taxonID	type	description	language	source
FA188782FF87FFED1283FB2B7BC7FE3C.taxon	description	(Figs 1 A – F, 2, 3, 12 A)	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFED1283FB2B7BC7FE3C.taxon	description	Redescription. Body elongate; head, thorax and abdomen dark brown; antennae dark reddish brown, with basal part paler; mouth parts and legs reddish yellow; elytra reddish yellow with posterolateral portion darker; head, thorax, abdominal sternites, elytra, and legs covered with setae rather densely; abdominal tergites sparsely covered with setae; dorsal surface of head and pronotum densely covered with reticulation; elytra weakly rugose; abdominal tergites glossy. Head circular, slightly longer than or as long as wide; postocular parts roundly narrowed toward base; eyes ovate. Antennae slender, longer than a combination of head and pronotum; segments IV – V slightly longer than wide; segments VI – VII as long as wide; segments VIII – IX slightly transverse; segments X as long as wide; segments XI ovate. Pronotum almost hexagonal, slightly wider than long; anterior margin widely rounded; lateral margin moderately rounded, narrowed more strongly to posterior than anterior; anterior corners rounded; posterior corners obtusely roundly angulated; hypomera visible in lateral view, with posterior part of carina (Fig. 12 A) directed posteriorly almost parallel to lateral margin. Mesoventrite (Fig. 2 A) shortly carinate at base of midline; inner coxal process narrow, elongate, reaching to metaventrite process. Metaventrite (Fig. 2 A) with inner coxal process short, moderately narrow. Elytra wider than long, longer and wider than pronotum, weakly diverging to posteriorly, widest near posterolateral corners; hind margin sinuate near posterolateral corners. Hind wings developed. Abdomen moderately elongate, parallel sided; tergites III – VII with small pores at basal impression; VIII tergite (Fig. 2 B) with posterior margin smooth, shallowly emarginate medially in both sexes. VIII sternum with posterior margin smooth, widely rounded in both sexes (Fig. 2 C). Legs slender, rather long (HTL / PL = 1.13 – 1.20); surface of fore and middle tibia not covered with spines; hind tarsal segments I as long as or slightly longer than a combination of II and III. Male: Median lobe of aedeagus (Fig. 1 B – E) rather small; apical lobe dorso-laterally rather thick, weakly curved dorsally at apex, bisinuate at ventral margin, dilated laterally at apical, rather strongly constricted at middle in ventral view; sclerites 1 (sc 1) with posterior margin strongly convex and curved laterally; anterior margin widely emarginate. Female: Spermatheca (Fig. 1 F) with capsule spherical. Measurements. BL 4.3 – 4.9, FBL 2.60 – 2.77, HW 0.68 – 0.70, HL 0.72 – 0.75, PW 0.85 – 0.97, PL 0.83 – 0.86, EW 1.15 – 1.39, EL 1.01 – 1.15, HTL 0.92 – 1.01.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFED1283FB2B7BC7FE3C.taxon	materials_examined	Type material. Ocalea japonica: lectotype, here designated, male, “ Type [RRL] // Japan (YRLS) // Japan. / G. Lewis // Sharp Coll. // 1905 - 313. // Ocalea / japonica / type D. S. // Lectotype / Ocalea japonica / Sharp, 1874 / des. Maruyama, 2011 ” (abdominal segments VIII-X and aedeagus were dissected and mounted in Euparal by MM) (PL, 0.85; PW, 0.86; HTL, 1.07) (BMNH). Specimens examined. JAPAN: Honshu: [Gunma Pref.]: 2 unsexed, Nishiyama-rindô, Sôri, Azuma-mura, Seta-gun, 21. V. 2005, T. Horiguchi leg. (KUM); 1 unsexed, Nagamine-shizen koen, Ikaho-chô., 13 – 18. IV. 2003, K. Kanai leg. (KUM); 1 male, Ôya, Annaka-shi, 22. IV. 2003, T Watanabe leg. (KUM); 1 female, 3 unsexed, Motojuku, Agatsuma-machi, 15. V. 2005, T. Watanabe leg. (KUM); [Saitama Pref.]: 1 male, Irikawa, Ôtaki, Chichibu-shi, 3. XI. 1991, M. Uchida leg. (KUM); 1 female, 1 unsexed, Mt. Kumakura-san, Chichibu-shi, 3. XI. 1990, H. Oda leg. (KUM); 3 unsexed, Kuroyama-santaki, Ogose-machi, Iruma-shi, 3. IV. 1983, H. Oda leg. (KUM); [Tokyo Met.]: 1 unsexed, Mt. Takao-san, Hachiôji-shi, 16. IV. 1962. Y. Shibata leg. (KUM); 1 unsexed, ditto, 24. V. 1963, Y. Shibata leg. (KUM); 1 unsexed, Setagaya-ku, 25. II. 1960, Y. Shibata leg. (KUM); 1 female, 1 unsexed, Machida-shi, 29. XI. 1986, M. Tao leg. (KUM); 1 male, Mt. Mitake-san, Ôme-shi, 29. VI. 1962, Y. Shibata leg. (KUM); [Chiba Pref.]: 2 males, 10 unsexed, Yata, Shiroi-shi, 14. XI. 1989, T. Takeda leg. (KUM); 1 unsexed, Koura, Onjuku-machi, 23. XII. 1989, T. Takeda leg. (KUM); 2 unsexed, Mt. Nagara, Nagara-machi, 14. XI. 1992, T. Takeda leg. (KUM); 1 unsexed, Kuramochi, Chônan-machi, 4. XI. 1992, T. Takeda leg. (KUM); 2 unsexed, ditto, 23. XI. 1992, T. Takeda leg. (KUM); 1 unsexed, Mineoka-asama-yama, Kamogawa-shi, 27. IV. 1997, S. Nomura leg. (KUM); 6 unsexed, Niaorikawa, Kiwadabata, Kimitsu-shi, 13. III – 5. VI. 