files/figure.png	http://doi.org/10.5281/zenodo.6839871	Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014, S Turkmenistan, Sary- Yazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-e- Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis & van Oorschot, 2014, S Turkmenistan, Sary- Yazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-e- Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.	2022-07-15	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.		Zenodo	biologists	Kolesnichenko, Kirill A.;Kotlobay, Anatoly A.			