2013, A. Saito leg., by underground trap (NHMC); 5 unsexed, Honzawa. Kiyosumi, Kamogawa-shi, 4. XII. 2013 – 22. V. 2014, A. Saito leg. (NHMC); 1 unsexed, Maezawa Hodou, Kiwadabata, Kimitsu-shi, 17. IV. 2013, A. Saito leg. (NHMC); [Kanagawa Pref.]: 1 unsexed, Jike-chô, Aoba-ku, Yokohama-shi, 23. XI. 2015, H. Ooki leg. (KUM); 1 unsexed, Motoishikawa, Aoba-ku, Yokohama-shi, 24. III. 2019, H. Ooki leg. (KUM); 1 unsexed, ditto, 1. III. 2021, H. Ooki leg. (KUM); 1 unsexed, Kurogane-chô, Aoba-ku, Yokohama-shi, 14. III. 2014, H. Ooki leg. (KUM); 3 unsexed, Niiharu-chô, Midori-ku, Yokohama-chô, 19. XI. 2010, H. Ooki leg. (KUM); 1 unsexed, ditto, 20. XI. 2011, H. Ooki leg. (KUM); 1 unsexed, Miho, Midori-ku, Yokohama-shi, 27. XI. 2013, H. Ooki leg. (KUM); 1 unsexed, Midori-ku, Yokohama-shi, 5. V. 1987, M. Tao leg.; 1 unsexed, Sengenyama, Nakai-machi, 7. XII. 2005, T. Watanabe leg. (KUM); 2 unsexed, Mt. Kôbo-yama, Hadano-shi, 15. XI. 1989, T. Watanabe leg. (KUM); 2 unsexed, near Bodai, Tanzawa, Hadano-shi, 29. IV. 1970, Y. Shibata leg. (KUM); 1 unsexed, Near Ôkura, Tanzawa, Hadano-shi, 29. IV. 1973, Y. Shibata leg. (KUM); 1 male, 3 unsexed, Mt. Ishihodoyama, Nishi-tanzawa, Yamakita-machi, 9. X. 1995, T. Watanabe leg. (KUM); 2 male, 7 unsexed, Fudakake, Tanzawa, Kiyokawa-mura, 6. XI. 1993, T. Watanabe leg., from animal dropping (KUM); 5 unsexed, ditto, 19. X. 2000, T. Watanabe leg. (KUM); 1 unsexed, Dôdaira, Tanzawa, Kiyokawa-mura, 10. X. 1987, Y. Hirano leg. (KUM); 1 unsexed, Near Dohdaira, Tanzawa, Kiyokawa-mura, 10. X. 1987, Y. Shibata leg. (KUM); 2 unsexed, Mt. Ôyama, 6. XI. 1960, Y. Shibata leg. (KUM); 1 unsexed, ditto, 23. XI. 1967, Y. Shibata leg. (KUM); 1 unsexed, ditto, 23. X. 1969, R. Kiryu leg. (KUM); [Ishikawa Pref.]: 2 unsexed, Komagaeri, Kanazawa-shi 20. IV. 1961, Y. Hayashi leg. (KUM); 1 unsexed, 1 female, Kodatsuno, Kanawaza-shi, 13. III. 1962, Y. Hayashi leg. (KUM); [Nagano Pref.]: 1 unsexed, Utsukushi, 1. V. 1960, Y. Shibata leg.; 1 unsexed, Shimashimadani, Matsumoto-shi, 800 m, 4 – 7. V. 2008, H. Asaki leg. (KUM); 1 unsexed, ditto, 800 m, 21 – 26. V. 2008, H. Asaki leg. (KUM); 2 unsexed, ditto, 800 m, 10 – 16, X. 2008, H. Asaki leg. (KUM); [Yamanashi Pref.]: 2 unsexed, Nagasaku, Kosuge-mura, 19. X. 1990, T. Watanabe leg. (KUM); 8 unsexed, Maruno-machi, 25. IV. 1993, K. Hosoda leg. (cI); [Gifu Pref.]: 1 unsexed, Hongô-chô, Seki-shi, 17. III. 1980, Y. Takai leg. (KUM); [Shizuoka Pref.]: 1 male, 3 unsexed, Mt. Amagi-san, Higashiizu-chô, 18. V. 1959, K. Mizusawa leg. (KUM); 2 unsexed, same data, Y. Shibata leg. (KUM); 1 unsexed, ditto, 4. V. 1959, Y. Nagashima leg. (KUM); 1 female, same data, K. Ishida leg. (KUM); 1 male, Amagi-tôge pass, 27. X. 1985, S. Nomura leg. (KUM); 1 unsexed, Abe-river, 17. XI. 1984, M. Tao leg.; [Mie Pref.]: Hirakura, Tsu-shi, 20. XI. 1994, T. Ito leg., from mushroom (cI); 2 unsexed, ditto, 19. XI. 1994, K. Mizuno leg. (cI); [Kyoto Pref.]: 1 unsexed, Maizuru-shi, 11. IV. 1998, T. Ito leg. (cI); 2 unsexed, Nyou, Maizuru-shi, 5. XI. 2003, K. Yasukawa and T. Murao leg. (cI); 1 unsexed, Iwakura, Sakyô-ku, Kyoto-shi, 5. XI. 1980, T. Ogata leg. (KUM); 1 male, 1 female, Kurama, Sakyô-ku, Kyoto-shi, 1. XI. 1964, T. Ito leg. (KUM); 1 unsexed, Kibune, Sakyô-ku, Kyoto-shi, 29. IV. 1958, Y. Shibata leg. (KUM); 1 unsexed, ditto, 2. V. 1959, T. Ito leg. (KUM); 2 unsexed, ditto, 1. XI. 1964, T. Ito leg. (KUM); 1 male, Kibune, Yamashiro, 19. VI. 1929, T. Esaki leg.; 2 male, 1 female, 23 unsexed, Otokoyama, Yawata-shi, 24. II. 2011, T. Ito leg., by rotten vegetables trap (KUM, cI); 5 unsexed, Hanase, Sakyô-ku, Kyoto-shi, 26. X. 1991, T. Ito leg, from mushroom. (KUM); [Ôsaka Pref.]: 1 unsexed, Minoo-shi, 2. IV. 1959, K. Ueda leg. (KUM); 1 male, 1 female, 1 unsexed, ditto, 4. IV. 1959, K. Ueda leg. (KUM); 3 unsexed, ditto, 13. IV. 1958, Y. Hayashi leg. (KUM); 1 female, Mt. Makio-san, Izumi-shi, 23. IV. 1960, Y. Kimura leg. (KUM); 1 unsexed, Hiraoka, Sakai-shi, 26. III. 1961, Y. Hayashi leg. (KUM); [Nara Pref.]: 1 male, Kasuga, 4. XI. 1964, T. Ito leg. (KUM); 20 unsexed, Nara-park, Nara-shi, 10. XI. 2009, T. Ito leg., from mushroom (KUM); [Wakayama Pref.]: 1 male, Onoda, Kainan-shi, 13. IV. 1993, M. Kitabata leg. (KUM); 3 unsexed, Fujishirozaka, Kainan-shi, 23. III. 1993, K. Harusawa leg. (cI); 1 unsexed, Higashihonjô, Minabe-chô, 7. III. 2019, I. Matoba leg. (cI); 1 unsexed, Ôsumi, Kimino-chô, 7. IV. 1979, H. Hiramatsu leg. (cI); [Tottori Pref.]: 1 male, Mt. Daisen, 8. XI. 1977, S. Naomi leg. (KUM); [Okayama Pref.]: 1 male. 1 female, 8 unsexed, Okutsugawakeikoku, Kamo-chô, Tomata-gun, 26. X. 2003, Y. Fujitani leg. (KUM); 1 unsexed, ditto, 23. XI. 2003, Y. Fujitani leg. (KUM); 1 unsexed, ditto, 7. XII. 2003, Y. Fujitani leg. (KUM); 1 unsexed, Mumyô-dani, Tetta-chô, Atetsu-gun, 19. X. 2003, Y. Fujitani leg. (KUM); [Yamaguchi Pref.]: 1 female, 4 unsexed, Nodani, Tokuji-chô, Saba-gun, 31. III. 2004, Y. Fujitani leg. (KUM); 4 unsexed, ditto, 10. IV. 2004, Y. Fujitani leg. (KUM); 1 female, ditto, 15. IV. 2004, Y. Fujitani leg. (KUM); 1 male, 1 unsexed, ditto, 30. IV. 2004, Y. Fujitani leg. (KUM); Shikoku: [Tokushima Pref.]: 1 female, Mt. Tsurugi-san, 15 – 17. X. 1980, S. Naomi leg. (KUM); 1 female, Mt. Maruzasa-yama (alt. 1410 m), Ichiu-son, Tsurugi-machi, 11. VI – 6. VIII. 2000, M. Yoshida leg., fowl trap (KUM). [Ehime Pref.]: 1 female, 1 unsexed, Riverside of Fukumigawa river, Matsuyama-shi, 27. XI – 3. XII. 1986, M. Hiraoka leg. (KUM); 3 unsexed, ditto, 5 – 9. XI. 1986, M. Hiraoka leg. (KUM); 3 unsexed, Fukumi, Matsuyama-shi, 5 – 9. XI. 1986, M. Hiraoka leg. (KUM); Kyushu: [Fukuoka Pref.]: 32 male, 12 female, 111 unsexed, Tachibana-yama, Fukuoka-shi, 27. XI. 1983, S. Naomi leg. (KUM); 33 unsexed, ditto, 7. V. 1980, S. Naomi leg. (KUM); 3 male, 3 unsexed, Mt. Hiko-san, Soedamachi, 24. X. 1974, M. T. Chujo leg. (KUM); 1 unsexed, ditto, 4. XI. 1971, K. Takeno leg., from Omphalotus japonicus (Kawam.) Kircham. & O. K. Mill. (KUM); 1 unsexed, ditto, 19. XI. 1980, T. Goto leg. (KUM); 1 unsexed, ditto, 20. IV. 1969, K. Kanmiya leg. (KUM); 1 unsexed, Mt. Wakasugi-yama, 17. IV. 1932, K. Yasumatsu leg. (KUM); 1 unsexed, Chikushi-yabakei, Nakagawa-shi, 16. III. 1986, S. Nomura leg. (KUM); 1 unsexed, Kôrasan, Kurume-shi, 1 – 7. IV. 2011, S. Imasaka leg. (cI); [Saga Pref.]: 2 unsexed, Makishima-yama, Seto-chô, Imari-shi, 35 m, 24. II. 2021, S. Kakizoe leg., from dog dung (KUM); 2 unsexed, Yuzuriha, Mitsuse-mura, Saga-shi, 400 m, 25. II. 2021, S. Kakizoe leg., from dog dung (KUM); [Nagasaki Pref.]: 1 unsexed, Shimabara, 13. IV. 1979, S. Imasaka leg. (KUM); 2 unsexed, Senbuki, Shimabara-shi, 4. XI. 1977, S. Imasaka leg. (KUM); 1 unsexed, ditto, 27. XII. 1977, S. Imasaka leg. (KUM); 1 male, Shitonezaki, Shikamachi-chô, 1. IV. 1988, S. Nomura leg. (KUM); 1 unsexed, Suwa-shrine, Nagasaki-shi, 21. I. 1984, S. Nomura leg. (KUM); [Kumamoto Pref.]: 1 male, 1 female, 2 unsexed, Ueki-chô, 22. XI. 1988, S. Naomi leg. (KUM); 1 unsexed, Aso-shimizu-dani, 27. X. 1993, Y. Tomishima leg. (KUM); 1 unsexed, Mt. Daikan-zan, Seiwa-mura, 3. XI. 1994, I. Ohtsuka leg. (KUM); 1 unsexed, Tenshu-zan, Yabe-chô, 17. X. 1994, Y. Tomishima leg. (KUM); 1 unsexed, Naidaijin, Yamato-machi, 22. X. 1982, M. Matsuzaki leg. (KUM); 1 unsexed, Mt. Ichifusa-yama, Minakami-mura, 31. X. 1982, M. Matsuzaki leg. (KUM); 1 unsexed, ditto, 13. X. 1995, Y. Tomishima leg. (KUM); [Ôita Pref.]: 1 unsexed, Taiko-bashi, Kadota, Takeda-shi, 12. IV. 2008, T. Miyazaki leg. (KUM); 1 female, Taiko-bashi-no-ana cave, Kadota, Takeda-shi, 322 m, 26. IV – 10. V. 2008, K. Ito leg. (KUM); 1 female, Kawanishi, Yufuin-chô, 6. XII. 2003, T. Miyazaki leg. (KUM); 1 unsexed, Shin-yabakei, Nakatsu-shi, 30. III. 1985, S. Nomura leg. (KUM); 9 unsexed, Mt. Kurotake, Yufu-shi, 1. XI. 2021, S. Kakizoe leg., from rotten mushroom (KUM); [Miyazaki Pref.]: 1 unsexed, Mt. Takachiho-no-mine, 24. X. 1977, M. T. Chujo leg. (KUM).	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFED1283FB2B7BC7FE3C.taxon	distribution	Distribution. Japan: Honshu, Shikoku, and Kyushu (Fig. 3). Bionomics. The collecting records show that Plesiochara japonica is distributed in lowlands to mountainous areas. Adults appear in fall, winter, and spring (October – June), and are found in mushroom (e. g. Omphalotus japonicus (Kawam.) Kircham. & O. K. Mill), rotten vegetables, animal dropping, and fallen leaves.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFED1283FB2B7BC7FE3C.taxon	diagnosis	Diagnosis. Plesiochara japonica is similar to P. nitida and P. inflexa, but can be distinguished by the following characters: legs shorter (HTL / PL = 1.13 – 1.20); hypomera with posterior part of carina directed posteriorly almost parallel to lateral margin; median lobe of aedeagus with apical lobe thick, bisinuate at ventral margin; sclerites 1 strongly convex and curved laterally at posterior margin; spermatheca with capsule spherical.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFED1283FB2B7BC7FE3C.taxon	discussion	Remarks. According to the original description (Sharp 1874), there should be three syntypes from “ Copper Temple, Nagasaki ”. However, only a male syntype was found in the Natural History Museum, London (BMNH), which MM photographed and dissected in 2011; it is designated as the lectotype herein. We compared the photograph of the dorsal habitus and drawing of the aedeagus of the lectotype examined by MM and other specimens from Japan and concluded that P. fusca is a junior synonym of O. japonica.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFFB1283FDF67DBEFAC7.taxon	vernacular_names	Japanese common name: Kusabira-hanekakushi-zoku	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFFB1283FDF67DBEFAC7.taxon	type_taxon	type species: Plesiochara fusca Sawada, 1989].	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFFB1283FDF67DBEFAC7.taxon	diagnosis	Diagnosis. This genus is distinguished from the other genera of Aleocharini by the combination of the following character states: body elongate; head, pronotum, mesoventrite, metaventrite, abdominal sternites, elytra, and legs covered with setae rather densely; labial palpus with segment II longer, with setae e, f located far remote from mp (Sawada 1989: Fig. 14 C); pronotum almost hexagonal; hypomera visible in lateral view; elytra longer than pronotum; inner coxal process of mesoventrite narrow, elongate, reaching beyond the half of mesocoxal cavities; surface of fore and middle tibia not covered with spines; median lobe of aedeagus fusiform; apical lobe rather thin from lateral view, dilated at apical in ventral view; internal sac composed of multiple sclerites; one pair of sclerites 1 (sc 1) located near apex of apical lobe; one pair of sclerites 2 (sc 2) located dorsal side of sclerites 1, oval at anterior part and arm-shaped at posterior part; sclerite 3 (sc 3) located basal side of sclerites 2; flagellum long, curved ventrally and bifurcated at base; spermatheca with capsule (ca) spherical or ovate: chamber (ch) short, simple.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF87FFFB1283FDF67DBEFAC7.taxon	discussion	Remarks. This genus is well characterized by the structure of aedeagus: shapes of sclerites 1 – 3 and flagellum as mentioned above though their polarities are unknown. Plesiochara is similar to Pseudocalea Luze in general appearance, but can be separated by the structure of aedeagus and short chamber of spermatheca. Key to species of Plesiochara	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF80FFE21283FDE279EBFE88.taxon	description	(Figs 4 A – F, 5, 12 B)	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF80FFE21283FDE279EBFE88.taxon	description	Redescription. Body elongate; head, thorax and abdomen dark brown; antennae dark reddish brown, with basal part paler; mouth parts and legs reddish yellow; elytra reddish yellow with posterolateral portion darker; head, thorax, abdominal sternites, elytra, and legs covered with setae rather densely; abdominal tergites sparsely covered with setae; dorsal surface of head and pronotum densely covered with reticulation; elytra weakly rugose; abdominal tergites glossy. Head circular, slightly longer than or as long as wide; postocular parts roundly narrowed toward base; eyes ovate. Antennae slender, longer than a combination of head and pronotum; segments IV – V slightly longer than wide; segments VI – VII as long as wide; segments VIII – IX slightly transverse; segments X as long as wide; segments XI ovate. Pronotum almost hexagonal, slightly wider than long; anterior margin widely rounded; lateral margin moderately rounded, narrowed more strongly to posterior than anterior; anterior corners rounded; posterior corners obtusely angulated; hypomera visible in lateral view, with posterior part of carina (Fig. 12 B) directed to posterolateral corners obliquely. Mesoventrite shortly carinate at base of midline; inner coxal process narrow, elongate, reaching to metaventrite process. Metaventrite with inner coxal process short, moderately narrow. Elytra wider than long, longer and wider than pronotum, weakly diverging to posteriorly, widest near posterolateral corners; hind margin sinuate near posterolateral corners. Hind wings developed. Abdomen moderately elongate, parallel sided; tergites III – VII with pores at basal impression; VIII tergite with posterior margin smooth, shallowly emarginate medially in both sexes. VIII sternum with posterior margin smooth, widely rounded in both sexes. Legs slender, long (HTL / PL = 1.35 – 1.47); surface of fore and middle tibia not covered with spines; hind tarsal segments I longer than a combination of II and III. Male: Median lobe of aedeagus (Fig. 4 B – E) rather large; apical lobe rather thin, weakly curved dorsally at apex, narrower, rather strongly constricted in ventral view at basal 1 / 3; apical lobe in lateral view; sclerites 1 convex at posterior margin, widely emarginate at anterior margin; sclerites 4 (sc 4) rather indistinct, located at posterior apex of sclerites 2. Female: Spermatheca (Fig. 4 F) with capsule ovate, longer than chamber. Measurements. BL 4.7 – 5.9, FBL 2.57 – 3.20, HW 0.75 – 0.85, HL 0.76 – 0.86, PW 0.84 – 1.13, PL 0.80 – 1.03, EW 1.23 – 1.48, EL 0.99 – 1.27, HTL 1.10 – 1.39. Specimens examined. JAPAN: Honshu: [Aomori Pref.]: 1 unsexed, Shiroiwa, Osaki, Hirakawa-shi, 21. V. 1994, T. Ozaki leg. (cI); [Yamagata Pref.]: 1 unsexed, Shirakozawa, Oguni-machi, 5. VI. 2010, I, Matoba leg. (cI); [Fukushima Pref.]: 1 male, Noji-onsen, Fukushima-shi, 25. X. 2003, T. Watanabe leg. (KUM); 1 male, 2 female, Futamata-onsen, Tenei-mura, 19. X. 2003, S. Nomura leg. (KUM); [Gunma Pref.]: 1 male, 1 female, Tamahara-kogen, Kamihocchi-machi, Numata-shi, 26. X. 2003, N. Kanai leg. (KUM); 1 female, Narahara, Ueno-mura, 8. IV. 2007, T. Horiguchi leg. (KUM); 1 female, Akegasawa, Narahara, Ueno-mura, 30. IV. 2007, T. Horiguchi leg. (KUM); 1 unsexed, Nagamine-koen, Shibukawa-shi, 16. IV. 2003, N. Kanai leg. (KUM); [Tochigi Pref.]: 1 female, Mt. Shibakusa-yama, Nikko-shi, 9. IV. 1995, H. Ohkawa leg. (KUM); [Kanagawa Pref.]: 1 female, Mt. Ôyama, 25. XII. 1966, Y. Shibata leg. (KUM); [Ishikawa Pref.]: 1 male, Hiraguri, Kanazawa-shi, 21. IV. 1983, Y. Hayashi leg. (KUM); [Fukui Pref.]: 1 female, Hakuba-dô cave, Ôno-shi, 22. VII. 2001, W. Takahashi leg. (KUM); [Yamanashi Pref.]: 1 male, near Dôsaka Tunnel, Dôshi-mura, 16. XI. 1996, T. Watanabe leg. (KUM); [Nagano Pref.]: 1 unsexed, Shimashimadani, Matsumoto-shi, 900 m, 7 – 10. VI. 2008, H. Asaki leg. (KUM); 1 unsexed, ditto, 800 m, 21 – 26. V. 2008, H. Asaki leg. (KUM); 2 unsexed, ditto, 800 m, 10 – 16, X. 2008, H. Asaki leg. (KUM); 2 male, 3 unsexed, Nagawa, Matsumoto-shi, 1500 m, 31. X – 6. XI. 2008, H. Asaki leg. (KUM); 1 male, 1 female, Konami, Suwa-shi, 6. III. 2021, T. Hashizume leg. (KUM); [Kyoto Pref.]: 1 unsexed, Otokoyama, Yawata-shi, Kyoto Pref., 19. II. 2011, T. Ito leg., by rotten vegetables trap (cI); [Nara Pref.]: 1 unsexed, Nara park, Nara-shi, 13. XII. 1987, S. Takahashi leg. (cI); 1 male, Ôdaigahara, Kamikitayama-mura, 25. V – 6. VI. 1986, S. Naomi leg. (KUM); [Wakayama Pref.]: 1 unsexed, Hirai, Kozagawa-chô, 22. V. 2008, I. Matoba leg. (cI); [Okayama Pref.]: 1 male, 1 female, Mt. Taki-yama, Nagi-chô, Katsuta-gun, 500 – 700 m, 16. III. 2003, Y. Fujitani leg. (KUM); 1 male, ditto, 23. XI. 2003, Y. Fujitani leg. (KUM); 1 female, ditto, 7. XII. 2003, Y. Fujitani leg. (KUM); 1 female, ditto, 28. III. 2004, Y. Fujitani leg. (KUM); [Yamaguchi Pref.]: 1 female, Nodani, Tokuji-chô, 31. III. 2004, Y. Fujitani leg. (KUM); Shikoku: [Tokushima Pref.]: 2 male, 2 unsexed, Mt. Tsurugi-san, 15 – 17. X. 1980, S. Naomi leg. (KUM); unsexed, ditto, 19 – 20. VI. 1981, S. Naomi leg. (KUM); 11 unsexed, Mt. Maruzawa-yama, Ichiu-son, 1410 m, 11. VI – 6. VIII. 2000, M. Yoshida leg. (KUM); 1 unsexed, Nishinô-no-Mizuana cave, Aioi-chô, Naka, 26. IV. 1986, H. Bando leg. (KUM); [Ehime Pref.]: 1 female, riverside of Fukumi-gawa river, Matsuyama-shi, 27. XI – 3. XII. 1986, M. Hiraoka leg. (KUM); 2 unsexed, ditto, 27. XI – 3. XII. 1986, M. Hiraoka leg. (KUM); Kyushu: [Saga Pref.]: 1 unsexed, Makishima-yama, Seto-chô, Imari-shi, 35 m, 24. II. 2021, S. Kakizoe leg., from dog dung (KUM); [Ôita Pref.]: 1 male, Taiko-bashi-no-ana cave, Kadota, Takeda-shi, 322 m, 26. IV – 10. V. 2008, K. Ito leg. (KUM) (KUM); Hirado Is.: [Nagasaki Pref.]: 1 male, 1 female, 2 unsexed, Mt. Yasuman-dake, Yamanaka-chô, Hirado-shi, 420 m, 16. II. 2021, S. Kakizoe & S. Inoue leg., from dog dung (KUM); Tsushima Is.: [Nagasaki Pref.]: 1 female, Meishinodan-yama, Uchiyama, Izuhara-machi, Tsushima-shi, 175 m, 20 – 21. II. 2022, S. Kakizoe leg., from dog dung (KUM).	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF80FFE21283FDE279EBFE88.taxon	distribution	Distribution. Japan: Honshu, Shikoku, Kyushu, Hirado Is., and Tsushima Is. (Fig. 5). Bionomics. The collecting records show that Plesiochara nitida is distributed in lowlands to mountainous areas. Adults appear in fall, winter, and spring (October – June) and are found in animal dropping and fallen leaves. Some specimens were collected in caves.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF80FFE21283FDE279EBFE88.taxon	diagnosis	Diagnosis. Plesiochara nitida is similar to P. japonica and P. inflexa, but can be distinguished by genital parts: median lobe of aedeagus with apical lobe thin, weakly curved dorsally at apex; spermatheca with capsule ovate, large, longer than chamber.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF8FFFE71283FE6B7D16F808.taxon	description	(Figs 6 A – F, 7 A – D, 8, 12 D – F) Description. Body elongate; head, thorax and abdomen dark brown; antennae dark reddish brown, with basal part paler; mouth parts and legs reddish yellow; elytra reddish yellow with posterolateral portion darker; head, thorax, abdominal sternites, elytra, and legs covered with setae rather densely; abdominal tergites sparsely covered with setae; dorsal surface of head and pronotum densely covered with reticulation; elytra weakly rugose; abdominal tergites glossy. Head circular, slightly longer than or as long as wide; postocular parts roundly narrowed toward base; eyes ovate. Antennae slender, longer than a combination of head and pronotum; segments IV – V slightly longer than wide; segments VI – VII as long as wide; segments VIII – IX slightly transverse; segments X as long as wide; segments XI ovate. Pronotum almost hexagonal, slightly wider than long; anterior margin widely rounded; lateral margin moderately rounded, narrowed more strongly to posterior than anterior; anterior corners rounded; posterior corners obtusely roundly angulated; hypomera visible in lateral view, with posterior part of carina (Fig. 12 D – F) directed to posterolateral corners obliquely and curved inward at posterior end in most cases. Mesoventrite shortly carinate at base of midline; inner coxal process narrow, elongate, reaching to metaventrite process. Metaventrite with inner coxal process short, moderately narrow. Elytra wider than long, longer and wider than pronotum, weakly diverging to posteriorly, widest near posterolateral corners; hind margin sinuate near posterolateral corners. Hind wings developed. Abdomen moderately elongate, parallel sided; tergites III – VII with small pores at basal impression; VIII tergite with posterior margin smooth, shallowly emarginate medially in both sexes. VIII sternum with posterior margin smooth, widely rounded in both sexes. Legs slender, rather long (HTL / PL = 1.27 – 1.40); surface of fore and middle tibia not covered with spines; hind tarsal segments I longer than a combination of II and III. Male: Median lobe of aedeagus (Fig. 6 B – E) rather large; apical lobe rather thin, curved ventrally; sclerites 1 oblong, narrowed ventrally, curved posteriorly at ventral apex; one pair of sclerites 4 (sc 4) rather distinct, located near apex of sclerites 2, strongly curved inward. Female: Spermatheca (Fig. 6 F) with capsule ovate, small, almost as long as chamber. Measurements. BL 4.5 – 5.5, FBL 2.63 – 3.10, HW 0.74 – 0.77, HL 0.76 – 0.80, PW 0.93 – 1.07, PL 0.83 – 0.98, EW 1.33 – 1.44, EL 1.01 – 1.21, HTL 1.05 – 1.32. Variation. Tanzawa mountains (Mt. Ishihodo-yama, Kanagawa Pref.; near Dôsaka tunnel, Yamanashi, Pref.): Hypomera with carina not curved inward at posterior end (Fig. 12 F); legs longer; median lobe of aedeagus with apical lobe thin, rather weakly curved ventrally, narrower in ventral view (Fig. 7 A – B). Shikoku (Mt. Tsurugi-san, Tokushima Pref.): median lobe of aedeagus with apical lobe thin, rather weakly curved ventrally, narrower in ventral view (Fig. 7 C – D). Hypomeral carina curved weakly or not curved in eastern localities (Fig. 12 E), curved strongly in northern localities (Fig. 12 D).	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF8FFFE71283FE6B7D16F808.taxon	materials_examined	Type materials. Holotype: male, Wakasugi forest, Nishiawakura-son, Okayama Pref., JAPAN, 22. X. 2009, T. Ito leg., from mushroom (KUM). Paratype. JAPAN: Hokkaido: [Hokkaido Pref.]: 1 unsexed, Kiyosato-chô, 27. X. 2009, K. Matsumoto leg. (cI); 1 unsexed, Kamimuri, Maruseppu-chô, 21. V. 2000, M. Maruyama leg., from bear dung (KUM); 1 unsexed, Nagahashi, Naebo, Otaru-shi, 1. V. 1992, M. Ôhara leg., by bait trap (KUM); 5 male, 3 female, 5 unsexed, ditto, 8. X. 1992, M. Ôhara leg. By bait trap (OM, KUM); 2 unsexed, ditto, 15. X. 1992, M. Ôhara leg. By bait trap (KUM); 3 female, 1 unsexed, ditto, 22. X. 1992, M. Ôhara leg. By bait trap (OM, KUM); 2 male, 2 female, 1 unsexed, ditto, 29. X. 1992, M. Ôhara leg. By bait trap (KUM); 1 male, ditto, 5. XI. 1992, M. Ôhara leg. By bait trap (KUM); 2 female, Kakkumi-chô, Hakodate-shi, 12. IV. 2020, Y. Tasaku leg. (KUM); Honshu: [Aomori Pref.]: 1 male, Oirase, Fukaura-chô, 9. VII. 1989, A. Abe leg. (KUM); [Yamagata Pref.]: 1 male, 1 unsexed, Ichino-taki, Chôkai-zan, Sakata-shi, 10. X. 2015, H. Ooki leg. (KUM, cO); [Fukushima Pref.]: 1 male, 1 unsexed, Futamata-onsen, Ten-ei-mura, 850 m, 19. X. 2003, S. Nomura leg. (KUM); [Tochigi Pref.]: 1 female, Okukinu, Kuriyama-mura, 30. V. 1991, S. Naomi leg. (KUM); [Gunma Pref.]: 1 male, 3 unsexed, Marunuma, Katashina-mura, 23. IX. 1969, Y. Shibata leg. (KUM); 4 male, 1 female, Tamahara-kogen, Kamihocchi-machi, Numata-shi, 26. X. 2003, N. Kanai leg. (KUM); 2 unsexed, Aratosawa, Mt. Hotaka-san, Katashina-mura, 950 m, 15. X. 2000, H. Kamezawa leg. (KUM); 2 male, 1 unsexed, Kirizumi spa, Yasunaka-shi, 29. IV. 1958, Y. Shibata leg. (KUM); 1 unsexed, ditto, 18. VI. 1958, Y. Shibata leg. (KUM); [Kanagawa Pref.]: 6 males, 1 female, 3 unsexed, Mt. Ishihodo-yama, Yamakita-machi, Kanagawa Pref., 9. X. 1995, T. Watanabe leg. (KUM). [Toyama Pref.]: 1 unsexed, Masagosawa, Mt. Tsurugi-dake, Tateyama-machi, 18. VII. 1962, Y. Hayashi leg. (KUM); [Fukui Pref.]: 6 male, 2 female, 2 unsexed, Kôkura, Imajo-chô, 14 – 18. X. 1996, K. Ishida et al. leg. (KUM); 1 male, Mt. Hyakuri-ga-take, Obama-shi, 5. V. 1979, H. Sasaji leg. (KUM); [Yamanashi Pref.]: 1 unsexed, Norogawa-deai, Minami-Alps-shi, 3. VII. 1980, H. Yamazaki leg. (KUM); 1 female, Kitazawa-tôge pass, Minami-Alps-shi, 28 – 30. VI. 1980, H. Yamazaki leg. (KUM); 1 male, near Dôsaka tunnel, Dôshi-mura, 16. XI. 1996, T. Watanabe leg. (KUM); [Nagano Pref.]: 1 unsexed, Hakuba-mura, 20. X. 2007, T. Ito leg. (cI); 3 unsexed, Norikura, Matsumoto-shi, 1500 m, 26. V – 1. VI. 2008, H. Asaki leg. (KUM); 1 unsexed, ditto, 2100 m, 27. VIII – 02. IX. 2008, H. Asaki leg. (KUM); 5 unsexed, ditto, 1800 m, 18 – 22. IX 2008, H. Asaki leg. (KUM); 1 unsexed, Nagawa, Matsumoto-shi, 1500 m, 11 – 14. VI. 2008, H. Asaki leg. (KUM); 4 unsexed, ditto, 1600 m, 10 – 16. IX. 2008, H. Asaki leg. (KUM); 2 unsexed, ditto, 1500 m, 31. X – 6. XI. 2008, H. Asaki leg. (KUM); [Gifu Pref.]: 2 unsexed, Hirayu, Takayama-shi, 12. X. 1980, K. Konishi leg. (KUM); 1 unsexed, Nishihotaka-guchi, Takayama-shi, 11. VI. 1980 (KUM); 2 unsexed, Mannami River, Miyagawa, Hida-shi, 5. XI. 1980, S. Saito leg. (KUM); 1 male, 1 female, 4 unsexed, Ooshirakawa, Shirakawa-mura, 26. X. 1980, Y. Takai leg. (KUM); 3 unsexed, Shirakawa-super-rindsuô, Shirakawa-mura, 11. X. 1997, Y. Takai leg.; 3 unsexed, Magari – Sanpôiwa, Hakusan, Shirakawa-mura, 13. X. 1996, Y. Takai leg.; 1 unsexed, Shirakawa-mura, 22. X. 2006, K. Toyoshima leg. (cI); 1 unsexed, Mt. Fukube-ga-take, Minami-mura, Gujô-shi, 12. V. 2002, A. Sugimura leg. (KUM); [Kyoto Pref.]: 1 male, 1 female, Sugio-tôge pass, Ashiu, Nantan-shi, 15. X. 1993, K. Setsuda leg., from Omphalotus japonicus (Kawam.) Kircham. & O. K. Mill. (KUM); [Nara Pref.]: Kamikitayama-mura, 4. X. 2009, T. Ito leg., from mushroom (KUM, cI); [Tottori Pref.]: 3 female, 2 unsexed, Daisen, 8. XI. 1977, S. Naomi leg. (KUM); [Okayama Pref.]: 1 male, 2 female, 4 unsexed, Wakasugi forest, Nishiawakura-son, 22. X. 2009, T. Ito leg., from mushroom (KUM, cI); 1 male, Wakasugi-tôge pass, Nishiawakura-son 8. V. 1977, Watanabe leg. (KUM); [Hiroshima Pref.]: 4 unsexed, Mt. Tateeboshi-yama, Saijô-chô, Shôbara-shi, 30. IV. 2022, S. Inoue leg., by shifting leaf litter (KUM); Oki Isls.: [Shimane Pref.]: 1 male, Minamidani-rindo, Dougo, Okinoshima-chô, 10. XI. 2004, T. Watanabe leg. (KUM); Shikoku: [Tokushima Pref.]: 4 male, 3 unsexed, Mt. Tsurugi-san, 15 – 17. X. 1980, S. Naomi leg. (KUM).	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF8FFFE71283FE6B7D16F808.taxon	distribution	Distribution. Japan: Hokkaido, Honshu, Oki Isls., and Shikoku (Fig. 8). Bionomics. The collecting records show that P. inflexa is distributed in lowlands to mountainous areas of Hokkaido, and mountainous areas to subalpine areas in Honshu and Shikoku. Adults appear in fall and spring to early summer (late September – November, April – early July), and are found in mushroom (e. g. Omphalotus japonicus (Kawam.) Kircham. & O. K. Mill), animal droppings, and fallen leaves.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF8FFFE71283FE6B7D16F808.taxon	diagnosis	Diagnosis. Plesiochara inflexa is similar to P. japonica and P. nitida but can be distinguished by the sexual characters median lobe of aedeagus with apical lobe thin, entirely curved ventrally; sclerites 1 oblong narrowed ventrally, curved posteriorly at ventral apex; spermatheca with capsule ovate, almost as long as chamber.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF8FFFE71283FE6B7D16F808.taxon	etymology	Etymology. The specific name alludes to the shape of sclerites 4, which are strongly curved inwards.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF88FFFB1283FF577B7FFD44.taxon	description	(Figs 9 A – F, 10 A – D, 11) Description. Body elongate, head, thorax and abdomen reddish brown to light brown, darker in head and posterior part of abdomen; antennae reddish brown, with basal part paler; mouth parts and legs reddish yellow; elytra reddish yellow with posterolateral portion darker; head, thorax, abdominal sternites, elytra, and legs covered with setae rather densely; abdominal tergites sparsely covered with setae; dorsal surface of head and pronotum without reticulation; elytra weakly rugose; abdominal tergites glossy. Head circular, slightly longer than or as long as wide; postocular parts roundly narrowed toward base; eyes ovate. Antennae slender, longer than a combination of head and pronotum; segments IV – V slightly longer than wide; segments VI – VII as long as wide; segments VIII – IX slightly transverse; segments X as long as wide; segments XI ovate. Pronotum almost hexagonal, slightly wider than long; anterior margin widely rounded; lateral margin moderately rounded, narrowed more strongly to posterior than anterior; anterior corners rounded; posterior corners obtusely roundly angulated; hypomera visible in lateral view, with posterior part of carina directed to posterolateral corners obliquely. Mesoventrite shortly carinate at base of midline; inner coxal process narrow, elongate, reaching to metaventrite process. Metaventrite with inner coxal process short, moderately narrow. Elytra wider than long, longer and wider than pronotum, weakly diverging to posteriorly, widest near posterolateral corners; hind margin sinuate near posterolateral corners. Hind wings developed. Abdomen moderately elongate, parallel sided; tergites III – VII with small pores at basal impression; VIII tergite with posterior margin smooth, shallowly emarginate medially in both sexes. VIII sternum with posterior margin smooth, widely rounded in both sexes. Legs slender, rather long (HTL / PL = 1.41 – 1.54); surface of fore and middle tibia not covered with spines; hind tarsal segments I distinctly longer than combination of II and III. Male: Median lobe of aedeagus (Figs 9 B – E, 10 A – D), rather small; apical lobe rather thin, curved ventrally; sclerites 1 convex at posterior margin; sclerites 4 indistinct; flagellum long. Female: Spermatheca (Fig. 9 F) with capsule ovate, almost as long as chamber. Variation. Honshu: Median lobe of aedeagus with apical lobe not thickened at apex, moderately constricted in ventral view (Fig. 10 A – B). Shikoku: Median lobe of aedeagus with apical lobe thickened at apex, moderately constricted in ventral view (Fig. 9 B – D). Kyushu: Median lobe of aedeagus with apical lobe not thickened at apex, constricted rather strongly in ventral view (Fig. 10 C – D). Measurements. BL 4.4 – 5.5, FBL 2.50 – 3.07, HW 0.65 – 0.75, HL 0.69 – 0.78, PW 0.85 – 0.98, PL 0.78 – 0.90, EW 1.17 – 1.36, EL 0.92 – 1.14, HTL 1.12 – 1.35.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF88FFFB1283FF577B7FFD44.taxon	materials_examined	Type materials. Holotype: 1 male, Mt. Narabara-yama, 3. XI. 1968, M. Takagi leg. (KUM). Paratype: JAPAN: Honshu: [Fukui Pref.]: 2 female, Kôkura, Imajo-chô, 14 – 18. X. 1996, K. Ishida leg. (KUM); [Shiga Pref.]: 1 unsexed, Oisugi forest, Kutsuki, 28. X. 2008, T. Ito leg. (cI); [Kyoto Pref.]: 5 male, 6 female, 1 unsexed, Sugiotôge pass, Ashiu, Nantan-shi, 15. X. 1993, K. Setsuda leg., from Omphalotus japonicus (Kawam.) Kircham. & O. K. Mill. (KUM); 1 unsexed, Sasari-tôge pass, 26. X. 2006, T. Ito leg. (cI); [Okayama Pref.]: 1 male, 3 unsexed, Wakasugi Forest, Nishiawakura-son, 22. X. 2009, T. Ito leg., from mushroom (KUM, cI); [Hiroshima Pref.]: 2 male, 90 unsexed, Mt. Tateeboshi-yama, Saijô-chô, Shôbara-shi, 1175 m, 14. X. 2019, Y. Senda leg., from Omphalotus japonicus (Kawam.) Kircham. & O. K. Mill. (HMNH, KUM); 1 unsexed, ditto, 30. IV. 2022, S. Inoue leg., by shifting leaf litter (KUM); Shikoku: [Ehime Pref.]: 3 male, 3 female, 5 unsexed, Mt. Narabara-yama, 3. XI. 1968, M. Takagi leg. (HMNH, KUM); 1 male, 1 unsexed, Mt. Shiratsue, Matsuyama-shi, 780 m, 10 – 23. X. 1986, T. Nagata leg. (HMNH, KUM); 1 male, 1 unsexed, Mt. Koya, Nomura-chô, Seiyo City, 1250 – 1300 m, 28. XI. 2008, T. Kurihara leg. (HMNH, KUM); 2 unsexed, Saragamine, Kumakôgen Town, 1100 – 1270 m, 4. X. 2008, T. Kurihara leg. (HMNH, KUM); Kyushu: [Kumamoto Pref.]: 3 male, 2 female, 4 unsexed, Kirihagi, Yamato-chô, 15. X. 1993, Y. Tomishima leg. (KUM); 2 unsexed, Mt. Heike-yama, Gokanosho, Yashiro-shi, 17. X. 1988, S. Naomi leg. (KUM); 1 male, Naidaijin, Yamato-chô, 22. X. 1982, M. Matsuzaki leg. (KUM); 1 male, 1 female, 3 unsexed, Mt. Ichifusayama, Minakami-mura, 13. X. 1995, Y. Tomishima leg. (KUM).	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF88FFFB1283FF577B7FFD44.taxon	distribution	Distribution. Japan: Honshu, Shikoku, and Kyushu (Fig. 11). Bionomics. The collecting records show that P. rufula is distributed in mountainous areas. Adults appear in fall and spring (October – November, April), and are found in mushroom (e. g. Omphalotus japonicus) and fallen leaves.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF88FFFB1283FF577B7FFD44.taxon	diagnosis	Diagnosis. P. rufula can be easily distinguished from other congeners by the following character states: head, thorax, and abdomen reddish brown to light brown, darker in head and posterior part of abdomen; dorsal surface of head and pronotum without reticulation; basal impressions of abdominal tergites III – VII with large pores. P. rufula is similar to P. inflexa regarding the median lobe of the aedeagus, but can be distinguished by the following characters: size smaller; sclerites 1 convex at posterior margin, not curved posteriorly at ventral apex; sclerites 4 indistinct.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
FA188782FF88FFFB1283FF577B7FFD44.taxon	etymology	Etymology. The specific name alludes to the reddish body.	en	Inoue, Shûgo, Maruyama, Munetoshi (2022): Revision of the genus Plesiochara Sawada (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 5165 (4): 501-519, DOI: https://doi.org/10.11646/zootaxa.5165.4.3
